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<document id="1F4FA5037EF8CF44DC268EAD2E73B0E6" ID-DOI="10.5281/zenodo.197793" ID-GBIF-Dataset="8f84f706-94cc-43a5-9027-efd2138aa784" ID-ISSN="1175-5326" ID-Zenodo-Dep="197793" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460278674581" checkinUser="plazi" docAuthor="Braasch, Dietrich &amp; Boonsoong, Boonsatien" docDate="2010" docId="716BF555FF99FFD98FF0FA207EAE67FD" docLanguage="en" docName="zt02610p026.pdf" docOrigin="Zootaxa 2610" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Epeorus nguyenbaeorum Braasch and Boonsoong, sp. nov." docType="treatment" docVersion="9" lastPageNumber="20" masterDocId="8D528D2DFF8BFFCA8F67FF8F7D58656A" masterDocTitle="A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia" masterLastPageNumber="26" masterPageNumber="1" pageNumber="19" updateTime="1698249041875" updateUser="plazi">
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<mods:title id="6CA4A02FDBD2AA9179E3602F66C5CBDB">A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia</mods:title>
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<mods:namePart id="62BECC256AD1C734B6D796A26EBFEF06">Braasch, Dietrich</mods:namePart>
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<mods:namePart id="B4461480EA6331BA456BE98C67B6980D">Boonsoong, Boonsatien</mods:namePart>
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<mods:date id="FFFCAB402F419542DBCBE9DB76425F7D">2010</mods:date>
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<treatment id="716BF555FF99FFD98FF0FA207EAE67FD" ID-DOI="http://doi.org/10.5281/zenodo.6211093" ID-GBIF-Taxon="127804273" ID-Zenodo-Dep="6211093" LSID="urn:lsid:plazi:treatment:716BF555FF99FFD98FF0FA207EAE67FD" httpUri="http://treatment.plazi.org/id/716BF555FF99FFD98FF0FA207EAE67FD" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="B1D817C8FF99FFD88FF0FA207F2563A5" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="F97D4443FF99FFD88FF0FA207E3960A3" blockId="18.[151,865,1455,1481]" box="[151,865,1455,1481]" pageId="18" pageNumber="19">
<heading id="A235F32FFF99FFD88FF0FA207E3960A3" bold="true" box="[151,865,1455,1481]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="CBB69851FF99FFD88FF0FA207E3960A3" bold="true" box="[151,865,1455,1481]" pageId="18" pageNumber="19">
<taxonomicName id="3EC23FC0FF99FFD88FF0FA207FA560A3" ID-CoL="6FNY3" authority="Braasch and Boonsoong" authorityName="Braasch and Boonsoong" box="[151,765,1455,1481]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nguyenbaeorum" status="sp. nov.">
<emphasis id="CBB69851FF99FFD88FF0FA207C9060A3" bold="true" box="[151,456,1455,1481]" italics="true" pageId="18" pageNumber="19">Epeorus nguyenbaeorum</emphasis>
Braasch and Boonsoong
</taxonomicName>
<taxonomicNameLabel id="D085252AFF99FFD88C63FA207E3960A3" box="[772,865,1455,1481]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F97D4443FF99FFD88FF0FA757F2563A5" blockId="18.[151,1438,1530,2004]" pageId="18" pageNumber="19">
This species was figured by Nguyen and Bae (2004) as
<taxonomicName id="3EC23FC0FF99FFD88C7BFA74780F637E" authority="Braasch &amp; Soldan, 1984" authorityName="Braasch &amp; Soldan" authorityYear="1984" box="[796,1367,1530,1556]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hieroglyphicus">
<emphasis id="CBB69851FF99FFD88C7BFA74796A637E" box="[796,1074,1531,1556]" italics="true" pageId="18" pageNumber="19">Epeorus hieroglyphicus</emphasis>
Braasch &amp; Soldán, 1984
</taxonomicName>
, from
<collectingCountry id="81D504D3FF99FFD88FF0F9AF7C5A6350" box="[151,258,1568,1594]" name="Vietnam" pageId="18" pageNumber="19">Vietnam</collectingCountry>
. Prof. Dr. Bae, being corresponding author, was informed by personal communication (Braasch,
<date id="8D7C6283FF99FFD88FF0F9CA7C766335" box="[151,302,1605,1631]" pageId="18" pageNumber="19" value="2004-09-15">15.IX.2004</date>
) about the misinterpretation of the species in question, of which he has given kindly acknowledgement (Bae,
<date id="8D7C6283FF99FFD88EB7F9E57F3863EE" box="[464,608,1642,1668]" pageId="18" pageNumber="19" value="2004-09-30">30.IX.2004</date>
personal communication). In honour of Dr. Van Vhin Nguyen as discoverer of the new species and Prof. Dr. Yoon Jae Bae as his adviser and co-author, we name the new species
<taxonomicName id="3EC23FC0FF99FFD88F94F93A7F4063A4" box="[243,536,1717,1742]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nguyenbaeorum" status="sp. nov.">
<emphasis id="CBB69851FF99FFD88F94F93A7F4063A4" box="[243,536,1717,1742]" italics="true" pageId="18" pageNumber="19">Epeorus nguyenbaeorum</emphasis>
</taxonomicName>
<emphasis id="CBB69851FF99FFD88D47F93A7F2563A5" bold="true" box="[544,637,1717,1743]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="D085252AFF99FFD88D47F93A7F2563A5" box="[544,637,1717,1743]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="B1D817C8FF99FFD88FA2F9557DA362E0" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="F97D4443FF99FFD88FA2F9557DA362E0" blockId="18.[151,1438,1530,2004]" pageId="18" pageNumber="19">
<emphasis id="CBB69851FF99FFD88FA2F9557C18639E" bold="true" box="[197,320,1754,1780]" pageId="18" pageNumber="19">Material:</emphasis>
<typeStatus id="2679FAE1FF99FFD88E2EF9557CB5639E" box="[329,493,1754,1780]" pageId="18" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
:
<emphasis id="CBB69851FF99FFD88E9EF9557FDB639E" bold="true" box="[505,643,1754,1780]" pageId="18" pageNumber="19">Female La</emphasis>
(Figs. 19–24),
<collectingCountry id="81D504D3FF99FFD88C25F9557EF4639E" box="[834,940,1754,1780]" name="Vietnam" pageId="18" pageNumber="19">Vietnam</collectingCountry>
, Ha Giang Prov., Vi Xuyen, Than Thuy,
<date id="8D7C6283FF99FFD88FF0F88F7C736270" box="[151,299,1792,1818]" pageId="18" pageNumber="19" value="2000-12-08">08.XII.2000</date>
, leg. Nguyen. Described by
<bibRefCitation id="9D5339B2FF99FFD88DE7F88F7ECB6270" author="Nguyen" box="[640,915,1792,1818]" pageId="18" pageNumber="19" refString="Nguyen, V. V. &amp; Bae, Y. J. (2004 a) Larvae of the Heptageniid Mayfly Genus Epeorus (Ephemeroptera: Heptageniidae) from Vietnam. Journal of Asia-Pacific Entomology, 7, 19 - 28." type="book chapter" year="2004" yearSuffix="a">Nguyen &amp; Bae (2004a</bibRefCitation>
: p. 19–28, Figs. 19–24: head and pronotum (Fig. 19), foreleg, dorsal (Fig. 20); abdomen, dorsal (Fig. 21), gill I (Fig. 22), gill III (Fig. 23), gill VII (Fig. 24).
<typeStatus id="2679FAE1FF99FFD88FABF8C57C2B620E" box="[204,371,1866,1892]" pageId="18" pageNumber="19" type="paratype">PARATYPES</typeStatus>
: 2 La, same locality; 2 La, Ha Giang Prov., Vi Xuyen, Tay Con Linh.
<date id="8D7C6283FF99FFD88BFCF8C57875620E" box="[1179,1325,1866,1892]" pageId="18" pageNumber="19" value="2000-12-09">09.XII.2000</date>
, leg. V.V. Nguyen.
</paragraph>
</subSubSection>
<subSubSection id="B1D817C8FF99FFD98FA2F81A7EBD674D" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="F97D4443FF99FFD98FA2F81A7EBD674D" blockId="18.[151,1438,1530,2004]" lastBlockId="19.[151,1437,152,663]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="CBB69851FF99FFD88FA2F81A7C1F62C5" bold="true" box="[197,327,1941,1967]" pageId="18" pageNumber="19">Diagnosis:</emphasis>
The head of
<taxonomicName id="3EC23FC0FF99FFD88E85F81A7FE662C4" box="[482,702,1941,1966]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nguyenbaeorum">
<emphasis id="CBB69851FF99FFD88E85F81A7FE662C4" box="[482,702,1941,1966]" italics="true" pageId="18" pageNumber="19">E. nguyenbaeorum</emphasis>
</taxonomicName>
with its quadrangular shape and conspicuous pattern contrasts strikingly with that of
<taxonomicName id="3EC23FC0FF99FFD88ECDF8347FDA62BE" box="[426,642,1978,2004]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hieroglyphicus">
<emphasis id="CBB69851FF99FFD88ECDF8347CE462BE" box="[426,444,1979,2004]" italics="true" pageId="18" pageNumber="19">E</emphasis>
.
<emphasis id="CBB69851FF99FFD88EABF8347FDA62BE" box="[460,642,1979,2004]" italics="true" pageId="18" pageNumber="19">hieroglyphicus</emphasis>
</taxonomicName>
, which has a rhomboid head with a central marking according to
<bibRefCitation id="9D5339B2FF98FFD98FF0FF177CB765D8" author="Braasch" box="[151,495,152,178]" pageId="19" pageNumber="20" refString="Braasch, D. &amp; Soldan, T. (1984 a) Eintagsfliegen Gattungen Epeorus und Iron) aus Vietnam (Ephemeroptera, Heptageniidae). In: Landa, V., Soldan, T. &amp; Tonner, M. (Eds), Proceedings of the Fourth International Conference Ephemeroptera. C zechoslovak Academy of Science, C eske Budejovice, pp. 109 - 114." type="book chapter" year="1984" yearSuffix="a">Braasch &amp; Soldán (1984a</bibRefCitation>
: Fig. 22, p.112); marks of the pronotum occupy the anterior half in
<taxonomicName id="3EC23FC0FF98FFD98AE6FF177C0F65BC" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hieroglyphicus">
<emphasis id="CBB69851FF98FFD98AE6FF177C0F65BC" italics="true" pageId="19" pageNumber="20">E. hieroglyphicus</emphasis>
</taxonomicName>
, whereas
<taxonomicName id="3EC23FC0FF98FFD98EBBFF327F8965BC" box="[476,721,189,214]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nguyenbaeorum">
<emphasis id="CBB69851FF98FFD98EBBFF327F8965BC" box="[476,721,189,214]" italics="true" pageId="19" pageNumber="20">E. nguyenbaeorum</emphasis>
</taxonomicName>
has only an anteromedian brown mark. Contrary to
<taxonomicName id="3EC23FC0FF98FFD98AE6FF327C156591" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hieroglyphicus">
<emphasis id="CBB69851FF98FFD98AE6FF327C156591" italics="true" pageId="19" pageNumber="20">E. hieroglyphicus</emphasis>
</taxonomicName>
, having a submedian row of oblique streaks along sterna, the new species is without any pattern. Microscopic structure of abdominal terga in
<taxonomicName id="3EC23FC0FF98FFD98C7DFE877EA6644B" box="[794,1022,264,289]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nguyenbaeorum">
<emphasis id="CBB69851FF98FFD98C7DFE877EA6644B" box="[794,1022,264,289]" italics="true" pageId="19" pageNumber="20">E. nguyenbaeorum</emphasis>
</taxonomicName>
is quite different in showing hind margin fringes with relatively long setae alternating with short acute bristles (
<bibRefCitation id="9D5339B2FF98FFD98B27FEA27813642D" author="Nguyen" box="[1088,1355,301,327]" pageId="19" pageNumber="20" refString="Nguyen, V. V. &amp; Bae, Y. J. (2004 a) Larvae of the Heptageniid Mayfly Genus Epeorus (Ephemeroptera: Heptageniidae) from Vietnam. Journal of Asia-Pacific Entomology, 7, 19 - 28." type="book chapter" year="2004" yearSuffix="a">Nguyen &amp; Bae 2004a</bibRefCitation>
: p. 24, Fig. 21), lacking in
<emphasis id="CBB69851FF98FFD98E18FEDD7F0D6401" box="[383,597,338,363]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="3EC23FC0FF98FFD98E18FEDD7F086401" box="[383,592,338,363]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hieroglyphicus">E. hieroglyphicus</taxonomicName>
,
</emphasis>
which has 2 submedian tubercles covered with 2 minute, spiny bristles and scattered with short hairs at the posterior margin of each tergum (
<bibRefCitation id="9D5339B2FF98FFD98CB0FEF7785064F8" author="Braasch" box="[983,1288,376,402]" pageId="19" pageNumber="20" refString="Braasch, D. &amp; Soldan, T. (1984 a) Eintagsfliegen Gattungen Epeorus und Iron) aus Vietnam (Ephemeroptera, Heptageniidae). In: Landa, V., Soldan, T. &amp; Tonner, M. (Eds), Proceedings of the Fourth International Conference Ephemeroptera. C zechoslovak Academy of Science, C eske Budejovice, pp. 109 - 114." type="book chapter" year="1984" yearSuffix="a">Braasch &amp; Soldán 1984a</bibRefCitation>
: p. 112, Fig. 27). The gill set offers distinct differences in gill I being egg-shaped (
<bibRefCitation id="9D5339B2FF98FFD98CA1FE12799364DD" author="Nguyen" box="[966,1227,413,439]" pageId="19" pageNumber="20" refString="Nguyen, V. V. &amp; Bae, Y. J. (2004 a) Larvae of the Heptageniid Mayfly Genus Epeorus (Ephemeroptera: Heptageniidae) from Vietnam. Journal of Asia-Pacific Entomology, 7, 19 - 28." type="book chapter" year="2004" yearSuffix="a">Nguyen &amp; Bae 2004a</bibRefCitation>
: p. 24, Fig. 22) in the new species, but pear-shaped in true
<taxonomicName id="3EC23FC0FF98FFD98D1DFE4D7E1164B1" box="[634,841,450,475]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hieroglyphicus">
<emphasis id="CBB69851FF98FFD98D1DFE4D7E1164B1" box="[634,841,450,475]" italics="true" pageId="19" pageNumber="20">E. hieroglyphicus</emphasis>
</taxonomicName>
(
<bibRefCitation id="9D5339B2FF98FFD98C3EFE4D79D064B6" author="Braasch" box="[857,1160,450,476]" pageId="19" pageNumber="20" refString="Braasch, D. &amp; Soldan, T. (1984 a) Eintagsfliegen Gattungen Epeorus und Iron) aus Vietnam (Ephemeroptera, Heptageniidae). In: Landa, V., Soldan, T. &amp; Tonner, M. (Eds), Proceedings of the Fourth International Conference Ephemeroptera. C zechoslovak Academy of Science, C eske Budejovice, pp. 109 - 114." type="book chapter" year="1984" yearSuffix="a">Braasch &amp; Soldán 1984a</bibRefCitation>
: p.112, Fig. 28), and in gill VII being obovate, whereas that of
<taxonomicName id="3EC23FC0FF98FFD98DF9FE677EDB676B" box="[670,899,488,513]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hieroglyphicus">
<emphasis id="CBB69851FF98FFD98DF9FE677EDB676B" box="[670,899,488,513]" italics="true" pageId="19" pageNumber="20">E. hieroglyphicus</emphasis>
</taxonomicName>
is somewhat oblong and distally with an asymmetrically pointed tip; gill III is large and elliptical in both species.
</paragraph>
</subSubSection>
<subSubSection id="B1D817C8FF98FFD98FA2FDBD7EAE67FD" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="F97D4443FF98FFD98FA2FDBD7EAE67FD" blockId="19.[151,1437,152,663]" pageId="19" pageNumber="20">
<emphasis id="CBB69851FF98FFD98FA2FDBD7CD36726" bold="true" box="[197,395,562,588]" pageId="19" pageNumber="20">
<typeStatus id="2679FAE1FF98FFD98FA2FDBD7C5A6726" box="[197,258,562,588]" pageId="19" pageNumber="20">Type</typeStatus>
deposition
</emphasis>
:
<typeStatus id="2679FAE1FF98FFD98EFCFDBD7F626726" box="[411,570,562,588]" pageId="19" pageNumber="20" type="holotype">HOLOTYPE</typeStatus>
:
<emphasis id="CBB69851FF98FFD98D22FDBD7E556726" bold="true" box="[581,781,562,588]" pageId="19" pageNumber="20">mature male La</emphasis>
(Figs. 19–24 see above), in Department of Invertebrate Zoology, Faculty of Biology, Hanoi University of Science,
<collectingCountry id="81D504D3FF98FFD98C36FDD77EE06718" box="[849,952,600,626]" name="Vietnam" pageId="19" pageNumber="20">Vietnam</collectingCountry>
;
<typeStatus id="2679FAE1FF98FFD98CA3FDD779336718" box="[964,1131,600,626]" pageId="19" pageNumber="20" type="paratype">PARATYPES</typeStatus>
: 1 La at the same place; 2 La in College of Life Science &amp; Biotechnology,
<collectingCountry id="81D504D3FF98FFD98DA8FDF27E4F67FD" box="[719,791,637,663]" name="South Korea" pageId="19" pageNumber="20">Korea</collectingCountry>
University,
<collectingCountry id="81D504D3FF98FFD98CC0FDF27EAA67FD" box="[935,1010,637,663]" name="South Korea" pageId="19" pageNumber="20">Korea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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