171 lines
17 KiB
XML
171 lines
17 KiB
XML
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<mods:title id="5728453E7DC6F126276EC92DAF3A7620">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
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<mods:namePart id="C0073B087351AD43749BB26BCB6424EC">Chamorro, William</mods:namePart>
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<mods:namePart id="1F0953E3C6A78272091E224632914EDB">Marin-Armijos, Diego</mods:namePart>
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<mods:namePart id="CAF2F68F85EC5D76E59AAEADFF5B0307">senjo, Angelico</mods:namePart>
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<mods:namePart id="5805C015711EBE1E0A5D2BE48684A0AF">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:date id="4309A68DFE82D643A8BF70E7492938BC">2019</mods:date>
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<mods:number id="D8D58278B112EE6629DC706A62F3839D">826</mods:number>
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<mods:classification id="9C94696302F31094C32F2E0D5BD3CB2F">journal article</mods:classification>
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<mods:identifier id="8E3A397B14C9516F905C1F4B77BF56B2" type="DOI">http://dx.doi.org/10.3897/zookeys.826.26488</mods:identifier>
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<treatment id="0EE1A86A45F78324852D7A98ECD7176C" ID-GBIF-Taxon="154127661" LSID="urn:lsid:plazi:treatment:0EE1A86A45F78324852D7A98ECD7176C" httpUri="http://treatment.plazi.org/id/0EE1A86A45F78324852D7A98ECD7176C" lastPageId="82" lastPageNumber="83" pageId="80" pageNumber="81">
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<subSubSection id="E0BACC9A997F6ECD47D932E7D7FE4B06" pageId="80" pageNumber="81" type="nomenclature">
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<paragraph id="1CA0932E3A99042F5B27976A86C9A01A" pageId="80" pageNumber="81">
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<taxonomicName id="F11F4F52C88B95DA929F993A02C8D4B3" ID-CoL="6CJ5C" authority="Paulian, 1939" authorityName="Paulian" authorityYear="1939" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum (Calhyboma) arrowi" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="arrowi" subGenus="Calhyboma">Deltochilum (Calhyboma) arrowi Paulian, 1939</taxonomicName>
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<taxonomicNameLabel id="471ED9F31D252D96C976306D866D3565" pageId="80" pageNumber="81">stat. n.</taxonomicNameLabel>
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Plates 19C, 57A
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</paragraph>
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</subSubSection>
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<subSubSection id="9AD442F17FEB4C84048B014F3BE80D9D" pageId="81" pageNumber="82" type="reference_group">
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<paragraph id="D58640BE8C90B127EFADF71DED7322BD" pageId="81" pageNumber="82">
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<taxonomicName id="7D9197E7E8EB58D4AD0C11F2A5BEF680" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum (E.) tessellatum" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tessellatum" subGenus="E.">
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<pageBreakToken id="FEAB2A37B194514E571667CED762FB07" pageId="81" pageNumber="82" start="start">Deltochilum</pageBreakToken>
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(E.) tessellatum
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</taxonomicName>
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var. Arrowi Paulian, 1939: 18 (original description. Type locality:
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<normalizedToken id="C622D975487E6BB8C1C5D3A13C08E0F8" originalValue="Équateur">Equateur</normalizedToken>
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[= Ecuador],
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<normalizedToken id="F2D16717556E24DC3FE7781C4CAD0242" originalValue="Río">Rio</normalizedToken>
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Pescado).
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</paragraph>
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<paragraph id="B50964632EF1C12C2DDCE4234D1512E4" pageId="81" pageNumber="82">
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<taxonomicName id="618A3B7D22B1CEE2D1025B9EB84A5C68" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Deltochilum tessellatum var. arrowi" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="variety" species="tessellatum" variety="arrowi">Deltochilum tessellatum var. arrowi</taxonomicName>
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:
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<bibRefCitation id="791BB68D68AD70C001F49B51D828CC01" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 203 (list of species from Latin America);
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<bibRefCitation id="DCB092ED98DF1C2F9B4491139770C595" author="Pereira, FS" journalOrPublisher="Revista Brasileira de Entomologia" pageId="274" pageNumber="275" pagination="7 - 50" title="The species of Deltochilumof thesubgenusCalhyboma Kolbe." volume="4" year="1955 a">
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Pereira and
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<normalizedToken id="418DBF67A368FEBA1ECA5EC934A89091" originalValue="D’Andretta">D'Andretta</normalizedToken>
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1955a
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</bibRefCitation>
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: 43 (redescription);
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<bibRefCitation id="757841AB2468D216115C257221162712" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
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: 94 (cited for Ecuador).
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</paragraph>
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<paragraph id="7D6395664F37FC340BBDB7119297D5FE" pageId="81" pageNumber="82">
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<taxonomicName id="57CBC72EE13E16133FCDA34AF3D4AA85" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Deltochilum var. arrowi" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="variety" variety="arrowi">Deltochilum var. arrowi</taxonomicName>
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:
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<bibRefCitation id="195C8FB768DE652106363F21CF0BE6F0" author="Contreras, GC" journalOrPublisher="Revista de la Academia Colombiana de Ciencias Fisicas y Naturales" pageId="265" pageNumber="266" pagination="220 - 229" title="Catalogo de los coleopteros colombianos." volume="8" year="1951">Contreras 1951</bibRefCitation>
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: 222 (cited for Colombia).
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</paragraph>
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</subSubSection>
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<subSubSection id="295311AB69FDD9D5846126321D238CE4" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="types specimens">
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<paragraph id="B056BEEDCEEAC6F7AC8589C64189160E" pageId="81" pageNumber="82">Types specimens.</paragraph>
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<paragraph id="2EFFB808B839323B79CE639CCC126AEE" pageId="81" pageNumber="82">
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<taxonomicName id="D1F59B3C410C6494B63D724280D363ED" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum (E.) tessellatum" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tessellatum" subGenus="E.">Deltochilum (E.) tessellatum</taxonomicName>
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var.
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<taxonomicName id="D1D0C4630F2A089F65E836D5A37FE354" lsidName="var. Arrowi" pageId="81" pageNumber="82" rank="variety" variety="Arrowi">Arrowi</taxonomicName>
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Paulian, 1939. The holotype (♂) and one paratype are deposited at the NHML (see
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<bibRefCitation id="00A22484102D00E7797D543F345DD8EA" author="Paulian, R" journalOrPublisher=") Annales de la Societe de France" pageId="274" pageNumber="275" pagination="1 - 48" title="Contribution a l'etude des Canthonides americains (Coleopt. Lamellic." volume="108" year="1939">Paulian 1939</bibRefCitation>
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: 18). Locality: Ecuador
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<normalizedToken id="14198CFAA87BE87128F08B3A3711C1D0" originalValue="Río">Rio</normalizedToken>
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Pescado. Examined.
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</paragraph>
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<paragraph id="170E3AEFDDC64E2DAEDC33E309D485B7" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
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Holotype (♂): "Ecuador /
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<normalizedToken id="B20ED2E896D42A1BDF9E173C703A36C4" originalValue="Río-">Rio-</normalizedToken>
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/ Pescado [p]", "v-17-1922 / 15000 ft / GHTate [p]", "Frank R. Manson / Collection [p]", "Ex coll. / F.R.Manson. / Brit. Mus. / 1923-
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<pageBreakToken id="38781AA563C42C3B9B75CAFB83EFE76F" pageId="82" pageNumber="83" start="start">141</pageBreakToken>
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[p]", "Deltochilum / tessellatum / var. arrowi Paulian / Type [hw]", "Holo- / type [p, red margin]".
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</paragraph>
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<paragraph id="242A2507F194142BCBE3E4BD0341A927" pageId="82" pageNumber="83">
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Paratype (♀): "Ecuador /
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<normalizedToken id="41DAB4E390E5463C5A7018110ED632A3" originalValue="Río-">Rio-</normalizedToken>
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/ Pescado [p]", "v-17-1922 / 15000 ft / GHTate [p]", "Ex coll. / F.R.Manson. / Brit. Mus. / 1923-141 [p]", "Para- Type [p, yellow margin]".
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</paragraph>
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</subSubSection>
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<subSubSection id="BA519713476E856F87581A9A13FA88B4" pageId="82" pageNumber="83" type="distribution">
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<paragraph id="CD966461388293318F3BB55794CBF1C6" pageId="82" pageNumber="83">Distribution.</paragraph>
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<paragraph id="C808079081C6CA69489C3E1390D1D436" pageId="82" pageNumber="83">Colombia and Ecuador.</paragraph>
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</subSubSection>
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<subSubSection id="A0588D2A224C059F6C80A446B99EEA8B" pageId="82" pageNumber="83" type="records examined">
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<paragraph id="DC705DFCC239036F9BFB1DB0AE52E5AC" pageId="82" pageNumber="83">Records examined.</paragraph>
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<paragraph id="9E578DE1F541D4B5752225427CC932CE" pageId="82" pageNumber="83">
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<normalizedToken id="99676D65CC5C99439E276F32102A3E91" originalValue="CAÑAR">CANAR</normalizedToken>
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: Javin, 1400 m (4 specimens CEMT). COTOPAXI: San Francisco de las Pampas (1 specimen CEMT).
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<normalizedToken id="F4A4B4C4B3B67D3C8F400ABA094B1592" originalValue="MANABÍ">MANABI</normalizedToken>
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:
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<normalizedToken id="5A6FD43D89A9C16986B9387B489295D9" originalValue="Río">Rio</normalizedToken>
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Pescado (2 specimens NHML). PICHINCHA: E B La Hesperia, 1200 m (1 specimen CEMT); Pacto, 1000 m (1 specimen CEMT).
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</paragraph>
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</subSubSection>
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<subSubSection id="F86CB40265C4777005CF3DF609DF5090" pageId="82" pageNumber="83" type="temporal data">
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<paragraph id="05F8918735793360B93C5E984DC1E84F" pageId="82" pageNumber="83">Temporal data.</paragraph>
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<paragraph id="850C0BC020AB8D4C1D6C2B8F915B20A9" pageId="82" pageNumber="83">Collected in January, February, March, and October.</paragraph>
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</subSubSection>
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<subSubSection id="EEBB3EC13113E6071D71638C19D5DE3D" pageId="82" pageNumber="83" type="remarks">
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<paragraph id="4AA4AD700D371A218D7A2437EF21E794" pageId="82" pageNumber="83">Remarks.</paragraph>
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<paragraph id="3ACFD409316EB75025C2EBC9AAA92146" pageId="82" pageNumber="83">Inhabits coastal evergreen foothill forests from 850-1200 m a.s.l. Collected using pitfall traps baited with pig feces.</paragraph>
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<paragraph id="FCE5A2159F6A9C91494B7CA17AFE5750" pageId="82" pageNumber="83">
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Paulian in 1939 described
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<taxonomicName id="5DB1A0D4D16019EF356E07C7C9B2489D" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Deltochilum (E.) tessellatum var. arrowi" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="variety" species="tessellatum" subGenus="E." variety="arrowi">Deltochilum (E.) tessellatum var. arrowi</taxonomicName>
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(written
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<taxonomicName id="6C1A8A15600C2EFF8E4723D3CCCA3020" genus="Arrowi" lsidName="Arrowi" pageId="82" pageNumber="83" rank="genus">Arrowi</taxonomicName>
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, type locality
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<normalizedToken id="BB7FF6652F772BFD46D9B01CFB93543A" originalValue="Río">Rio</normalizedToken>
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Pescado, Ecuador) as a variety of
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<taxonomicName id="860C8BF3CA2F696D5F242634F1228F81" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum tessellatum" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="tessellatum">Deltochilum tessellatum</taxonomicName>
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Bates, 1870 (type locality Gualaquiza, Ecuador, Amazonian region), which made the former name available as a subspecific category (see:
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<bibRefCitation id="8BF21BC75FDB07C011F0A8C559A84428" author="Celi, J" journalOrPublisher="Madrid, Espana" pageId="265" pageNumber="266" title="Codigo Internacional de Nomenclatura Zoologica. Cuarta edicion adoptada por la Union Internacional de Ciencias Biologicas." year="1999">ICZN 1999</bibRefCitation>
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, Article 45.6.4). However, upon examining the external and genital morphology of the type specimens of
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<taxonomicName id="7A11D476486A9B25C2B94D249A2FEBF8" lsidName="D. tessellatum" pageId="82" pageNumber="83" rank="subSpecies" species="tessellatum" subSpecies="arrowi">D. tessellatum arrowi</taxonomicName>
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(holotype ♂, deposited at the NHML, Plate 57A) and
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<taxonomicName id="4868379DA50E2E35D039EE003782E9CA" lsidName="D. tessellatum" pageId="82" pageNumber="83" rank="species" species="tessellatum">D. tessellatum</taxonomicName>
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(lectotype ♂ here designated, deposited at the MNHN, Plate 57B) (specifically, the differences observed in the elytral striae and shape of the aedeagus) we determined them to be two distinct species. Therefore, to maintain the subspecific name originally proposed by
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<bibRefCitation id="50BA6B7365D6FA246B4792E7810F9377" author="Paulian, R" journalOrPublisher=") Annales de la Societe de France" pageId="274" pageNumber="275" pagination="1 - 48" title="Contribution a l'etude des Canthonides americains (Coleopt. Lamellic." volume="108" year="1939">Paulian (1939)</bibRefCitation>
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, we elevate the variation
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<taxonomicName id="60440A89ABEE1D78EFB94D7318515BDF" lsidName="arrowi" pageId="82" pageNumber="83" rank="species" species="arrowi">arrowi</taxonomicName>
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to species level as follows:
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<taxonomicName id="FE2C642E483F8B696BF37A54ACF3AE71" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum (Calhyboma) arrowi" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="arrowi" subGenus="Calhyboma">Deltochilum (Calhyboma) arrowi</taxonomicName>
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Paulian, 1939 stat. n. The holotype is here illustrated (♂ Plate 57A) and originates from the locality
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<normalizedToken id="276E7B0958390F0A677429CA581DF1AD" originalValue="Río">Rio</normalizedToken>
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Pescado, Ecuador (
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<normalizedToken id="C24ABAE4A765E756FCF2EC377347D1CB" originalValue="Manabí">Manabi</normalizedToken>
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province, in the coastal region).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |