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<document id="69BB920B3321CAC3EC207DC9DB32BBFB" ID-DOI="10.11646/zootaxa.3856.2.2" ID-GBIF-Dataset="24c01b66-c0d2-458b-a7fe-0e3de779a8e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="250932" ID-ZooBank="ABC0E7D7-2E22-4605-A97B-0F1E632F3930" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460889728808" checkinUser="plazi" docAuthor="Bota, Cornelio Andrés" docDate="2014" docId="4E56376DFFA6FF9B46CB62AE2B96FB13" docLanguage="en" docName="zt03856p210.pdf" docOrigin="Zootaxa 3856 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxyallagma" docType="key" docVersion="8" lastPageNumber="197" masterDocId="B26F4F15FFA3FF9E465C607B2E0DFFB6" masterDocTitle="A brief look at the Odonata from the Páramo ecosystems in Colombia, with the descriptions of Oxyallagma colombianum sp. nov. and Rhionaeschna caligo sp. nov. (Odonata: Coenagrionidae, Aeshnidae, Libellulidae)" masterLastPageNumber="210" masterPageNumber="192" pageNumber="197" updateTime="1698359463704" updateUser="plazi">
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<mods:title id="598A7DB2DDE2CB43BE286905D0238A29">A brief look at the Odonata from the Páramo ecosystems in Colombia, with the descriptions of Oxyallagma colombianum sp. nov. and Rhionaeschna caligo sp. nov. (Odonata: Coenagrionidae, Aeshnidae, Libellulidae)</mods:title>
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<mods:namePart id="4AEC3F8911BC4E3E8434D18A70B97120">Bota, Cornelio Andrés</mods:namePart>
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<mods:date id="D703FE99503DBD8C245E74FE05B5B635">2014</mods:date>
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<treatment id="4E56376DFFA6FF9B46CB62AE2B96FB13" ID-DOI="http://doi.org/10.5281/zenodo.6141375" ID-GBIF-Taxon="119586679" ID-Zenodo-Dep="6141375" LSID="urn:lsid:plazi:treatment:4E56376DFFA6FF9B46CB62AE2B96FB13" httpUri="http://treatment.plazi.org/id/4E56376DFFA6FF9B46CB62AE2B96FB13" lastPageNumber="197" pageId="5" pageNumber="197">
<subSubSection id="8EE5D5F0FFA6FF9B46CB62AE2F3DFCEE" pageId="5" pageNumber="197" type="nomenclature">
<paragraph id="C640867BFFA6FF9B46CB62AE2C0DFD59" blockId="5.[151,512,725,751]" box="[151,512,725,751]" pageId="5" pageNumber="197">
<heading id="9D083117FFA6FF9B46CB62AE2C0DFD59" bold="true" box="[151,512,725,751]" fontSize="11" level="1" pageId="5" pageNumber="197" reason="1">
<emphasis id="F48B5A69FFA6FF9B46CB62AE2C0DFD59" bold="true" box="[151,512,725,751]" pageId="5" pageNumber="197">
Key to species of
<taxonomicName id="01FFFDF8FFA6FF9B473162AE2C0DFD59" ID-CoL="6B83" box="[365,512,725,751]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="F48B5A69FFA6FF9B473162AE2C0DFD59" bold="true" box="[365,512,725,751]" italics="true" pageId="5" pageNumber="197">Oxyallagma</emphasis>
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</paragraph>
<paragraph id="C640867BFFA6FF9B46CB63672F3DFCEE" blockId="5.[151,1436,796,856]" pageId="5" pageNumber="197">
Reference to the illustrations of
<taxonomicName id="01FFFDF8FFA6FF9B445B63672C9FFC82" box="[519,658,796,820]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="197" phylum="Arthropoda" rank="species" species="dissidens">
<emphasis id="F48B5A69FFA6FF9B445B63672C9FFC82" box="[519,658,796,820]" italics="true" pageId="5" pageNumber="197">O. dissidens</emphasis>
</taxonomicName>
in
<bibRefCitation id="A26EFB8AFFA6FF9B44E463672DBDFC82" box="[696,944,796,821]" pageId="5" pageNumber="197" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World: An Illustrated and Annotated Key to the Zygoptera. The Johns Hopkins University Press, Baltimore, 528 pp." type="book">
Garrison
<emphasis id="F48B5A69FFA6FF9B457F63662D51FC82" box="[803,860,796,820]" italics="true" pageId="5" pageNumber="197">et al.</emphasis>
(2010)
</bibRefCitation>
is important in order to accurately identify these species.
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFA6FF9B46CB63FC2B96FB13" pageId="5" pageNumber="197" type="key">
<keyStep id="7D0B9EDEFFA6FF9B46CB63FC2B96FC0C" pageId="5" pageNumber="197">
<paragraph id="C640867BFFA6FF9B46CB63FC2B96FC2A" blockId="5.[151,1436,903,1189]" box="[151,1435,903,924]" pageId="5" pageNumber="197">
<keyLead id="7D0E254EFFA6FF9B46CB63FC2B96FC2A" box="[151,1435,903,924]" pageId="5" pageNumber="197">1. Females............................................................................................ 2</keyLead>
</paragraph>
<paragraph id="C640867BFFA6FF9B46CB63DE2B96FC0C" blockId="5.[151,1436,903,1189]" box="[151,1435,933,954]" pageId="5" pageNumber="197">
<keyLead id="7D0E254EFFA6FF9B46CB63DE2B96FC0C" box="[151,1435,933,954]" pageId="5" pageNumber="197">1. Males............................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="7D0B9EDEFFA6FF9B46CB63B92B96FB99" pageId="5" pageNumber="197">
<paragraph id="C640867BFFA6FF9B46CB63B92B91FC43" blockId="5.[151,1436,903,1189]" pageId="5" pageNumber="197">
<keyLead id="7D0E254EFFA6FF9B46CB63B92B91FC43" pageId="5" pageNumber="197">
2 (1). S2
<emphasis id="F48B5A69FFA6FF9B46A663B92F0BFC61" box="[250,262,962,983]" italics="true" pageId="5" pageNumber="197"></emphasis>
6 with middorsal black stripe, S9 with light blue middorsal stripe (
<figureCitation id="5EC49AFEFFA6FF9B45CB63B92DDEFC61" box="[919,979,962,983]" captionStart="FIGURE 4" captionStartId="13.[151,250,1954,1976]" captionTargetBox="[202,1384,193,1931]" captionTargetId="figure@13.[202,1385,193,1932]" captionTargetPageId="13" captionText="FIGURE 4. (a) Ischnura ramburii female (Páramo El Morro). (b) Oxyallagma colombianum, teneral male (Antioquia, Labores). Oxyallagma dissidens: (c) Female (Nariño, La Cocha); (d) Male (Cauca, Silvia)." httpUri="https://zenodo.org/record/250936/files/figure.png" pageId="5" pageNumber="197">Fig. 4</figureCitation>
c); cercus ivory white; middorsum of posterior lobe of pronotum strongly bent dorsally (Northern
<collectingCountry id="BEE8C6EBFFA6FF9B449463A42CFBFC42" box="[712,758,991,1012]" name="Peru" pageId="5" pageNumber="197">Peru</collectingCountry>
,
<collectingCountry id="BEE8C6EBFFA6FF9B455C63A42D5CFC42" box="[768,849,991,1012]" name="Ecuador" pageId="5" pageNumber="197">Ecuador</collectingCountry>
, and Southern
<collectingCountry id="BEE8C6EBFFA6FF9B45BC63A42A4DFC42" box="[992,1088,991,1012]" name="Colombia" pageId="5" pageNumber="197">Colombia</collectingCountry>
)..................
<taxonomicName id="01FFFDF8FFA6FF9B437A63A42B91FC43" ID-CoL="4BCCY" box="[1318,1436,991,1013]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="197" phylum="Arthropoda" rank="species" species="dissidens">
<emphasis id="F48B5A69FFA6FF9B437A63A42B91FC43" box="[1318,1436,991,1013]" italics="true" pageId="5" pageNumber="197">O. dissidens</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C640867BFFA6FF9B46CB63862B96FB99" blockId="5.[151,1436,903,1189]" pageId="5" pageNumber="197">
<keyLead id="7D0E254EFFA6FF9B46CB63862B96FB99" pageId="5" pageNumber="197">
2. No middorsal black stripe on S2
<emphasis id="F48B5A69FFA6FF9B444463862C29FBA4" box="[536,548,1021,1042]" italics="true" pageId="5" pageNumber="197"></emphasis>
6, nor light blue middorsal stripe on S9 (
<figureCitation id="5EC49AFEFFA6FF9B45F663862DE0FBA4" box="[938,1005,1021,1042]" captionStart="FIGURE 5" captionStartId="14.[151,250,1953,1975]" captionTargetBox="[202,1383,193,1930]" captionTargetId="figure@14.[202,1384,193,1932]" captionTargetPageId="14" captionText="FIGURE 5. Oxyallagma colombianum: (Páramo Sabanas): (a) Pair in wheel position; (b) Tandem pair; (Antioquia, Labores): (c) Ovipositing pair with microparasitoid close to the oviposition place; (d) Tandem pair (photograph by Cintia Moreno)." httpUri="https://zenodo.org/record/250937/files/figure.png" pageId="5" pageNumber="197">Figs. 5</figureCitation>
b, d); cercus dark brown; middorsum of pos- terior lobe of pronotum only slightly bent dorsally (
<figureCitation id="5EC49AFEFFA6FF9B449364612D05FB99" box="[719,776,1050,1071]" captionStart="FIGURE 6" captionStartId="15.[151,250,1831,1853]" captionTargetBox="[233,1352,206,1809]" captionTargetId="figure@15.[226,1360,193,1810]" captionTargetPageId="15" captionText="FIGURE 6. Oxyallagma colombianum (Páramo Sabanas): Female paratype: (a) Dorsal view of pronotum posterior lobe and mesostigmal plates. Male paratype (Páramo Las Baldías): (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio - dorsal view of caudal appendages. l-m. l.: Latero-medial lobes. m. p.: Mesostigmal plates. p. p. l.: Pronotum posterior lobe. e. v ap. p.: Ventroapical external process. v b t.: Ventrobasal tooth. i. d. p.: Inner dorsal process. p. d. b.: Paraproct dorsal branch. p. v. b.: Paraproct ventral branch. Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250938/files/figure.png" pageId="5" pageNumber="197">Fig. 6</figureCitation>
a) (Northern
<collectingCountry id="BEE8C6EBFFA6FF9B45D964612DE8FB99" box="[901,997,1050,1071]" name="Colombia" pageId="5" pageNumber="197">Colombia</collectingCountry>
)......................
<taxonomicName id="01FFFDF8FFA6FF9B42A064622B96FB99" ID-CoL="9X3X4" box="[1276,1435,1049,1071]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="197" phylum="Arthropoda" rank="species" species="colombianum">
<emphasis id="F48B5A69FFA6FF9B42A064622B96FB99" box="[1276,1435,1049,1071]" italics="true" pageId="5" pageNumber="197">O. colombianum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7D0B9EDEFFA6FF9B46CB644C2B96FB13" pageId="5" pageNumber="197">
<paragraph id="C640867BFFA6FF9B46CB644C2B91FBDC" blockId="5.[151,1436,903,1189]" pageId="5" pageNumber="197">
<keyLead id="7D0E254EFFA6FF9B46CB644C2B91FBDC" pageId="5" pageNumber="197">
3 (1). Genital ligula without small latero-medial pointed lobes; paraproct with swollen semicircular ventral branch as wide as cercus; cercus with two short ventroapical processes (Northern
<collectingCountry id="BEE8C6EBFFA6FF9B44AB642E2D28FBDC" box="[759,805,1109,1130]" name="Peru" pageId="5" pageNumber="197">Peru</collectingCountry>
,
<collectingCountry id="BEE8C6EBFFA6FF9B4573642E2D8CFBDC" box="[815,897,1109,1130]" name="Ecuador" pageId="5" pageNumber="197">Ecuador</collectingCountry>
, and Southern
<collectingCountry id="BEE8C6EBFFA6FF9B4253642E2A62FBDC" box="[1039,1135,1109,1130]" name="Colombia" pageId="5" pageNumber="197">Colombia</collectingCountry>
)..............
<taxonomicName id="01FFFDF8FFA6FF9B437A642F2B91FBDC" ID-CoL="4BCCY" box="[1318,1436,1108,1130]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="197" phylum="Arthropoda" rank="species" species="dissidens">
<emphasis id="F48B5A69FFA6FF9B437A642F2B91FBDC" box="[1318,1436,1108,1130]" italics="true" pageId="5" pageNumber="197">O. dissidens</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C640867BFFA6FF9B46CB64092B96FB13" blockId="5.[151,1436,903,1189]" pageId="5" pageNumber="197">
<keyLead id="7D0E254EFFA6FF9B46CB64092B96FB13" pageId="5" pageNumber="197">
3. Genital ligula with small latero-medial pointed lobes (
<figureCitation id="5EC49AFEFFA6FF9B44B164092D25FB31" box="[749,808,1138,1159]" captionStart="FIGURE 6" captionStartId="15.[151,250,1831,1853]" captionTargetBox="[233,1352,206,1809]" captionTargetId="figure@15.[226,1360,193,1810]" captionTargetPageId="15" captionText="FIGURE 6. Oxyallagma colombianum (Páramo Sabanas): Female paratype: (a) Dorsal view of pronotum posterior lobe and mesostigmal plates. Male paratype (Páramo Las Baldías): (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio - dorsal view of caudal appendages. l-m. l.: Latero-medial lobes. m. p.: Mesostigmal plates. p. p. l.: Pronotum posterior lobe. e. v ap. p.: Ventroapical external process. v b t.: Ventrobasal tooth. i. d. p.: Inner dorsal process. p. d. b.: Paraproct dorsal branch. p. v. b.: Paraproct ventral branch. Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250938/files/figure.png" pageId="5" pageNumber="197">Fig. 6</figureCitation>
b); paraproct with ventral branch not swollen, narrower than half of cercus (
<figureCitation id="5EC49AFEFFA6FF9B471964F42F8AFB12" box="[325,391,1167,1188]" captionStart="FIGURE 6" captionStartId="15.[151,250,1831,1853]" captionTargetBox="[233,1352,206,1809]" captionTargetId="figure@15.[226,1360,193,1810]" captionTargetPageId="15" captionText="FIGURE 6. Oxyallagma colombianum (Páramo Sabanas): Female paratype: (a) Dorsal view of pronotum posterior lobe and mesostigmal plates. Male paratype (Páramo Las Baldías): (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio - dorsal view of caudal appendages. l-m. l.: Latero-medial lobes. m. p.: Mesostigmal plates. p. p. l.: Pronotum posterior lobe. e. v ap. p.: Ventroapical external process. v b t.: Ventrobasal tooth. i. d. p.: Inner dorsal process. p. d. b.: Paraproct dorsal branch. p. v. b.: Paraproct ventral branch. Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250938/files/figure.png" pageId="5" pageNumber="197">Figs. 6</figureCitation>
c
<emphasis id="F48B5A69FFA6FF9B47CE64EB2F93FB13" box="[402,414,1168,1189]" italics="true" pageId="5" pageNumber="197"></emphasis>
d); cercus with one ventroapical process (
<figureCitation id="5EC49AFEFFA6FF9B457064F42D68FB12" box="[812,869,1167,1188]" captionStart="FIGURE 6" captionStartId="15.[151,250,1831,1853]" captionTargetBox="[233,1352,206,1809]" captionTargetId="figure@15.[226,1360,193,1810]" captionTargetPageId="15" captionText="FIGURE 6. Oxyallagma colombianum (Páramo Sabanas): Female paratype: (a) Dorsal view of pronotum posterior lobe and mesostigmal plates. Male paratype (Páramo Las Baldías): (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio - dorsal view of caudal appendages. l-m. l.: Latero-medial lobes. m. p.: Mesostigmal plates. p. p. l.: Pronotum posterior lobe. e. v ap. p.: Ventroapical external process. v b t.: Ventrobasal tooth. i. d. p.: Inner dorsal process. p. d. b.: Paraproct dorsal branch. p. v. b.: Paraproct ventral branch. Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250938/files/figure.png" pageId="5" pageNumber="197">Fig. 6</figureCitation>
d) (Northern
<collectingCountry id="BEE8C6EBFFA6FF9B45BF64F42A4EFB12" box="[995,1091,1167,1188]" name="Colombia" pageId="5" pageNumber="197">Colombia</collectingCountry>
)..............
<taxonomicName id="01FFFDF8FFA6FF9B42A064F42B96FB13" ID-CoL="9X3X4" box="[1276,1435,1167,1189]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="197" phylum="Arthropoda" rank="species" species="colombianum">
<emphasis id="F48B5A69FFA6FF9B42A064F42B96FB13" box="[1276,1435,1167,1189]" italics="true" pageId="5" pageNumber="197">O. colombianum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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