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<document id="10F16F1D8AE90857B697215A2E6E43FB" ID-DOI="10.11646/zootaxa.3856.2.2" ID-GBIF-Dataset="24c01b66-c0d2-458b-a7fe-0e3de779a8e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="250932" ID-ZooBank="ABC0E7D7-2E22-4605-A97B-0F1E632F3930" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460889728808" checkinUser="plazi" docAuthor="Bota, Cornelio Andrés" docDate="2014" docId="4E56376DFFABFF9646CB62642B6FFBBD" docLanguage="en" docName="zt03856p210.pdf" docOrigin="Zootaxa 3856 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhionaeschna joannisi Martin 1897" docType="treatment" docVersion="8" lastPageNumber="200" masterDocId="B26F4F15FFA3FF9E465C607B2E0DFFB6" masterDocTitle="A brief look at the Odonata from the Páramo ecosystems in Colombia, with the descriptions of Oxyallagma colombianum sp. nov. and Rhionaeschna caligo sp. nov. (Odonata: Coenagrionidae, Aeshnidae, Libellulidae)" masterLastPageNumber="210" masterPageNumber="192" pageNumber="200" updateTime="1698359463704" updateUser="plazi">
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<mods:title id="9D633FD6EAA2B62ACEC086F430478851">A brief look at the Odonata from the Páramo ecosystems in Colombia, with the descriptions of Oxyallagma colombianum sp. nov. and Rhionaeschna caligo sp. nov. (Odonata: Coenagrionidae, Aeshnidae, Libellulidae)</mods:title>
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<mods:namePart id="9FF44201BE3464E11933B73C8B7FA54F">Bota, Cornelio Andrés</mods:namePart>
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<mods:date id="2B113E37AF3B9D26DC8195B3E16C9296">2014</mods:date>
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<treatment id="4E56376DFFABFF9646CB62642B6FFBBD" ID-DOI="http://doi.org/10.5281/zenodo.6141391" ID-GBIF-Taxon="119586696" ID-Zenodo-Dep="6141391" LSID="urn:lsid:plazi:treatment:4E56376DFFABFF9646CB62642B6FFBBD" httpUri="http://treatment.plazi.org/id/4E56376DFFABFF9646CB62642B6FFBBD" lastPageNumber="200" pageId="8" pageNumber="200">
<subSubSection id="8EE5D5F0FFABFF9646CB62642C7CFDED" pageId="8" pageNumber="200" type="nomenclature">
<paragraph id="C640867BFFABFF9646CB62642C66FD8C" blockId="8.[151,625,543,603]" box="[151,619,543,570]" pageId="8" pageNumber="200">
<heading id="9D083117FFABFF9646CB62642C66FD8C" bold="true" box="[151,619,543,570]" fontSize="11" level="1" pageId="8" pageNumber="200" reason="1">
<taxonomicName id="01FFFDF8FFABFF9646CB62642C66FD8C" ID-CoL="78TG4" authority="Martin, 1897" authorityName="Martin" authorityYear="1897" box="[151,619,543,570]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="200" phylum="Arthropoda" rank="species" species="joannisi">
<emphasis id="F48B5A69FFABFF9646CB62642C66FD8C" bold="true" box="[151,619,543,570]" pageId="8" pageNumber="200">
<emphasis id="F48B5A69FFABFF9646CB62642FA3FD8F" bold="true" box="[151,430,543,569]" italics="true" pageId="8" pageNumber="200">Rhionaeschna joannisi</emphasis>
(Martin, 1897)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C640867BFFABFF9646CB62392C7CFDED" blockId="8.[151,625,543,603]" box="[151,625,578,603]" pageId="8" pageNumber="200">
<figureCitation id="5EC49AFEFFABFF9646CB62392EDAFDED" box="[151,215,578,603]" captionStart="FIGURE 2" captionStartId="10.[151,250,1953,1975]" captionTargetBox="[157,1433,199,1931]" captionTargetId="figure@10.[151,1436,193,1932]" captionTargetPageId="10" captionText="FIGURE 2. (a) Rhionaeschna peralta female (habitus). Rhionaeschna caligo: (b) Hamuli, ventroposterior view; (c) Caudal appendages lateral view; (d) Vesica spermalis ventral view; (e) left spine of anterior lamina. (f) Rhionaeschna joannisi male (habitus). Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250934/files/figure.png" pageId="8" pageNumber="200">Fig. 2</figureCitation>
f (Habitus ♂), Table 1 (coordinates).
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFABFF9646CB62F12ACCFD15" box="[151,1217,650,675]" pageId="8" pageNumber="200" type="materials_examined">
<paragraph id="C640867BFFABFF9646CB62F12ACCFD15" blockId="8.[151,1436,650,1035]" box="[151,1217,650,675]" pageId="8" pageNumber="200">
<emphasis id="F48B5A69FFABFF9646CB62F12FE5FD15" bold="true" box="[151,488,650,675]" pageId="8" pageNumber="200">
Examined
<taxonomicName id="01FFFDF8FFABFF96474B62F12F79FD15" box="[279,372,650,675]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="200" phylum="Arthropoda" rank="species" species="paramo">páramo</taxonomicName>
material.
</emphasis>
<specimenCount id="D0F94DF2FFABFF9647B262F02C14FD14" box="[494,537,651,675]" pageId="8" pageNumber="200" type="generic">1 ♂</specimenCount>
and 1♀:
<taxonomicName id="01FFFDF8FFABFF9644DC62F02CD8FD15" box="[640,725,651,675]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="210" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
Las Baldías,
<date id="B241A0BBFFABFF96453362F12DE4FD15" box="[879,1001,650,675]" pageId="8" pageNumber="200" value="2002-06-15">15.vi.2002</date>
, Leg: J. A. Posada.
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFABFF96469B62D52DD3FCE1" pageId="8" pageNumber="200" type="discussion">
<paragraph id="C640867BFFABFF96469B62D52DD3FCE1" blockId="8.[151,1436,650,1035]" pageId="8" pageNumber="200">
<emphasis id="F48B5A69FFABFF96469B62D52F36FD71" bold="true" box="[199,315,686,711]" pageId="8" pageNumber="200">Remarks.</emphasis>
In the male, and in two additional males collected outside the
<taxonomicName id="01FFFDF8FFABFF96427062D42A8DFD71" box="[1068,1152,687,711]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="200" phylum="Arthropoda" rank="species" species="paramo">páramo</taxonomicName>
ecosystem, the epiproct reaches to more than half of cercus length, while in the illustration of male S10 provided by von
<bibRefCitation id="A26EFB8AFFABFF96429262A92B91FD5D" author="Ellenrieder" box="[1230,1436,722,747]" pageId="8" pageNumber="200" refString="von Ellenrieder, N. (2003) A synopsis of the neotropical species of &quot; Aeshna &quot; Fabricius: Genus Rhionaeschna Forster. Tijdschrift voor Entomologie, 146, 67 - 207. http: // dx. doi. org / 10.1163 / 22119434 - 900000120" type="journal article" year="2003">Ellenrieder (2003)</bibRefCitation>
of a specimen of
<taxonomicName id="01FFFDF8FFABFF964707628C2FD8FCB8" box="[347,469,759,782]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="200" phylum="Arthropoda" rank="species" species="joannisi">
<emphasis id="F48B5A69FFABFF964707628C2FD8FCB8" box="[347,469,759,782]" italics="true" pageId="8" pageNumber="200">R. joannisi</emphasis>
</taxonomicName>
from
<collectingCountry id="BEE8C6EBFFABFF964447628D2C75FCB9" box="[539,632,758,783]" name="Ecuador" pageId="8" pageNumber="200">Ecuador</collectingCountry>
the epiproct is about as long as half of cercus length. In all Colombian specimens there is a metepisternal stripe (
<figureCitation id="5EC49AFEFFABFF9644D763612CD9FC85" box="[651,724,794,819]" captionStart="FIGURE 2" captionStartId="10.[151,250,1953,1975]" captionTargetBox="[157,1433,199,1931]" captionTargetId="figure@10.[151,1436,193,1932]" captionTargetPageId="10" captionText="FIGURE 2. (a) Rhionaeschna peralta female (habitus). Rhionaeschna caligo: (b) Hamuli, ventroposterior view; (c) Caudal appendages lateral view; (d) Vesica spermalis ventral view; (e) left spine of anterior lamina. (f) Rhionaeschna joannisi male (habitus). Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250934/files/figure.png" pageId="8" pageNumber="200">Fig. 2</figureCitation>
f), which is incomplete or broken into separate spots in the specimens from
<collectingCountry id="BEE8C6EBFFABFF96470C63452FA0FCE1" box="[336,429,830,855]" name="Ecuador" pageId="8" pageNumber="200">Ecuador</collectingCountry>
and
<collectingCountry id="BEE8C6EBFFABFF9647B863452C3AFCE1" box="[484,567,830,855]" name="Bolivia" pageId="8" pageNumber="200">Bolivia</collectingCountry>
illustrated by von
<bibRefCitation id="A26EFB8AFFABFF96455463452DD4FCE1" author="Ellenrieder" box="[776,985,830,855]" pageId="8" pageNumber="200" refString="von Ellenrieder, N. (2003) A synopsis of the neotropical species of &quot; Aeshna &quot; Fabricius: Genus Rhionaeschna Forster. Tijdschrift voor Entomologie, 146, 67 - 207. http: // dx. doi. org / 10.1163 / 22119434 - 900000120" type="journal article" year="2003">Ellenrieder (2003)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFABFF96469B63192A50FC75" pageId="8" pageNumber="200" type="distribution">
<paragraph id="C640867BFFABFF96469B63192A50FC75" blockId="8.[151,1436,650,1035]" pageId="8" pageNumber="200">
<emphasis id="F48B5A69FFABFF96469B63192F6DFCCD" bold="true" box="[199,352,866,891]" pageId="8" pageNumber="200">Distribution.</emphasis>
From
<collectingCountry id="BEE8C6EBFFABFF9647F663192FF0FCCD" box="[426,509,866,891]" name="Bolivia" pageId="8" pageNumber="200">Bolivia</collectingCountry>
to
<collectingCountry id="BEE8C6EBFFABFF96447C63192C83FCCD" box="[544,654,866,891]" name="Colombia" pageId="8" pageNumber="200">Colombia</collectingCountry>
(von
<bibRefCitation id="A26EFB8AFFABFF96448C63192D80FCCD" author="Ellenrieder" box="[720,909,866,891]" pageId="8" pageNumber="200" refString="von Ellenrieder, N. (2003) A synopsis of the neotropical species of &quot; Aeshna &quot; Fabricius: Genus Rhionaeschna Forster. Tijdschrift voor Entomologie, 146, 67 - 207. http: // dx. doi. org / 10.1163 / 22119434 - 900000120" type="journal article" year="2003">Ellenrieder 2003</bibRefCitation>
) across the Andean region from
<quantity id="01072B9EFFABFF9642A763182EBDFC28" metricMagnitude="0" metricUnit="m" metricValue="2.115" metricValueMax="3.13" metricValueMin="1.1" pageId="8" pageNumber="200" unit="m" value="2.115" valueMax="3.13" valueMin="1.1">1,100 to 3,130 m</quantity>
. These are the first records from
<collectingCountry id="BEE8C6EBFFABFF96447763FD2C94FC29" box="[555,665,902,927]" name="Colombia" pageId="8" pageNumber="200">Colombia</collectingCountry>
since Needhams record from Popayán in 1912 (
<bibRefCitation id="A26EFB8AFFABFF96429A63FD2B52FC29" author="Calvert" box="[1222,1375,902,927]" pageId="8" pageNumber="200" refString="Calvert, P. P. (1956) The Neotropical species of the ' subgenus Aeshna ' sensu Selysii 1883 (Odonata). Memoirs of the American Entomological Society, 15, 1 - 251." type="journal article" year="1956">Calvert 1956</bibRefCitation>
), and the known distributional range is expanded by four degrees north from 2°40 to 6°19.
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFABFF96469B63B52B6FFBBD" pageId="8" pageNumber="200" type="biology_ecology">
<paragraph id="C640867BFFABFF96469B63B52B6FFBBD" blockId="8.[151,1436,650,1035]" pageId="8" pageNumber="200">
<emphasis id="F48B5A69FFABFF96469B63B52F2AFC51" bold="true" box="[199,295,974,999]" pageId="8" pageNumber="200">Biology.</emphasis>
Despite being widely distributed, the species seems to be rare. I have never seen a live specimen. It inhabits small ponds in the
<taxonomicName id="01FFFDF8FFABFF96479663882C13FBBD" box="[458,542,1011,1035]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="200" phylum="Arthropoda" rank="species" species="paramo">páramo</taxonomicName>
and high mountain forests of the northern Central Cordillera in
<collectingCountry id="BEE8C6EBFFABFF9642B063892B52FBBD" box="[1260,1375,1010,1035]" name="Colombia" pageId="8" pageNumber="200">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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