137 lines
11 KiB
XML
137 lines
11 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.285.4502" ID-PMC="PMC3690755" ID-Pensoft-Pub="1313-2970-285-1" ID-Pensoft-UUID="FFCFFFB91706FF856810FFA7FFD2FFCE" ID-PubMed="23798894" ID-Zenodo-Dep="578334" ModsDocID="1313-2970-285-1" checkinTime="1451247459543" checkinUser="pensoft" docAuthor="Blakemore, Robert J." docDate="2013" docId="3EF2D7D90F0152B196FF57DCEBD59755" docLanguage="en" docName="ZooKeys 285: 1-21" docOrigin="ZooKeys 285" docPubDate="2013-04-05" docSource="http://dx.doi.org/10.3897/zookeys.285.4502" docTitle="Eisenia nordenskioldi subsp. onon Blakemore 2013, ssp. n." docType="treatment" docVersion="3" id="FFCFFFB91706FF856810FFA7FFD2FFCE" lastPageNumber="6" masterDocId="FFCFFFB91706FF856810FFA7FFD2FFCE" masterDocTitle="Earthworms newly from Mongolia (Oligochaeta, Lumbricidae, Eisenia)" masterLastPageNumber="21" masterPageNumber="1" pageNumber="6" updateTime="1668155597120" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Earthworms newly from Mongolia (Oligochaeta, Lumbricidae, Eisenia)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Blakemore, Robert J.</mods:namePart>
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<mods:affiliation>National Institute of Biological Resources (NIBR), Incheon, 404 - 708, Korea</mods:affiliation>
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<mods:nameIdentifier type="email">rob.blakemore@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-04-05</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>285</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>21</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.285.4502</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.285.4502</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-285-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFCFFFB91706FF856810FFA7FFD2FFCE</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578334</mods:identifier>
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</mods:mods>
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<subSection pageId="5" pageNumber="6" type="multiple">
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<treatment ID-GBIF-Taxon="182235593" LSID="urn:lsid:plazi:treatment:3EF2D7D90F0152B196FF57DCEBD59755" httpUri="http://treatment.plazi.org/id/3EF2D7D90F0152B196FF57DCEBD59755" lastPageNumber="6" pageId="5" pageNumber="6">
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<subSubSection pageId="5" pageNumber="6" type="nomenclature">
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<paragraph pageId="5" pageNumber="6">
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<taxonomicName LSID="3EF2D7D9-0F01-52B1-96FF-57DCEBD59755" authority="Blakemore, 2013" authorityName="Blakemore" authorityYear="2013" class="Clitellata" family="Lumbricidae" genus="Eisenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Eisenia nordenskioldi subsp. onon" order="Crassiclitellata" pageId="5" pageNumber="6" phylum="Annelida" rank="subSpecies" species="nordenskioldi" status="ssp. n." subSpecies="onon">Eisenia nordenskioldi onon</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="6">ssp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A Eisenia nordenskioldi onon ssp. n. Holotype sketched as for Figs 1 - 2 B Allolobophora harbinensis Kobayashi, 1940: fig. 6 (incertae sedis) for comparison." httpUri="https://binary.pensoft.net/fig/29791" pageId="5" pageNumber="6">Fig. 3</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="distribution">
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<paragraph pageId="5" pageNumber="6">Material examined.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Holotype (H) NIBR IV0000261277 (mature, dissected, providing DNA sample - wo65) plus six sub-adults provisionally listed as paratypes (IV000061278) and a
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<normalizedToken originalValue="‘tail’">'tail'</normalizedToken>
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, all poorly-preserved in same batch from "
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<emphasis italics="true" pageId="5" pageNumber="6">2012-7-20 Dadal</emphasis>
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". Unidentifiable were ca. 20 specimens (IV0000261279) some having clitella ca. 24-33 and TP ca. 28-30, also poorly-preserved from crowding in a single tube, labelled "
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<emphasis italics="true" pageId="5" pageNumber="6">2012-7-21 Dadal</emphasis>
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". All specimens collected by
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<normalizedToken originalValue="NIBR’s">NIBR's</normalizedToken>
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Mr T.-S. Park.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="etymology">
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<paragraph pageId="5" pageNumber="6">Etymology.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Nominative noun in apposition after sample region where Dadal and the upper Onon River are supposed birthplace and likely final resting place of
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<normalizedToken originalValue="Temüjin">Temuejin</normalizedToken>
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(otherwise known as Genghis Khan).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="description">
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<paragraph pageId="5" pageNumber="6">Description.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Body medium sized, H 100 mm. Segments 170. Reddish pink anterior-dorsum to segment 15 otherwise unpigmented. Epilobous. Pale laterally around
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<emphasis italics="true" pageId="5" pageNumber="6">cd</emphasis>
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in 8-11 and slightly tumid
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<emphasis italics="true" pageId="5" pageNumber="6">ab</emphasis>
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on 11-12 and possibly somewhat on clitellum. First dorsal pore 4/5. Spermathecal pores in 9/10/11 mid-dorsally. Female and male pores slight, lateral of
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<emphasis italics="true" pageId="5" pageNumber="6">b</emphasis>
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setae on 14 and 15, respectively. Nephridia sporadically visible lateral of
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<emphasis italics="true" pageId="5" pageNumber="6">b</emphasis>
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lines near intersegments (at least on clitellum) otherwise near
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<emphasis italics="true" pageId="5" pageNumber="6">d</emphasis>
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lines? Clitellum, pale from 24 dorsally or laterally 25-33, i.e., 24,25-33. TP longitudinally lenticular lateral of
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<emphasis italics="true" pageId="5" pageNumber="6">b</emphasis>
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28-31. External features rather unclear due to poor preservation.
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</paragraph>
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<paragraph pageId="5" pageNumber="6">Internally similar to nominal subspecies. Seminal vesicles in 9-12. Testis iridescent, free in 10 & 11. Calciferous glands in 11 & 12, vascularized and extending slightly into adjacent segments. Nephridial bladders sausage-shaped. Gizzard 17-18 and thin inverted T-shaped typhlosole present. Soil with coarse organic debris in gut. No parasites were noted.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="remarks">
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<paragraph pageId="5" pageNumber="6">Remarks.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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The current taxon differs from previously described subspecies (
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Characters of Eisenia nordenskioldi sub-species after Kobayashi (1940), Perel' (1969), Zicsi (1972) and pers. obs (cf. other sub-species in Appendix 2)." httpUri="http://table.plazi.org/id/FD4AC38FD47DCE1B365EAED9DEC2ACFE" pageId="5" pageNumber="6" tableUuid="FD4AC38FD47DCE1B365EAED9DEC2ACFE">Tab. 1</tableCitation>
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and Appendix 2) on its clitellum, TP and tumescences; moreover it appears fertile. Fresher and better preserved material should confirm this analysis. In the meantime, although physically closest to
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<taxonomicName authorityName="Blakemore" authorityYear="2013" class="Clitellata" family="Lumbricidae" genus="Eisenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Crassiclitellata" pageId="5" pageNumber="6" phylum="Annelida" rank="subSpecies" species="nordenskioldi" subSpecies="mongol">
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<emphasis italics="true" pageId="5" pageNumber="6">Eisenia nordenskioldi mongol</emphasis>
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</taxonomicName>
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, it is clearly separated objectively on mtDNA data (Appendix 1). This compares to its sibling species-complex: European
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<taxonomicName authorityName="species-complex. s. Blakemore" authorityYear="2010" baseAuthorityName="Savigny" baseAuthorityYear="1826" class="Clitellata" family="Lumbricidae" genus="Eisenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Crassiclitellata" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="fetida">
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<emphasis italics="true" pageId="5" pageNumber="6">Eisenia fetida</emphasis>
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</taxonomicName>
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(Savigny, 1826)
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<emphasis italics="true" pageId="5" pageNumber="6">vs</emphasis>
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.
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<taxonomicName baseAuthorityName="Bouché" baseAuthorityYear="1972" class="Clitellata" family="Lumbricidae" genus="Eisenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Crassiclitellata" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="andrei">
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<emphasis italics="true" pageId="5" pageNumber="6">Eisenia andrei</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Bouché">Bouche</normalizedToken>
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, 1972 that is claimed to differ molecularly on enzyme gel electrophoresis, e.g. by
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<bibRefCitation author="Jaenike, J" journalOrPublisher="Megadrilogica" pageId="11" pageNumber="12" pagination="6 - 8" refId="B32" refString="Jaenike, J, 1982. ' Eisenia foetida ' is two biological species. Megadrilogica 4: 6 - 8" title="' Eisenia foetida ' is two biological species." volume="4" year="1982">Jaenike (1982)</bibRefCitation>
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based on material from New York, but never yet on respective types of either taxon (see Appendix 1 and Discussion).
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/29791" pageId="5" pageNumber="6" start="Figure 3" startId="F3">
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<paragraph pageId="5" pageNumber="6">
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<emphasis bold="true" pageId="5" pageNumber="6">Figure 3.</emphasis>
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<emphasis bold="true" pageId="5" pageNumber="6">A</emphasis>
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<taxonomicName authorityName="Blakemore" authorityYear="2013" class="Clitellata" family="Lumbricidae" genus="Eisenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Crassiclitellata" pageId="5" pageNumber="6" phylum="Annelida" rank="subSpecies" species="nordenskioldi" subSpecies="onon">
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<emphasis italics="true" pageId="5" pageNumber="6">Eisenia nordenskioldi onon</emphasis>
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</taxonomicName>
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ssp. n. Holotype sketched as for Figs
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<emphasis bold="true" pageId="5" pageNumber="6">1-2</emphasis>
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<emphasis bold="true" pageId="5" pageNumber="6">B</emphasis>
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<taxonomicName authorityName="Kobayashi" authorityYear="1940" class="Clitellata" family="Lumbricidae" genus="Allolobophora" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Crassiclitellata" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="harbinensis">
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<emphasis italics="true" pageId="5" pageNumber="6">Allolobophora harbinensis</emphasis>
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</taxonomicName>
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Kobayashi, 1940: fig. 6 (
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<emphasis italics="true" pageId="5" pageNumber="6">incertae sedis</emphasis>
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) for comparison.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</subSection>
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</document> |