211 lines
26 KiB
XML
211 lines
26 KiB
XML
<document ID-DOI="10.11646/zootaxa.5256.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7745281" ID-ZooBank="CE0AC7CD-E55D-484C-9346-86F16C985DBE" approvalRequired="29" approvalRequired_for_taxonomicNames="9" approvalRequired_for_textStreams="19" approvalRequired_for_treatments="1" checkinTime="1679054310250" checkinUser="plazi" docAuthor="Serrano, Artur R. M." docDate="2023" docId="AE0487C8F526FFCD0986A07B65A0F9DD" docLanguage="en" docName="zootaxa.5256.1.1.pdf" docOrigin="Zootaxa 5256 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrocheilus proximus Peringuey 1896" docType="treatment" docVersion="2" lastPageNumber="22" masterDocId="523DFFB0F533FFD80911A31C612BFFA8" masterDocTitle="Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes" masterLastPageNumber="35" masterPageNumber="1" pageNumber="22" updateTime="1679055118150" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Serrano, Artur R. M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-4282-6073</mods:nameIdentifier>
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<mods:affiliation>cE 3 c-Center for Ecology, Evolution and Environmental Changes & CHANGE - Global Change and Sustainability Institute, Departamento de Biologia Animal, Faculdade de Ciências da Universidade de Lisboa, R. Ernesto de Vasconcelos, Ed. C 2, Campo Grande, 1749 - 016 Lisboa, PORTUGAL. aserrano @ fc. ul. pt; http: // orcid. org / 0000 - 0003 - 4282 - 6073</mods:affiliation>
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<mods:nameIdentifier type="email">aserrano@fc.ul.pt</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5256</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:start>1</mods:start>
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<mods:end>35</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.1.1</mods:url>
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<mods:identifier type="DOI">10.11646/zootaxa.5256.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7745281</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7745382" ID-Zenodo-Dep="7745382" LSID="urn:lsid:plazi:treatment:AE0487C8F526FFCD0986A07B65A0F9DD" httpUri="http://treatment.plazi.org/id/AE0487C8F526FFCD0986A07B65A0F9DD" lastPageNumber="22" pageId="21" pageNumber="22">
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<subSubSection box="[151,645,870,898]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph blockId="21.[151,645,870,934]" box="[151,645,870,898]" pageId="21" pageNumber="22">
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<heading bold="true" box="[151,645,870,898]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
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<taxonomicName ID-CoL="72SCJ" authority="Peringuey, 1896" authorityName="Peringuey" authorityYear="1896" box="[151,645,870,898]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="proximus">
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<emphasis bold="true" box="[151,645,870,898]" pageId="21" pageNumber="22">
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<emphasis bold="true" box="[151,439,871,898]" italics="true" pageId="21" pageNumber="22">Macrocheilus proximus</emphasis>
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<bibRefCitation author="Peringuey, L." box="[446,645,870,897]" pageId="21" pageNumber="22" pagination="125 - 623" refId="ref25799" refString="Peringuey, L. (1896) A descriptive catalogue of the Coleoptera of South Africa. Part II. Carabidae. Transactions of the South African Philosophical Society, 7, 125 - 623, pls. III - X. https: // doi. org / 10.5962 / bhl. title. 9230" type="journal article" year="1896">Péringuey, 1896</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,374,907,934]" pageId="21" pageNumber="22" type="description">
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<paragraph blockId="21.[151,645,870,934]" box="[151,374,907,934]" pageId="21" pageNumber="22">
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<figureCitation box="[151,267,907,934]" captionStart="FIGURE 9" captionStartId="22.[152,255,1916,1940]" captionTargetBox="[238,1445,163,1876]" captionTargetId="figure-82@22.[219,1454,151,1889]" captionTargetPageId="22" captionText="FIGURE 9. Habitus of Macrocheilus (dorsal view). a, M. longicollis (holotype); b, M. longicollis, MOZ (PSC); c, M. longicollis, ZAM (TAC); d, M. persimilis (holotype, plus labels); e, M. persimilis, ZAM (TAC); f, M. proximus (holotype); g, M. proximus, ETI (TAC); h, M. quadrinotatus (holotype, plus labels); i, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745301" httpUri="https://zenodo.org/record/7745301/files/figure.png" pageId="21" pageNumber="22">Figs. 9f, g</figureCitation>
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,
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<figureCitation box="[279,321,907,933]" captionStart="FIGURE 10" captionStartId="23.[152,255,1844,1868]" captionTargetBox="[166,1426,707,1780]" captionTargetId="figure-241@23.[152,1436,697,1791]" captionTargetPageId="23" captionText="FIGURE 10. Labrum of Macrocheilus (dorsal view). a, M. longicollis, MOZ (PSC); b, M. persimilis, ZAM (TAC); c, M. proximus, ETI (TAC); d, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745303" httpUri="https://zenodo.org/record/7745303/files/figure.png" pageId="21" pageNumber="22">10c</figureCitation>
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,
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<figureCitation box="[333,374,907,933]" captionStart="FIGURE 14" captionStartId="27.[152,255,1916,1940]" captionTargetBox="[421,1245,991,1878]" captionTargetId="figure-120@27.[413,1253,983,1890]" captionTargetPageId="27" captionText="FIGURE 14. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. proximus, ETI (TAC);; b, M. quadrinotatus, ANG (ASC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, RSA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745313" httpUri="https://zenodo.org/record/7745313/files/figure.png" pageId="21" pageNumber="22">14a</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="materials_examined">
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<paragraph blockId="21.[151,1437,979,1653]" pageId="21" pageNumber="22">
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<emphasis bold="true" box="[151,359,979,1005]" pageId="21" pageNumber="22">Studied material.</emphasis>
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<materialsCitation box="[367,1084,979,1006]" collectingDate="1951-10-14" collectingDateMax="1951-10-18" collectingDateMin="1951-10-14" collectionCode="T, MRAC" collectorName="Janse" location="Barberton" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1">
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<specimenCount box="[367,448,979,1005]" pageId="21" pageNumber="22" type="male">1 male</specimenCount>
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: “
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<location LSID="urn:lsid:plazi:treatment:AE0487C8F526FFCD0986A07B65A0F9DD:23726005F526FFCD08C4A0CF636CFC45" box="[469,583,979,1005]" name="Barberton" pageId="21" pageNumber="22">Barberton</location>
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\
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<date box="[600,758,979,1006]" pageId="21" pageNumber="22" value="1951-10-14" valueMax="1951-10-18" valueMin="1951-10-14">
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<collectingDate box="[600,758,979,1006]" pageId="21" pageNumber="22" value="1951-10-14" valueMax="1951-10-18" valueMin="1951-10-14">14–18.X.1951</collectingDate>
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</date>
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\ A.J.
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<collectionCode box="[818,838,979,1005]" name="Tavera, Department of Geology and Geophysics" pageId="21" pageNumber="22">T</collectionCode>
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.
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<collectorName box="[849,910,980,1006]" pageId="21" pageNumber="22">Janse</collectorName>
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leg.” (
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<collectionCode box="[987,1076,979,1005]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="21" pageNumber="22" type="Museum">MRAC</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation collectingDate="1971-04" collectionCode="MRAC" collectorName="A. Allaen" location="Katanga" municipality="Kasenga" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1">
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<specimenCount box="[1097,1200,979,1005]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
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: “
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<location LSID="urn:lsid:plazi:treatment:AE0487C8F526FFCD0986A07B65A0F9DD:23726005F526FFCD0DD4A0CF6402FC45" box="[1221,1321,979,1005]" municipality="Kasenga" name="Katanga" pageId="21" pageNumber="22">Katanga</location>
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:
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<collectingMunicipality box="[1332,1428,979,1005]" pageId="21" pageNumber="22">Kasenga</collectingMunicipality>
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\?.
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<date box="[166,258,1015,1041]" pageId="21" pageNumber="22" value="1971-04">
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<collectingDate box="[166,258,1015,1041]" pageId="21" pageNumber="22" value="1971-04">IV.1971</collectingDate>
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</date>
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\ don
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<collectorName box="[327,438,1015,1041]" pageId="21" pageNumber="22">A. Allaen</collectorName>
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leg.” (
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<collectionCode box="[518,607,1015,1041]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="21" pageNumber="22" type="Museum">MRAC</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation collectingDate="1998-04-27" collectorName="Werner & Th. Assmann" country="Ethiopia" determinerName="Th. Assmann & A. Serrano" location="Ethiopia" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1">
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<specimenCount box="[629,734,1015,1041]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
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: “
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<collectingCountry box="[756,856,1015,1041]" name="Ethiopia" pageId="21" pageNumber="22">Ethiopia</collectingCountry>
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, 25/
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<date box="[907,1029,1015,1042]" pageId="21" pageNumber="22" value="1998-04-27">
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<collectingDate box="[907,1029,1015,1042]" pageId="21" pageNumber="22" value="1998-04-27">27.IV.1998</collectingDate>
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</date>
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\ near Negele Borana\ Sidamo, leg.
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<collectorName box="[151,236,1051,1077]" pageId="21" pageNumber="22">Werner</collectorName>
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” \\ “Collection\
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<collectorName box="[416,563,1051,1077]" pageId="21" pageNumber="22">Th. Assmann</collectorName>
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\ Bleckede, CAB” (yellow label)\\ “
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<taxonomicName authorityName="Serrano" authorityYear="2023" box="[961,1114,1051,1077]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis box="[961,1114,1051,1077]" italics="true" pageId="21" pageNumber="22">Macrocheilus</emphasis>
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</taxonomicName>
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||
\ pr
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||
<emphasis box="[1151,1230,1052,1077]" italics="true" pageId="21" pageNumber="22">oximus</emphasis>
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Péring.” [h]\ “det.
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<determinerName box="[151,302,1087,1113]" pageId="21" pageNumber="22">Th. Assmann</determinerName>
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||
, 2006” [t] \\ “
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<taxonomicName authorityName="Serrano" authorityYear="2023" box="[457,610,1087,1113]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis box="[457,610,1087,1113]" italics="true" pageId="21" pageNumber="22">Macrocheilus</emphasis>
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</taxonomicName>
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\
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<taxonomicName authorityName="Peringuey" authorityYear="1896" box="[625,728,1088,1113]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="proximus">
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<emphasis box="[625,728,1088,1113]" italics="true" pageId="21" pageNumber="22">proximus</emphasis>
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</taxonomicName>
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||
\
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||
<bibRefCitation author="Peringuey, L." box="[742,926,1087,1113]" pageId="21" pageNumber="22" pagination="125 - 623" refId="ref25799" refString="Peringuey, L. (1896) A descriptive catalogue of the Coleoptera of South Africa. Part II. Carabidae. Transactions of the South African Philosophical Society, 7, 125 - 623, pls. III - X. https: // doi. org / 10.5962 / bhl. title. 9230" type="journal article" year="1896">Péringuey, 1896</bibRefCitation>
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” [h]\ “
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<determinerName box="[1001,1118,1087,1113]" pageId="21" pageNumber="22">A.Serrano</determinerName>
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det. 2021” [t] [h]
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</materialsCitation>
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||
.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="discussion">
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<paragraph blockId="21.[151,1437,979,1653]" pageId="21" pageNumber="22">
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<emphasis bold="true" box="[199,315,1123,1149]" pageId="21" pageNumber="22">Remarks.</emphasis>
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The species was originally described from Natal (D`Urban) (
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<collectingCountry box="[1011,1299,1123,1149]" name="South Africa" pageId="21" pageNumber="22">Republic of South Africa</collectingCountry>
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) (
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<bibRefCitation author="Peringuey, L." pageId="21" pageNumber="22" pagination="125 - 623" refId="ref25799" refString="Peringuey, L. (1896) A descriptive catalogue of the Coleoptera of South Africa. Part II. Carabidae. Transactions of the South African Philosophical Society, 7, 125 - 623, pls. III - X. https: // doi. org / 10.5962 / bhl. title. 9230" type="journal article" year="1896">Péringuey 1896</bibRefCitation>
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) (
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<figureCitation box="[235,317,1159,1185]" captionStart="FIGURE 9" captionStartId="22.[152,255,1916,1940]" captionTargetBox="[238,1445,163,1876]" captionTargetId="figure-82@22.[219,1454,151,1889]" captionTargetPageId="22" captionText="FIGURE 9. Habitus of Macrocheilus (dorsal view). a, M. longicollis (holotype); b, M. longicollis, MOZ (PSC); c, M. longicollis, ZAM (TAC); d, M. persimilis (holotype, plus labels); e, M. persimilis, ZAM (TAC); f, M. proximus (holotype); g, M. proximus, ETI (TAC); h, M. quadrinotatus (holotype, plus labels); i, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745301" httpUri="https://zenodo.org/record/7745301/files/figure.png" pageId="21" pageNumber="22">Fig. 9f</figureCitation>
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). The MRAC specimens were determined by P. Basilewsky. The elytral black spot pattern of
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<taxonomicName authorityName="Peringuey" authorityYear="1896" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="proximus">
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<emphasis italics="true" pageId="21" pageNumber="22">M. proximus</emphasis>
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</taxonomicName>
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is slightly similar to that of the
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<taxonomicName box="[616,782,1195,1221]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="basilewskyi">
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<emphasis box="[616,782,1195,1221]" italics="true" pageId="21" pageNumber="22">M. basilewskyi</emphasis>
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</taxonomicName>
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, but the pronotum conformation and the aedeagus of both species are quite different (see
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<figureCitation box="[511,647,1231,1257]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="25.[152,255,1923,1947]" captionStartId-1="26.[152,255,1275,1299]" captionTargetBox-0="[359,1259,530,1904]" captionTargetBox-1="[176,1405,161,1252]" captionTargetId-0="figure-221@25.[333,1270,517,1916]" captionTargetId-1="figure-327@26.[166,1413,151,1264]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURE 11. Habitus of Macrocheilus (dorsal view). a, M. scapularis (MRAC, plus labels); b, M. spectandus (holotype); c, M. spectandus, TAN (TAC); d, M. taedatus (paratype); e, M. taedatus, TAN (TAC); f, M. viduatus katanganus (holotype, plus labels); g, M. viduatus katanganus, ZAM (PSC)." captionText-1="FIGURE 12. Labrum of Macrocheilus (dorsal view). a, M. scapularis (MRAC); b, M. spectandus, TAN (TAC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, ZAM (PSC)." figureDoi-0="http://doi.org/10.5281/zenodo.7745305" figureDoi-1="http://doi.org/10.5281/zenodo.7745307" httpUri-0="https://zenodo.org/record/7745305/files/figure.png" httpUri-1="https://zenodo.org/record/7745307/files/figure.png" pageId="21" pageNumber="22">Figs. 11‒12</figureCitation>
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,
|
||
<figureCitation box="[661,733,1231,1257]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStart-3="FIGURE 16" captionStartId-0="27.[152,255,872,896]" captionStartId-1="27.[152,255,1916,1940]" captionStartId-2="31.[152,255,1943,1967]" captionStartId-3="32.[152,255,1943,1967]" captionTargetBox-0="[370,1262,159,835]" captionTargetBox-1="[421,1245,991,1878]" captionTargetBox-2="[245,1343,189,1903]" captionTargetBox-3="[242,1350,192,1911]" captionTargetId-0="figure-117@27.[365,1272,151,843]" captionTargetId-1="figure-120@27.[413,1253,983,1890]" captionTargetId-2="figure-16@31.[221,1366,181,1920]" captionTargetId-3="figure-19@32.[226,1362,181,1920]" captionTargetPageId-0="27" captionTargetPageId-1="27" captionTargetPageId-2="31" captionTargetPageId-3="32" captionText-0="FIGURE 13. Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. persimilis, ZAM (TAC); b, M. scapularis (MRAC); c, M. spectandus, ZAM (TAC); d, M. viduatus katanganus, ZAM (PSC)." captionText-1="FIGURE 14. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. proximus, ETI (TAC);; b, M. quadrinotatus, ANG (ASC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, RSA (PSC)." captionText-2="FIGURE 15. Habitus of Macrocheilus (dorsal view). a, M. angustatus (holotype, plus labels); b, M. biplagiatus (MRAC, plus labels); c, M. burgeoni (holotype, plus labels); d, M. crampeli (paratype, plus labels); e, M. cruciatus (MRAC, plus labels); f, M. moraisi (holotype, plus labels)." captionText-3="FIGURE 16. Habitus of Macrocheilus (dorsal view). a, M. ocellatus (holotype, plus labels); b, M. overlaeti (holotype, plus labels); c, M. varians varians (holotype); d, M. varians eugrammus (holotype); e, M. vinctus (paratype)." figureDoi-0="http://doi.org/10.5281/zenodo.7745311" figureDoi-1="http://doi.org/10.5281/zenodo.7745313" figureDoi-2="http://doi.org/10.5281/zenodo.7745315" figureDoi-3="http://doi.org/10.5281/zenodo.7745317" httpUri-0="https://zenodo.org/record/7745311/files/figure.png" httpUri-1="https://zenodo.org/record/7745313/files/figure.png" httpUri-2="https://zenodo.org/record/7745315/files/figure.png" httpUri-3="https://zenodo.org/record/7745317/files/figure.png" pageId="21" pageNumber="22">13‒16</figureCitation>
|
||
,
|
||
<quantity box="[747,807,1231,1257]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="21" pageNumber="22" unit="in" value="18.0">18 in</quantity>
|
||
<bibRefCitation author="Serrano, A. R. M." box="[816,973,1231,1257]" pageId="21" pageNumber="22" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano 2000</bibRefCitation>
|
||
). The known distribution in
|
||
<collectingCountry name="South Africa" pageId="21" pageNumber="22">Republic of South Africa</collectingCountry>
|
||
and the new records for
|
||
<collectingCountry box="[568,959,1267,1293]" name="Democratic Republic of the Congo" pageId="21" pageNumber="22">Democratic Republic of the Congo</collectingCountry>
|
||
and
|
||
<collectingCountry box="[1012,1107,1267,1293]" name="Ethiopia" pageId="21" pageNumber="22">Ethiopia</collectingCountry>
|
||
points to a scattered distribution in Africa.
|
||
</paragraph>
|
||
<paragraph blockId="21.[151,1437,979,1653]" pageId="21" pageNumber="22">
|
||
<emphasis box="[199,365,1339,1365]" italics="true" pageId="21" pageNumber="22">Male genitalia</emphasis>
|
||
. Drawings of median lobe in lateral and dorsal views and left paramere were provided by
|
||
<bibRefCitation author="Serrano, A. R. M." pageId="21" pageNumber="22" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano (2000</bibRefCitation>
|
||
,
|
||
<figureCitation box="[288,381,1375,1402]" captionStart="FIGURE 15" captionStartId="31.[152,255,1943,1967]" captionTargetBox="[245,1343,189,1903]" captionTargetId="figure-16@31.[221,1366,181,1920]" captionTargetPageId="31" captionText="FIGURE 15. Habitus of Macrocheilus (dorsal view). a, M. angustatus (holotype, plus labels); b, M. biplagiatus (MRAC, plus labels); c, M. burgeoni (holotype, plus labels); d, M. crampeli (paratype, plus labels); e, M. cruciatus (MRAC, plus labels); f, M. moraisi (holotype, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745315" httpUri="https://zenodo.org/record/7745315/files/figure.png" pageId="21" pageNumber="22">Figs. 15</figureCitation>
|
||
,
|
||
<figureCitation box="[393,424,1375,1401]" captionStart="FIGURE 16" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[242,1350,192,1911]" captionTargetId="figure-19@32.[226,1362,181,1920]" captionTargetPageId="32" captionText="FIGURE 16. Habitus of Macrocheilus (dorsal view). a, M. ocellatus (holotype, plus labels); b, M. overlaeti (holotype, plus labels); c, M. varians varians (holotype); d, M. varians eugrammus (holotype); e, M. vinctus (paratype)." figureDoi="http://doi.org/10.5281/zenodo.7745317" httpUri="https://zenodo.org/record/7745317/files/figure.png" pageId="21" pageNumber="22">16</figureCitation>
|
||
). Median lobe short, slightly knee shaped at the base, dilated in the middle, apical lamella very short, slightly acuminated at apex (dorsal view), downward (lateral view), ostium with one pair of very large scaled and sclerotized stripes on latero-dorsal region, bending downward near apex (
|
||
<bibRefCitation author="Serrano, A. R. M." box="[1121,1278,1447,1473]" pageId="21" pageNumber="22" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano 2000</bibRefCitation>
|
||
,
|
||
<figureCitation box="[1291,1376,1447,1474]" captionStart="FIGURE 15" captionStartId="31.[152,255,1943,1967]" captionTargetBox="[245,1343,189,1903]" captionTargetId="figure-16@31.[221,1366,181,1920]" captionTargetPageId="31" captionText="FIGURE 15. Habitus of Macrocheilus (dorsal view). a, M. angustatus (holotype, plus labels); b, M. biplagiatus (MRAC, plus labels); c, M. burgeoni (holotype, plus labels); d, M. crampeli (paratype, plus labels); e, M. cruciatus (MRAC, plus labels); f, M. moraisi (holotype, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745315" httpUri="https://zenodo.org/record/7745315/files/figure.png" pageId="21" pageNumber="22">Fig. 15</figureCitation>
|
||
); left paramere rounded, slightly longer than wide, without a tooth in postero-dorsal edge (
|
||
<bibRefCitation author="Serrano, A. R. M." box="[1101,1256,1483,1509]" pageId="21" pageNumber="22" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano 2000</bibRefCitation>
|
||
,
|
||
<figureCitation box="[1267,1349,1483,1509]" captionStart="FIGURE 16" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[242,1350,192,1911]" captionTargetId="figure-19@32.[226,1362,181,1920]" captionTargetPageId="32" captionText="FIGURE 16. Habitus of Macrocheilus (dorsal view). a, M. ocellatus (holotype, plus labels); b, M. overlaeti (holotype, plus labels); c, M. varians varians (holotype); d, M. varians eugrammus (holotype); e, M. vinctus (paratype)." figureDoi="http://doi.org/10.5281/zenodo.7745317" httpUri="https://zenodo.org/record/7745317/files/figure.png" pageId="21" pageNumber="22">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="21.[151,1437,979,1653]" pageId="21" pageNumber="22">
|
||
<emphasis box="[199,390,1519,1545]" italics="true" pageId="21" pageNumber="22">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation box="[404,500,1519,1545]" captionStart="FIGURE 14" captionStartId="27.[152,255,1916,1940]" captionTargetBox="[421,1245,991,1878]" captionTargetId="figure-120@27.[413,1253,983,1890]" captionTargetPageId="27" captionText="FIGURE 14. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. proximus, ETI (TAC);; b, M. quadrinotatus, ANG (ASC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, RSA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745313" httpUri="https://zenodo.org/record/7745313/files/figure.png" pageId="21" pageNumber="22">Fig. 14a</figureCitation>
|
||
). Gonocoxite 1 subtriangular; gonocoxite 2 sickle shaped, apex hardly pointed; dorsal surface concave, almost all covered with numerous setae, inner margin with eight setae, outer margin with a sub marginal setose longitudinal rib; ventral surface slightly convex, some minor setae on surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1163,1627,1653]" pageId="21" pageNumber="22" type="distribution">
|
||
<paragraph blockId="21.[151,1437,979,1653]" box="[199,1163,1627,1653]" pageId="21" pageNumber="22">
|
||
<emphasis bold="true" box="[199,353,1627,1653]" pageId="21" pageNumber="22">Distribution.</emphasis>
|
||
<collectingCountry box="[359,754,1627,1653]" name="Democratic Republic of the Congo" pageId="21" pageNumber="22">Democratic Republic of the Congo</collectingCountry>
|
||
,
|
||
<collectingCountry box="[765,863,1627,1653]" name="Ethiopia" pageId="21" pageNumber="22">Ethiopia</collectingCountry>
|
||
,
|
||
<collectingCountry box="[874,1158,1627,1653]" name="South Africa" pageId="21" pageNumber="22">Republic of South Africa</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |