922 lines
101 KiB
XML
922 lines
101 KiB
XML
<document id="72DD60363F37D1D56E067F9D25F8B344" ID-CLB-Dataset="298084" ID-DOI="10.3897/zookeys.1204.117261" ID-GBIF-Dataset="4ee75651-5bbb-424e-bca6-97c5391ca8c5" ID-ZooBank="0704A2BB-6173-44BD-A1F7-CB3EDCFDC5DC" ID-publisher-id="117261" URI-arpha="29A78280-EE32-57DD-97A4-22DB0AEC910C" URI-zoobank="http://zoobank.org/0704A2BB-6173-44BD-A1F7-CB3EDCFDC5DC" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1717518118260" checkinUser="pensoft" docAuthor="Che Engku Abdullah, Che Engku Siti Mariam, Idris, Izwandy, Fahmi, Afiq Durrani Mohd, Flaxman, Beth & Hutchings, Pat" docDate="2024" docId="02B523B8A84257E99251CBE50DFBBA48" docLanguage="en" docName="ZooKeys 1204: 65-103" docOrigin="ZooKeys 1204" docSource="https://zookeys.pensoft.net/article/117261/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Marphysa ibaiensis Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings 2024, sp. nov." docType="treatment" docUuid="EFD572B1-A215-49FF-9C42-76DA2561E4D9" docUuidSource="ZooBank" docVersion="5" dtd-version="3.0" lastPageNumber="103" masterDocId="29A78280EE3257DD97A422DB0AEC910C" masterDocTitle="Four new species of Marphysa (Annelida, Eunicida, Eunicidae) from the east coast of Peninsular Malaysia" masterLastPageNumber="103" masterPageNumber="65" pageNumber="65" updateTime="1717537492882" updateUser="ExternalLinkService">
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<mods:titleInfo id="CCD6565F338A53645B9FE4FB10189038">
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<mods:title id="F762AB92E642769137CEA6E84E5981C7">Four new species of Marphysa (Annelida, Eunicida, Eunicidae) from the east coast of Peninsular Malaysia</mods:title>
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<mods:name id="B7CBCE0961F6EB947F845FFD62BDF740" type="personal">
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<mods:namePart id="19F3B6074A41F70790CB4E0FD0FD3044">Che Engku Abdullah, Che Engku Siti Mariam</mods:namePart>
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<mods:nameIdentifier id="2A47CA7F7AE7FFF00A9793876EA4AFA6" type="ORCID">https://orcid.org/0009-0002-8371-1141</mods:nameIdentifier>
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<mods:affiliation id="612B4B0BFE69BF43CDAA3FC957062986">Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:namePart id="AB2429CFAEDDC74DFA15866D59730609">Idris, Izwandy</mods:namePart>
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<mods:nameIdentifier id="0103C11021CB08CF5B7B24F9D8DA7989" type="ORCID">0000-0003-1516-8175</mods:nameIdentifier>
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<mods:affiliation id="A314345B30ED8A692E7652E60743D10C">Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:roleTerm id="FD1017F7F868A4E78AD485C27BE76A20">Author</mods:roleTerm>
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<mods:namePart id="6E74B03ACCCC271B8BE7A4BBFC6813B3">Fahmi, Afiq Durrani Mohd</mods:namePart>
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<mods:nameIdentifier id="8077F59CE80132B2EC68E9B14C4D5BA5" type="ORCID">0000-0002-5131-0098</mods:nameIdentifier>
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<mods:affiliation id="A41AD3ECCE8E78C1E3C02AE849366366">Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:namePart id="D07E25F8D9463A926EB7CA06C6499889">Flaxman, Beth</mods:namePart>
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<mods:nameIdentifier id="A2AA67133C6641E458D7169286F67148" type="ORCID">0000-0002-0329-9525</mods:nameIdentifier>
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<mods:affiliation id="250ECEE16B2A2C707AC0BE366B0B5514">South China Sea Repository and Reference Centre, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia & Mangrove Research Unit, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:namePart id="36E517EEFDCEAF1B6C4249ACD6376E10">Hutchings, Pat</mods:namePart>
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<mods:affiliation id="4EA65E830A7CAEF6B6EF2E65B9D093C4">Mangrove Research Unit, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia & Faculty of Science and Marine Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:title id="9499146C41F266F553D9A4288D55FEEA">ZooKeys</mods:title>
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<mods:date id="B77028E0BAE3E01165AFD537F58AA939">2024</mods:date>
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<mods:number id="49DA6F96319431A582160E1CDEB160A4">2024-06-04</mods:number>
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<mods:detail id="9B55AFB8C6BD5AA8FFC33D65C3DB852A" type="volume">
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<mods:number id="F0026BB1ED026FAB19C0216A415ED2D2">1204</mods:number>
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<mods:start id="788AE1BE8D0BB074DA0EDA197D7D045B">65</mods:start>
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<mods:end id="7DE96A8D49DD76F01A4AF4C1A03333E4">103</mods:end>
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<mods:classification id="84C8B6064714FE6DDD8DDD362D3F6F5C">journal article</mods:classification>
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<mods:identifier id="9E1B1B07F0AB72058149A6FF410EBD49" type="CLB-Dataset">298084</mods:identifier>
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<mods:identifier id="F1F12AADCEB7866B38A885A9D15B22C0" type="DOI">10.3897/zookeys.1204.117261</mods:identifier>
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<mods:identifier id="28383ECA17942B7457918B2953A79B62" type="ZooBank">0704A2BB-6173-44BD-A1F7-CB3EDCFDC5DC</mods:identifier>
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<treatment id="02B523B8A84257E99251CBE50DFBBA48" ID-DOI="http://doi.org/10.5281/zenodo.11479296" ID-GBIF-Taxon="232137204" ID-Zenodo-Dep="11479296" ID-arpha="02B523B8-A842-57E9-9251-CBE50DFBBA48" ID-zoobank="https://zoobank.org/EFD572B1-A215-49FF-9C42-76DA2561E4D9" LSID="urn:lsid:zoobank.org:act:EFD572B1-A215-49FF-9C42-76DA2561E4D9" LSID-ZBK="urn:lsid:zoobank.org:act:EFD572B1-A215-49FF-9C42-76DA2561E4D9" httpUri="http://treatment.plazi.org/id/02B523B8A84257E99251CBE50DFBBA48">
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<subSubSection id="BBFD0DD6EE74704813B68A755BEDF67F" type="nomenclature">
|
||
<paragraph id="5C238E4462563E3CCA7BA46451431410">
|
||
<taxonomicName id="DE324AE9B793B01CBF706F00EB9B9DC9" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="1661AED3203DC3F88DB7D193D5731674" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="62D89D393576B6A30741937F00637D4E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E2D2B13F7F20D4AC30B8D8D1EEB901FF" type="description">
|
||
<paragraph id="41B59F3E3A889384D1B98526776909FC">
|
||
<figureCitation id="B085626323D2B402B82ACE0DDE5806D8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval)." figureDoi="10.3897/zookeys.1204.117261.figure1" httpUri="https://binary.pensoft.net/fig/1062327">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="7B9C720558411C1215D2CA87430BCA53" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H)." figureDoi="10.3897/zookeys.1204.117261.figure2" httpUri="https://binary.pensoft.net/fig/1062328">2</figureCitation>
|
||
,
|
||
<figureCitation id="A8B8FAE254CBC23EBD28B9D025B3DEEC" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14</figureCitation>
|
||
,
|
||
<figureCitation id="A8AF8B30286CDF3B9CEA7D60E1F76BEC" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (B, C, E, F), paratype AM W. 54052 (A, D, G) A pectinate chaetae, chaetiger 97 B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 88 D pectinate chaetae, chaetiger 88 E – F pectinate chaetae, chaetiger 125 G serrations and projections on limbate chaetae, chaetiger 31. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 3. Thin, narrow heterodont; 4, 5. Thick, wide isodont; 7. Thick, wide anodont. Scale bars: 5 µm (A – B, D); 10 µm (C, F – H); 20 µm (E)." figureDoi="10.3897/zookeys.1204.117261.figure15" httpUri="https://binary.pensoft.net/fig/1062341">15</figureCitation>
|
||
,
|
||
<figureCitation id="4CEA833057722F6104147011361B5A4F" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sampling site in Kuala Ibai (estuary and lagoon area) A habitat of Marphysa ibaiensis B found in sediment deposited inside driftwood C live M. ibaiensis sp. nov." figureDoi="10.3897/zookeys.1204.117261.figure16" httpUri="https://binary.pensoft.net/fig/1062342">16</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E2TDI" type="material">
|
||
<paragraph id="55DD98DADE1D0CAA7D44C0BBEEAE8524">
|
||
<heading id="2B2198A807937975CE617D98F4B4B606" reason="title">Material examined.</heading>
|
||
</paragraph>
|
||
<paragraph id="CC0B7C0F4DCA6120C23E2895E0BA74D6">
|
||
<materialsCitation id="668719C544F38DE283216ECB0AB0CA74" ID-GBIF-Occurrence="4873238306" collectedFrom="lagoon, and estuary" collectingDate="2021-10" country="Malaysia" latitude="5.279667" location="Kuala Ibai" longLatPrecision="1" longitude="103.16895" specimenCode="UMTAnn 2179" specimenCount="1" stateProvince="Terengganu" typeStatus="holotype">
|
||
<emphasis id="588D3AB27AD01D812B42AE073B71A0FB" bold="true">
|
||
<emphasis id="BD0ABA82C6E3E1A840CD60363303E755" bold="true" italics="true">
|
||
<typeStatus id="2C6CABBA6AB8A6C917AC5B4CAD0B20BC" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="5A80CC1BEF9963FCB17534A7F95A91EF">UMTAnn 2179</specimenCode>
|
||
, complete, antero-ventrally dissected, some parapodia mounted for SEM
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="0A3F235243C56E396A8A19FA190469C4" ID-GBIF-Occurrence="4873238301" collectedFrom="lagoon, and estuary" collectingDate="2021-10" collectionCode="AM, ZRC" country="Malaysia" latitude="5.279667" location="Kuala Ibai" longLatPrecision="1" longitude="103.16895" specimenCode="AM W. 54052, LACM-AHF 13500, ZRC. ANN. 1610, SAM-MB-A 096022" specimenCount="1" stateProvince="Terengganu" typeStatus="paratype">
|
||
<emphasis id="8AD2831D42E324B66438C64BA0ACE822" bold="true">
|
||
<emphasis id="F6B0A01604C528EF5DCC63A689F51702" bold="true" italics="true">
|
||
<typeStatus id="CB8FD7AA34485BA0120929E3DFE57417" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="DDC0C7DFA95A658D3DA61E9E548CC1BD">
|
||
<named-content id="98BF037E130BD920EBCE220EEFAD2864" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/australian-museum" xlink_title="Australian Museum, Sydney, Australia">AM</named-content>
|
||
W. 54052
|
||
</specimenCode>
|
||
, complete, some parapodia mounted for SEM.
|
||
<specimenCode id="80DA1F37422BC1F4C07DD28A62091000">LACM-AHF 13500</specimenCode>
|
||
to 13502, complete, some parapodia removed;
|
||
<specimenCode id="C4CEB4BA273BAAB71B09F67065DC86CB">ZRC. ANN. 1610</specimenCode>
|
||
to 1612, complete, some parapodia removed;
|
||
<specimenCode id="C3248D57D2916044EB498D7A6259C09D">SAM-MB-A 096022</specimenCode>
|
||
, complete, some parapodia removed
|
||
</materialsCitation>
|
||
. All material was collected from the east coast of Peninsular
|
||
<collectingCountry id="F8755BBFFE6FC4015AB6596A9D27967E" name="Malaysia">Malaysia</collectingCountry>
|
||
,
|
||
<collectingRegion id="FD9D90F9F037D247DFB957618DF5ECDE" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
|
||
,
|
||
<location id="1F1FFDF56432CCBEE42339F396F9736C" LSID="urn:lsid:plazi:treatment:02B523B8A84257E99251CBE50DFBBA48:1F1FFDF56432CCBEE42339F396F9736C" country="Malaysia" latitude="5.279667" longLatPrecision="1" longitude="103.16895" name="Kuala Ibai" stateProvince="Terengganu">Kuala Ibai</location>
|
||
<collectedFrom id="A394BB9286F097FDB0603FDAF66A06E3">lagoon</collectedFrom>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESUDI" specific-use="{"type":"Point","coordinates":[103.169900,5.286633]}">
|
||
<geoCoordinate id="6704D45405365F7DD4E9ED1DEE843DDB" degrees="05" direction="north" minutes="17.198" orientation="latitude" precision="1" value="5.2866335">05 ° 17.198 ' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78BA1419F2F02C37F1EF7F361DC09E9D" degrees="103" direction="east" minutes="10.194" orientation="longitude" precision="1" value="103.1699">103 ° 10.194 ' E</geoCoordinate>
|
||
</named-content>
|
||
)
|
||
<collectedFrom id="FF2C2BA9E47400657DC3486C119184EE">and estuary</collectedFrom>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1UDI" specific-use="{"type":"Point","coordinates":[103.168950,5.279667]}">
|
||
<geoCoordinate id="ACC09010007C15BB7F7460D3A3357804" degrees="05" direction="north" minutes="16.780" orientation="latitude" precision="1" value="5.279667">05 ° 16.780 ' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E481B4AA9F1710653E47A6B72962902A" degrees="103" direction="east" minutes="10.137" orientation="longitude" precision="1" value="103.16895">103 ° 10.137 ' E</geoCoordinate>
|
||
</named-content>
|
||
),
|
||
<date id="7340706D795722E14B315E70A8D9034C" value="2021-10">
|
||
<collectingDate id="1B31C3F271B8DB5D9F75ADE3B577F719" value="2021-10">October 2021</collectingDate>
|
||
</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E6UDI" type="diagnosis">
|
||
<paragraph id="8FEEC7539326B70C0CCF38A0D3638A28">
|
||
<heading id="30BF1878646692F99FEDE7D9435CC894" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="3D3A86D1EB0B65E17C1B8F4630FF6629">Prostomium completely bilobed, five prostomial appendages without articulations; eyes absent. Peristomium without Peristomial cirri. Maxillary apparatus with four pairs of maxillae, an unpaired on the left side, MI with falcal arch extended at sub-right angle, basal outer edge arched, basal inner edge lacking curvature. MII with triangular teeth and without attachment lamella. MIII slightly curved, with equal-sized triangular teeth, without attachment lamella. MIV with curved attachment lamella. Branchiae distributed along entire body. Dorsal cirri without articulations; postchaetal lobe well developed in anterior regions. Ventral cirri with swollen, inflated base. Sub-aciculae black, blunt, and translucent at distal end, pale brown in posterior-most parapodia. Supra-acicular chaetae include limbate, pectinate, thin, narrow isodont with short and slender inner teeth, pectinate thin, narrow heterodont with short and slender inner teeth, pectinate thick, wide isodont with long or short and slender inner teeth, and pectinate thick, wide anodont with long and slender inner teeth. Subacicular chaetae include limbate and compound spinigers. Subacicular hook bidentate. Pygidium with two pairs of anal cirri, without articulation.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EEVDI" type="description">
|
||
<paragraph id="45A91B960D4E98BD6923A0CE60512061">
|
||
<heading id="2B4A415EEC1468DA70BE4209F44EE44F" reason="title">Description</heading>
|
||
</paragraph>
|
||
<paragraph id="64C14E2F60F11EA3A0BBF07F23225BC4">
|
||
<emphasis id="5830B836C4EF3D0D8D9B11C22112C31E" bold="true">
|
||
(based on
|
||
<typeStatus id="BF4B12EEBC507F50E59CEFA3AECE324D" type="holotype">holotype</typeStatus>
|
||
, with variation in parentheses for
|
||
<typeStatus id="09F400FFD9915D674513FD8F18EAC483" type="paratype">paratypes</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Preserved specimens beige (Fig.
|
||
<figureCitation id="4C4BF9AAEADA6395BCDC719EC719686B" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 A</figureCitation>
|
||
), ~ 195 (66–401) chaetigers and
|
||
<quantity id="81F81854B6B6C72FC787EB3297E131F1" metricMagnitude="-2" metricUnit="m" metricValue="5.2" unit="mm" value="52.0">52 mm</quantity>
|
||
(
|
||
<quantity id="4763DD8933C318CA4CACE0D4AEC8CD78" metricMagnitude="-2" metricUnit="m" metricValue="5.55" metricValueMax="9.1" metricValueMin="2.0" unit="mm" value="55.5" valueMax="91.0" valueMin="20.0">20–91 mm</quantity>
|
||
) long,
|
||
<abbrev id="ABBRID0EQVDI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
:
|
||
<quantity id="515822311600675EE30F4D4B8B81F2BA" metricMagnitude="-3" metricUnit="m" metricValue="4.5" unit="mm" value="4.5">4.5 mm</quantity>
|
||
(
|
||
<quantity id="A30AB95FF05D49FC545C222604A45E27" metricMagnitude="-3" metricUnit="m" metricValue="4.275" metricValueMax="6.3" metricValueMin="2.25" unit="mm" value="4.275" valueMax="6.3" valueMin="2.25">2.25–6.3 mm</quantity>
|
||
),
|
||
<abbrev id="ABBRID0EUVDI" xlink_title="width at chaetiger 10">W 10</abbrev>
|
||
:
|
||
<quantity id="E8479B5361C7D06D4410D2998ED80515" metricMagnitude="-3" metricUnit="m" metricValue="2.85" unit="mm" value="2.85">2.85 mm</quantity>
|
||
(
|
||
<quantity id="C167EBCC990C9FEC1F88A9242D6909EA" metricMagnitude="-3" metricUnit="m" metricValue="2.475" metricValueMax="3.75" metricValueMin="1.2" unit="mm" value="2.475" valueMax="3.75" valueMin="1.2">1.2–3.75 mm</quantity>
|
||
), excluding parapodia. Anterior region of body cylindrical, with shallow groove until median chaetigers (Fig.
|
||
<figureCitation id="771DEB484877645035753955D66FF07C" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 A</figureCitation>
|
||
); body depressed from chaetiger 30, elongated, and tapering at distal end. Live specimens red (Fig.
|
||
<figureCitation id="C06C33569D29F768779611228D2FF26F" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sampling site in Kuala Ibai (estuary and lagoon area) A habitat of Marphysa ibaiensis B found in sediment deposited inside driftwood C live M. ibaiensis sp. nov." figureDoi="10.3897/zookeys.1204.117261.figure16" httpUri="https://binary.pensoft.net/fig/1062342">16 C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="1922575A45BAB46F649C6375E1E5211A" ID-DOI="10.3897/zookeys.1204.117261.figure14" ID-arpha="9E0A7CBB-E1FB-5741-890C-0C75751EFC13" httpUri="https://binary.pensoft.net/fig/1062340" startId="F14">
|
||
<paragraph id="25746140A48B7FC062416005E29AD4D5">
|
||
<label id="77FBA5005320EBF301E1DBEA18AD87F0">Figure 14.</label>
|
||
</paragraph>
|
||
<paragraph id="F65712D85254BD4EFCE7EFC5A8297AF3">
|
||
<taxonomicName id="8ED13EA1E7F34A0277AB5F52E1E0F925" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="381EAE67A63891DE22F88F2E435C3CA6" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C69848B091A46DB83B44A01CF3F0F1F1" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype UMTAnn 2179 (
|
||
<emphasis id="E1E0E62C80F662FAE308ADAAB4AE18CF" bold="true">A – I</emphasis>
|
||
). Light microscopy images and digital drawing
|
||
<emphasis id="4B822AF3C0901F46B88247A341748EFF" bold="true">A</emphasis>
|
||
anterior end, lateral view. Arrow shows shallow groove
|
||
<emphasis id="CDC09595AFC1D8045B44D6D190DD5426" bold="true">B</emphasis>
|
||
anterior end, dorsal view
|
||
<emphasis id="A0EFFE6C4BA4B4F9236B52804D819C87" bold="true">C</emphasis>
|
||
maxillae, dorsal view
|
||
<emphasis id="1064F98867567058208972790131B54D" bold="true">D</emphasis>
|
||
mandibles, dorsal view
|
||
<emphasis id="CA61DD7658E29B1D4C886BE06DF97F0C" bold="true">E</emphasis>
|
||
bidentate hook, chaetiger 97
|
||
<emphasis id="F079FFDB5A3C20C218E036C379CBC533" bold="true">F</emphasis>
|
||
parapodium, chaetiger 10
|
||
<emphasis id="F3133F519DC61C5DE00207170D182C7B" bold="true">G</emphasis>
|
||
parapodium, chaetiger 97
|
||
<emphasis id="32D61069A1531915669E924AC5200495" bold="true">H</emphasis>
|
||
parapodium chaetiger 145
|
||
<emphasis id="BD423C476FFC0454405547FEE5D0A2CA" bold="true">I</emphasis>
|
||
posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (
|
||
<emphasis id="8C34F1B1910500D6E5EF1EE5EDBC746C" bold="true">A, D</emphasis>
|
||
); 1 mm (
|
||
<emphasis id="499D378D483ECADE8CB70A933A161C01" bold="true">B, C, I</emphasis>
|
||
); 20 µm (
|
||
<emphasis id="F84786EBDCA7CA038391877892FE74D6" bold="true">E</emphasis>
|
||
), 0.1 mm (
|
||
<emphasis id="1387C50EE5990B9FF4F2423923A692D0" bold="true">F – H</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D9BF622F1C929CE29337200C50F02B7">
|
||
Prostomium conically bilobed, with two dorsoventrally lobes separated by an anterior notch (Fig.
|
||
<figureCitation id="3E9D22E909010BFD8593905D683800DB" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 A, B</figureCitation>
|
||
). Prostomial appendages in a semicircle, median antennae separated by a gap. Palps, lateral and median antennae reaching first peristomium. Palpophores and ceratophores ring-shaped, short, and thin; palpostyles and ceratostyles tapering and slender. Prostomial appendage peduncles absent. Peristomium wider than prostomium; first ring 3 × longer than second ring, separation between rings distinct on all sides.
|
||
</paragraph>
|
||
<paragraph id="B97C7A3FCC96BC8557A8DEF88EAE873A">
|
||
Maxillae pale brown (Fig.
|
||
<figureCitation id="28E7D13DF51FA1087867684382B43ED8" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 C</figureCitation>
|
||
) and maxillary formula as follows:
|
||
<abbrev id="ABBRID0EAYDI" xlink_title="maxillary formula, roman numerals refer to number of maxilla">MF</abbrev>
|
||
= 1 + 1, 6 (5–6) + 7 (6–7), 7 (7–8) + 0, 4 + 10 (9–10), 1 + 1. Maxillary carrier ~ 2.2 × shorter than MI, rectangular anteriorly, triangular posteriorly. MI forceps-like, without attachment lamellae, falcal arch extended at sub-right angle, basal outer edge arched, basal inner edge lacking a curvature. Closing system is ~ 5.5 × shorter than MI. Ligament between MI and MII pale brown. MII without attachment lamella, teeth triangular, present on <1 / 2 of plate length. Ligament between MII and MIII pale brown. MIII single, longer than left MIV, slightly curved, with equal-sized triangular teeth, without attachment lamella. Left MIV short (<1 / 2 the size of right MIV) with curved attachment lamellae. Right MIV long, with teeth triangular and curved attachment lamellae, decreasing in size and teeth curved posteriorly. MV paired. Mandibles whitish with pale brown core, longer than MI; cutting longer than MI; cutting plates whitish (Fig.
|
||
<figureCitation id="4E7FF5C6AA4A71D512BD4C8382769C56" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EE0683D0990E794E244C601E04705596">
|
||
First few parapodia inserted ventrolaterally, but then becoming lateral in anterior region and dorsolaterally in subsequent segments. Chaetal lobes rounded on all chaetigers (Fig.
|
||
<figureCitation id="ADE32C2FB9C33475D1DBF082D919F536" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 F – H</figureCitation>
|
||
). Prechaetal lobe shorter than chaetal lobe along the entire body. Postchaetal lobe rounded and longer than chaetal lobe in anterior chaetigers and mid-body onwards (Fig.
|
||
<figureCitation id="DF0691EB5DE148419FD719ACFBE7CC3B" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 F – H</figureCitation>
|
||
), becoming shorter and absent in the posterior-most chaetigers. Dorsal cirri digitiform and slender, longer than ventral cirri anteriorly, as long as or shorter from mid-body and shorter in posterior chaetigers (Fig.
|
||
<figureCitation id="C2D70079317D6DFD3400F711C70B0B06" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 F – H</figureCitation>
|
||
). Ventral cirri digitiform in first chaetigers, basally inflated with digitiform tip from chaetiger six onwards (Fig.
|
||
<figureCitation id="517CCC463F4528002AFE7CD3915FBEB5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 F – H</figureCitation>
|
||
). Branchiae pectinate, starting from chaetiger 20 (11–65) and continuing to near end (~ 13 last chaetigers without branchiae), branchial filament 3 × longer than dorsal cirri where best developed; number of filaments increasing from one anteriorly to eight in mid-body, decreasing to six in last several chaetigers. Pygidial cirri attached to ventral side of pygidium, dorsal pair ~ 4 × longer than ventral (Fig.
|
||
<figureCitation id="69BFE4C35CC6AD13157ACA18860E864E" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="0F0C0803AF23BC31E90C33DBCCD83AA1">
|
||
Notoaciculae absent. Neuroaciculae black, blunt, and translucent at distal end along most of body, pale brown in posterior-most parapodia; ~ 3 or 4 per parapodium in anterior, one or two per parapodium in median and one per parapodium in posterior chaetigers (Fig.
|
||
<figureCitation id="DA59B0A17B629CE604ED23DFF8B5AC9C" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 F – H</figureCitation>
|
||
). Supra-acicular chaetae with limbate capillaries and pectinates.
|
||
<specimenCount id="CD9AF43CCC53337596783FBE71C1491C" count="5" type="generic" typeStatus="types">Five types</specimenCount>
|
||
of pectinate chaetae were identified (types 1, 3, 4, 5, 7) (see Fig.
|
||
<figureCitation id="5C50E0D8272E6F7A7942D7D1BCF0B0D0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H)." figureDoi="10.3897/zookeys.1204.117261.figure2" httpUri="https://binary.pensoft.net/fig/1062328">2</figureCitation>
|
||
): type 1: thin, narrow isodont with 12–15 short and slender inner teeth, present in anterior and median region (Fig.
|
||
<figureCitation id="EA79A86972474611E5201C35652B86AB" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (B, C, E, F), paratype AM W. 54052 (A, D, G) A pectinate chaetae, chaetiger 97 B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 88 D pectinate chaetae, chaetiger 88 E – F pectinate chaetae, chaetiger 125 G serrations and projections on limbate chaetae, chaetiger 31. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 3. Thin, narrow heterodont; 4, 5. Thick, wide isodont; 7. Thick, wide anodont. Scale bars: 5 µm (A – B, D); 10 µm (C, F – H); 20 µm (E)." figureDoi="10.3897/zookeys.1204.117261.figure15" httpUri="https://binary.pensoft.net/fig/1062341">15 A</figureCitation>
|
||
); type 3: thin, narrow heterodont with 12 short and slender inner teeth, outer teeth longer on one side, present in the anterior body region (Fig.
|
||
<figureCitation id="42F73CF815EFD68B3F01D92FF17462B2" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (B, C, E, F), paratype AM W. 54052 (A, D, G) A pectinate chaetae, chaetiger 97 B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 88 D pectinate chaetae, chaetiger 88 E – F pectinate chaetae, chaetiger 125 G serrations and projections on limbate chaetae, chaetiger 31. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 3. Thin, narrow heterodont; 4, 5. Thick, wide isodont; 7. Thick, wide anodont. Scale bars: 5 µm (A – B, D); 10 µm (C, F – H); 20 µm (E)." figureDoi="10.3897/zookeys.1204.117261.figure15" httpUri="https://binary.pensoft.net/fig/1062341">15 B</figureCitation>
|
||
); type 4: thick, wide isodont with 18–29 short and slender teeth, outer teeth different length to inner teeth, only present in median and posterior region (Fig.
|
||
<figureCitation id="AFEF0E2F2D087E9877542931C965D0E1" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (B, C, E, F), paratype AM W. 54052 (A, D, G) A pectinate chaetae, chaetiger 97 B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 88 D pectinate chaetae, chaetiger 88 E – F pectinate chaetae, chaetiger 125 G serrations and projections on limbate chaetae, chaetiger 31. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 3. Thin, narrow heterodont; 4, 5. Thick, wide isodont; 7. Thick, wide anodont. Scale bars: 5 µm (A – B, D); 10 µm (C, F – H); 20 µm (E)." figureDoi="10.3897/zookeys.1204.117261.figure15" httpUri="https://binary.pensoft.net/fig/1062341">15 C – F</figureCitation>
|
||
): type 5: thick, wide isodont with 15–18 long and slender inner teeth, present only posteriorly (Fig.
|
||
<figureCitation id="C53D48384E36A1F4D649A954A96D5308" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (B, C, E, F), paratype AM W. 54052 (A, D, G) A pectinate chaetae, chaetiger 97 B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 88 D pectinate chaetae, chaetiger 88 E – F pectinate chaetae, chaetiger 125 G serrations and projections on limbate chaetae, chaetiger 31. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 3. Thin, narrow heterodont; 4, 5. Thick, wide isodont; 7. Thick, wide anodont. Scale bars: 5 µm (A – B, D); 10 µm (C, F – H); 20 µm (E)." figureDoi="10.3897/zookeys.1204.117261.figure15" httpUri="https://binary.pensoft.net/fig/1062341">15 E, F</figureCitation>
|
||
); type 7: thick, wide anodont with ~ 15 long and slender inner teeth, only present in posterior parapodia (Fig.
|
||
<figureCitation id="51005E3F3DDDEDA04B8A82869CD46FA7" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (B, C, E, F), paratype AM W. 54052 (A, D, G) A pectinate chaetae, chaetiger 97 B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 88 D pectinate chaetae, chaetiger 88 E – F pectinate chaetae, chaetiger 125 G serrations and projections on limbate chaetae, chaetiger 31. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 3. Thin, narrow heterodont; 4, 5. Thick, wide isodont; 7. Thick, wide anodont. Scale bars: 5 µm (A – B, D); 10 µm (C, F – H); 20 µm (E)." figureDoi="10.3897/zookeys.1204.117261.figure15" httpUri="https://binary.pensoft.net/fig/1062341">15 F</figureCitation>
|
||
). Subacicular chaetae with compound spinigers and limbate capillaries in median and posterior chaetigers. Some limbate chaetae with inconspicuous serrations and numerous projections (Fig.
|
||
<figureCitation id="241A63716FE3CE050423CF02C76CBC04" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. SEM images of Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (B, C, E, F), paratype AM W. 54052 (A, D, G) A pectinate chaetae, chaetiger 97 B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 88 D pectinate chaetae, chaetiger 88 E – F pectinate chaetae, chaetiger 125 G serrations and projections on limbate chaetae, chaetiger 31. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 3. Thin, narrow heterodont; 4, 5. Thick, wide isodont; 7. Thick, wide anodont. Scale bars: 5 µm (A – B, D); 10 µm (C, F – H); 20 µm (E)." figureDoi="10.3897/zookeys.1204.117261.figure15" httpUri="https://binary.pensoft.net/fig/1062341">15 G</figureCitation>
|
||
). Subacicular hooks pale brown, translucent at distal end, from chaetiger 22 (22–46), 1–3 per parapodium; subacicular hooks bidentate present throughout (Fig.
|
||
<figureCitation id="FE98E7933C3BEA087BECB94EFCD8C509" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 E</figureCitation>
|
||
). Pygidium with crenulated margin, with two pairs of tapering pygidial cirri attached to ventral side of pygidium, dorsal pair ~ 4 × longer than ventral one (Fig.
|
||
<figureCitation id="D4D2E123EEB896170E09D1D9AA9570FB" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H)." figureDoi="10.3897/zookeys.1204.117261.figure14" httpUri="https://binary.pensoft.net/fig/1062340">14 I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="6A5085D5903F0E1BC0E61DBADDE2375A" ID-DOI="10.3897/zookeys.1204.117261.figure15" ID-arpha="F07FB15C-9783-5EC0-AF16-3DA63ABD91F3" httpUri="https://binary.pensoft.net/fig/1062341" startId="F15">
|
||
<paragraph id="79275CAFC6FAC89BC8E183444CF18989">
|
||
<label id="A5DF1A731F435116B474D4DDB0BC49FE">Figure 15.</label>
|
||
</paragraph>
|
||
<paragraph id="72FD4AC7D8F5C1E571BE6C992373A05E">
|
||
SEM images of
|
||
<taxonomicName id="477BC7F83477FCE46D3B6FE2A70601F9" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="5B0DC7479933772DE104FA0ED15478F6" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AFBB082A86E4AD9BA503B5E6F378CDA5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype UMTAnn 2179 (
|
||
<emphasis id="AA7019C1A33563EED0A77AF82468A3C1" bold="true">B, C, E, F</emphasis>
|
||
), paratype
|
||
<named-content id="31FC69BC65E07EE3D7C7BC1B737341F4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/australian-museum" xlink_title="Australian Museum, Sydney, Australia">AM</named-content>
|
||
W. 54052 (
|
||
<emphasis id="08A67D3C10A12DC8958F8CEF5D3FA8BE" bold="true">A, D, G</emphasis>
|
||
)
|
||
<emphasis id="B509A72B10D70B3FFA57578D59B841CF" bold="true">A</emphasis>
|
||
pectinate chaetae, chaetiger 97
|
||
<emphasis id="EB9F2C9D0592C84D3AAFB2AC57FAAE2B" bold="true">B</emphasis>
|
||
pectinate chaetae, chaetiger 10
|
||
<emphasis id="4B9376D72083EE70DEB202DEE2F2E665" bold="true">C</emphasis>
|
||
pectinate chaetae, chaetiger 88
|
||
<emphasis id="8A026A81CF764CD9B96AC27547DBBAA2" bold="true">D</emphasis>
|
||
pectinate chaetae, chaetiger 88
|
||
<emphasis id="8AF2EFDB2233022E98D232C5DC57F58E" bold="true">E – F</emphasis>
|
||
pectinate chaetae, chaetiger 125
|
||
<emphasis id="43D43680910280841D6095B2092F6281" bold="true">G</emphasis>
|
||
serrations and projections on limbate chaetae, chaetiger 31. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 3. Thin, narrow heterodont; 4, 5. Thick, wide isodont; 7. Thick, wide anodont. Scale bars: 5 µm (
|
||
<emphasis id="D7097D9D516EBD32F723930922CFCE3B" bold="true">A – B, D</emphasis>
|
||
); 10 µm (
|
||
<emphasis id="253FCDBB1715F0D80A2DB1EA58E8EDA1" bold="true">C, F – H</emphasis>
|
||
); 20 µm (
|
||
<emphasis id="889B9D5385C6A90DD5DF395AEC113D9E" bold="true">E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5544D0B58285CB46137AE8349347C571" ID-DOI="10.3897/zookeys.1204.117261.figure16" ID-arpha="14E99A8E-2794-54F4-91EE-F370CDAC5624" httpUri="https://binary.pensoft.net/fig/1062342" startId="F16">
|
||
<paragraph id="51EE7D7B5E8B4E61BA7A68151A122C6E">
|
||
<label id="1FBB4315D5E93BCB4A97FEFECCD072CB">Figure 16.</label>
|
||
</paragraph>
|
||
<paragraph id="049886BE11157F18D52F7E18BD59F644">
|
||
Sampling site in Kuala Ibai (estuary and lagoon area)
|
||
<emphasis id="8DB452A5A55430C1CA6835D67A3310AE" bold="true">A</emphasis>
|
||
habitat of
|
||
<taxonomicName id="B7DD6A0D0E6C9965A1E7081DEC95150F" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis">
|
||
<emphasis id="5A0CF093B253922BF6266CB435E421F4" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BBF05F88E3AA69FCC0D60F7A59A62EC5" bold="true">B</emphasis>
|
||
found in sediment deposited inside driftwood
|
||
<emphasis id="615B7A9519B192340D2D8B16391B91E0" bold="true">C</emphasis>
|
||
live
|
||
<taxonomicName id="422D966EC6119568BF1AC8AC8B563911" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="2E87F7E9D0637383B852B57DB2C9B9DE" italics="true">M. ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CB72FEDB6BA97C10251914CA11374E00" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EE4DI" type="etymology">
|
||
<paragraph id="B19A077CCA4DE265D6B510F7BA735154">
|
||
<heading id="DC695638B81432B0C822DAE54966B784" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="007800BCA66D4CA698695390B5060D51">Name refers to the type locality Kuala Ibai River.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EJ4DI" type="type locality">
|
||
<paragraph id="8145BD1810BB231FD9209D7C0C9E3AD7">
|
||
<heading id="D6A7906904CDF091FE59791B5788B928" reason="title">Type locality.</heading>
|
||
</paragraph>
|
||
<paragraph id="F6800882C2139857B4088284E1471584">
|
||
South
|
||
<collectingCountry id="1F924B76349C3A778576356F14D1930A" name="China">China</collectingCountry>
|
||
Sea,
|
||
<collectingCountry id="15D4F179A65B7E4E28BBFFC5B32A65B8" name="Malaysia">Malaysia</collectingCountry>
|
||
, east coast of Peninsular,
|
||
<collectingRegion id="6D38A58B7DF023517E632F51136A74BC" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
|
||
, Kuala Ibai river estuary and lagoon (see Fig.
|
||
<figureCitation id="01720FEDF74F43ACC0B4496585226CA7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval)." figureDoi="10.3897/zookeys.1204.117261.figure1" httpUri="https://binary.pensoft.net/fig/1062327">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ET4DI" type="distribution">
|
||
<paragraph id="1689DA781298924F449095085462670B">
|
||
<heading id="047993C1A72E7A2C74344AFCA8664195" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="1C9C4786E04590C934F75D4B8DA9CF35">Known only from the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EY4DI" type="habitat">
|
||
<paragraph id="A43E2019EF314F9FD6E092255CFB0284">
|
||
<heading id="080DFA7396673922B1EF2F867E4347A5" reason="title">Habitat.</heading>
|
||
</paragraph>
|
||
<paragraph id="72B7F5B2DF40BE4C5C9D948E9DF5C6E9">
|
||
Slightly gravelly sand sediment (Table
|
||
<tableCitation id="E2A8E29900DEE55855D05E832D062592" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Distance of sampling sites from river mouth, type of sediment textures and total organic matter content of sampling sites in Terengganu mangrove forest." httpUri="http://table.plazi.org/id/A7F5E5E96E47328E73CF7254E753C455" tableUuid="A7F5E5E96E47328E73CF7254E753C455">4</tableCitation>
|
||
) associated with oyster clumps within
|
||
<taxonomicName id="9E505C94144C58A6EE7CCBA95232FC69" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="5914D75B3B87D4B67255BED3435DBC9B" italics="true">Rhizophora</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="934BA69EAA7784BDFC419356222765A5" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sampling site in Kuala Ibai (estuary and lagoon area) A habitat of Marphysa ibaiensis B found in sediment deposited inside driftwood C live M. ibaiensis sp. nov." figureDoi="10.3897/zookeys.1204.117261.figure16" httpUri="https://binary.pensoft.net/fig/1062342">16 A</figureCitation>
|
||
), burrowing in sediment deposited inside driftwood bark (Fig.
|
||
<figureCitation id="39B0BA21D5C376803CEADE311465355D" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sampling site in Kuala Ibai (estuary and lagoon area) A habitat of Marphysa ibaiensis B found in sediment deposited inside driftwood C live M. ibaiensis sp. nov." figureDoi="10.3897/zookeys.1204.117261.figure16" httpUri="https://binary.pensoft.net/fig/1062342">16 B</figureCitation>
|
||
) with salinity 26 ‰ (estuary) and 18 ‰ (lagoon) during spring low tide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ER5DI" type="remarks">
|
||
<paragraph id="ECAA6FE2A3F3E65457E2620279D4DEB8">
|
||
<heading id="3DD7279092FD4CA72D5D3A2560FC6339" reason="title">Remarks.</heading>
|
||
</paragraph>
|
||
<paragraph id="D42A0587AECDAC5AAAE101FEAF95ABA9">
|
||
With the presence of subacicular limbate and compound spinigers in the median and posterior region,
|
||
<taxonomicName id="3C68CD8ECD40C0B0C00991D9D41D1F8C" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="4D879696A87B1688A78F7B8C9E6FABFF" italics="true">M. ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="182B35295EF185A6269CB51F901C5F64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belongs to Group E (Gravelyi). There are four
|
||
<taxonomicName id="5BA9E0AD80A2C7E0BE310D462DBEE60C">
|
||
<emphasis id="0C4522ED9835B03D4922EF28851071C3" italics="true">Marphysa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
belonging to this group;
|
||
<taxonomicName id="26FD7F12D81DCEA8D5E9EEDE11764394" authority="Pillai, 1958" authorityName="Pillai" authorityYear="1958" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="borradailei">
|
||
<emphasis id="CDF379DC2A333472A535659C43B108B2" italics="true">M. borradailei</emphasis>
|
||
<bibRefCitation id="8F4B4724C85FE7CE0EB92BB0FF3AD07D" author="Pillai TG" journalOrPublisher="Ceylon Journal of Science" pagination="94-106" refId="B33" refString="Pillai TG (1958) Studies on a brackish-water polychaetous annelid, Marphysa borradailei sp. n. from Ceylon. Ceylon Journal of Science 1: 94 - 106." title="Studies on a brackish-water polychaetous annelid, Marphysa borradailei sp. n. from Ceylon." volume="1" year="1958">Pillai, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
(type locality: Negombo Lagoon,
|
||
<collectingCountry id="8A8F24A1156B72D1A7515B1F6EC0CD61" name="Sri Lanka">Sri Lanka</collectingCountry>
|
||
),
|
||
<taxonomicName id="4ABEEAF409AB5E6BA0149E86ACE58259" authority="Glasby & Hutchings, 2010" authorityName="Glasby & Hutchings" authorityYear="2010" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="fauchaldi">
|
||
<emphasis id="EDCFBFF5E49DCD41BB26879739A22062" italics="true">M. fauchaldi</emphasis>
|
||
<bibRefCitation id="4A543861A4AED73E6976BD2581CE6FC0" DOI="10.11646/zootaxa.2352.1.2" author="Glasby CJ & Hutchings P" issue="1" journalOrPublisher="Zootaxa" pagination="29-45" refId="B13" refString="Glasby CJ, Hutchings P (2010) A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from Northern Australia and a review of similar taxa from the Indo-West Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352 (1): 29 - 45. https: // doi. org / 10.11646 / zootaxa. 2352.1. 2" title="A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from Northern Australia and a review of similar taxa from the Indo-West Pacific, including the genus Nauphanta Kinberg, 1865." volume="2352" year="2010">Glasby & Hutchings, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
(type locality: Ardatek Barrumundi farm, Darwin,
|
||
<collectingCountry id="162D6A264E8F84ED9F95C1D7271CCE22" name="Australia">Australia</collectingCountry>
|
||
),
|
||
<taxonomicName id="1599C1D01BB1C98FFC3336B75A83E57A" authorityName="Southern" authorityYear="1921" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="gravelyi">
|
||
<emphasis id="DA47BC0A02AC9294F18D6C005480DA8C" italics="true">M. gravelyi</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="570AE483A85C60413CEF4C915530D8F7" author="Southern R" refId="B37" refString="Southern R (1921) Polychaeta of the Chilka Lake, and also of fresh and brackish waters in other parts of India. Memoirs of the Indian Museum 5: 563 - 659. Available from: https: // www. biodiversitylibrary. org / part / 168161" url="https://www.biodiversitylibrary.org/part/168161" year="1921">Southern, 1921</bibRefCitation>
|
||
(type locality: Chilka Lake,
|
||
<collectingCountry id="3112E2FCFBEC987221B2ABDDA3A77FD3" name="India">India</collectingCountry>
|
||
) and
|
||
<taxonomicName id="776702DD75AE6232516DFED5ED09130E" authority="Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby, 2020" authorityName="Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby" authorityYear="2020" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="madrasi">
|
||
<emphasis id="144F56C085751719069E8C8C8713A6AA" italics="true">M. madrasi</emphasis>
|
||
<bibRefCitation id="742C52E3E596D10E72DC0B24028A45B0" DOI="10.11646/zootaxa.4852.3.2" author="Hutchings P & Lavesque N & Priscilla L & Daffe G & Malathi E & Glasby CJ" issue="3" journalOrPublisher="Zootaxa" pagination="285-308" refId="B17" refString="Hutchings P, Lavesque N, Priscilla L, Daffe G, Malathi E, Glasby CJ (2020) A new species of Marphysa (Annelida: Eunicida: Eunicidae) from India, with notes on previously described or reported species from the region. Zootaxa 4852 (3): 285 - 308. https: // doi. org / 10.11646 / zootaxa. 4852.3. 2" title="A new species of Marphysa (Annelida: Eunicida: Eunicidae) from India, with notes on previously described or reported species from the region." volume="4852" year="2020">Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby, 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
(type locality: Chennai,
|
||
<collectingCountry id="05724BD5E06F91FB930F74545C7A7121" name="India">India</collectingCountry>
|
||
). The morphological features of these species are given in Table
|
||
<tableCitation id="C384B7345D2C1BE22D2875AB3F630967" captionStart="Table 7" captionStartId="T7" captionText="Table 7. Morphological features comparison between Marphysa Group E (Gravelyi) described in this study and species occurring worldwide. The features for new species are based on the holotype, with variation in parentheses for paratypes. Abbreviations: MF: maxillary formula, roman numerals refer to number of maxilla; PR-I: first peristomial ring; PR-II: second peristomial ring; p / a: present / absent; NIA: no information available. The major differences between the species are marked with asterisk (*)." httpUri="http://table.plazi.org/id/AF48FD41F4F3E2896DF72709ABBA25E3" tableUuid="AF48FD41F4F3E2896DF72709ABBA25E3">7</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="AF48FD41F4F3E2896DF72709ABBA25E3" ID-Table-UUID="AF48FD41F4F3E2896DF72709ABBA25E3" httpUri="http://table.plazi.org/id/AF48FD41F4F3E2896DF72709ABBA25E3" startId="T7">
|
||
<paragraph id="531B69FCF9226448AAB1967539060E05">
|
||
<label id="BF1ED01997A3C9D7362EDE91ACFFCA72">Table 7.</label>
|
||
</paragraph>
|
||
<paragraph id="7AA57BDB57F2C07AC85FD1821E08844A">
|
||
Morphological features comparison between
|
||
<taxonomicName id="6C6CD5677539A9ACF54ADC7DE67C23CD" authorityName="de Quatrefages" authorityYear="1866" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="genus">
|
||
<emphasis id="F61C0F7CD0A0D3ABE4360B0721A3FF2B" italics="true">Marphysa</emphasis>
|
||
</taxonomicName>
|
||
Group E (Gravelyi) described in this study and species occurring worldwide. The features for new species are based on the holotype, with variation in parentheses for paratypes. Abbreviations:
|
||
<abbrev id="ABBRID0EKBAK" xlink_title="maxillary formula, roman numerals refer to number of maxilla">MF</abbrev>
|
||
: maxillary formula, roman numerals refer to number of maxilla;
|
||
<abbrev id="ABBRID0EOBAK" xlink_title="first peristomial ring">PR-I</abbrev>
|
||
: first peristomial ring;
|
||
<abbrev id="ABBRID0ESBAK" xlink_title="second peristomial ring">PR-II</abbrev>
|
||
: second peristomial ring;
|
||
<abbrev id="ABBRID0EWBAK" xlink_title="present/absent">p / a</abbrev>
|
||
: present / absent;
|
||
<abbrev id="ABBRID0E1BAK" xlink_title="no information available">NIA</abbrev>
|
||
: no information available. The major differences between the species are marked with asterisk (*).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1785654C6BF4F56B57C88E3B6C1E9028">
|
||
<table id="TID0EYOCI" rules="all">
|
||
<tbody id="113B71D98A324C88BF2E673D1C2378FC">
|
||
<tr id="C3B53DD4100DAAAE5641137FC9CB2B47">
|
||
<th id="4D3295185D7D304429A8E60EE72A3C1C" colspan="1" rowspan="1">Morphological feature</th>
|
||
<th id="CFFC6096DC7BCDD7B39D0F96614F7B15" colspan="1" rowspan="1">
|
||
<taxonomicName id="C30188978C8DB6434978FE6B60EC9FA2" authority="Pillai, 1958" authorityName="Pillai" authorityYear="1958" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="borradailei">
|
||
<emphasis id="202ABCB6D50638959C800A69B44F5233" italics="true">M. borradailei</emphasis>
|
||
<bibRefCitation id="270C8FE171F1EB97FF5524FE34BBBC9A" author="Pillai TG" journalOrPublisher="Ceylon Journal of Science" pagination="94-106" refId="B33" refString="Pillai TG (1958) Studies on a brackish-water polychaetous annelid, Marphysa borradailei sp. n. from Ceylon. Ceylon Journal of Science 1: 94 - 106." title="Studies on a brackish-water polychaetous annelid, Marphysa borradailei sp. n. from Ceylon." volume="1" year="1958">Pillai, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="33F3AE3CEB5606F052E568341C695106" colspan="1" rowspan="1">
|
||
<taxonomicName id="C846707094941581F1801F54D7BC8659" authorityName="Southern" authorityYear="1921" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="gravelyi">
|
||
<emphasis id="63130E7E95A9B6A822409C1503D3790F" italics="true">M. gravelyi</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="CC21C94EE210F3748A14D9F5114F38EF" author="Southern R" refId="B37" refString="Southern R (1921) Polychaeta of the Chilka Lake, and also of fresh and brackish waters in other parts of India. Memoirs of the Indian Museum 5: 563 - 659. Available from: https: // www. biodiversitylibrary. org / part / 168161" url="https://www.biodiversitylibrary.org/part/168161" year="1921">Southern, 1921</bibRefCitation>
|
||
</th>
|
||
<th id="21B6D69D8B045148B18718BA8016E77E" colspan="1" rowspan="1">
|
||
<taxonomicName id="610B07A124951F52D88B8BBE72949B0C" authority="Glasby & Hutchings, 2010" authorityName="Glasby & Hutchings" authorityYear="2010" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="fauchaldi">
|
||
<emphasis id="BC49064CB76D3FEB0B418C59B9B80558" italics="true">M. fauchaldi</emphasis>
|
||
<bibRefCitation id="538F4697AAC52057096A4F9F72016D36" DOI="10.11646/zootaxa.2352.1.2" author="Glasby CJ & Hutchings P" issue="1" journalOrPublisher="Zootaxa" pagination="29-45" refId="B13" refString="Glasby CJ, Hutchings P (2010) A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from Northern Australia and a review of similar taxa from the Indo-West Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352 (1): 29 - 45. https: // doi. org / 10.11646 / zootaxa. 2352.1. 2" title="A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from Northern Australia and a review of similar taxa from the Indo-West Pacific, including the genus Nauphanta Kinberg, 1865." volume="2352" year="2010">Glasby & Hutchings, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="79E79F4D3E254F7E05E60FC1D0B0FFC1" colspan="1" rowspan="1">
|
||
<taxonomicName id="1E9DC579444176D8F66BE69C585BCF20" authority="Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby, 2020" authorityName="Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby" authorityYear="2020" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="madrasi">
|
||
<emphasis id="695A7B6C8BEA05E32B323FCE4EC4F66D" italics="true">M. madrasi</emphasis>
|
||
<bibRefCitation id="34CA45EF6BF5233A105BEDBCEDE8A432" DOI="10.11646/zootaxa.4852.3.2" author="Hutchings P & Lavesque N & Priscilla L & Daffe G & Malathi E & Glasby CJ" issue="3" journalOrPublisher="Zootaxa" pagination="285-308" refId="B17" refString="Hutchings P, Lavesque N, Priscilla L, Daffe G, Malathi E, Glasby CJ (2020) A new species of Marphysa (Annelida: Eunicida: Eunicidae) from India, with notes on previously described or reported species from the region. Zootaxa 4852 (3): 285 - 308. https: // doi. org / 10.11646 / zootaxa. 4852.3. 2" title="A new species of Marphysa (Annelida: Eunicida: Eunicidae) from India, with notes on previously described or reported species from the region." volume="4852" year="2020">Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby, 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="3E4E763F85F95FEFCFD2EC932DCF0BD8" colspan="1" rowspan="1">
|
||
<taxonomicName id="C501EA1E71DDEC1C7E5BCEF5500B4AC3" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="535753773839BF782EACB450D7CC10D0" italics="true">M. ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="28A79042A6FCE1176367A1963DAEB5AC" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
</tr>
|
||
<tr id="31249585C47A415494D4A993494C3162">
|
||
<td id="EC7040BC8DA8CC081C729139CD768A47" colspan="1" rowspan="1">Source of Information</td>
|
||
<td id="BCFB997CB235385B049B7348281FC154" colspan="1" rowspan="1">
|
||
Lectotype BMNH 1960.3. 13.6 (
|
||
<bibRefCitation id="33306897D4162480176B92B4A79133E2" DOI="10.11646/zootaxa.2352.1.2" author="Glasby and Hutchings" firstAuthor="Glasby" issue="1" journalOrPublisher="Zootaxa" pagination="29-45" refId="B13" refString="Glasby CJ, Hutchings P (2010) A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from Northern Australia and a review of similar taxa from the Indo-West Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352 (1): 29–45. https://doi.org/10.11646/zootaxa.2352.1.2" title="A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from Northern Australia and a review of similar taxa from the Indo-West Pacific, including the genus Nauphanta Kinberg, 1865." volume="2352" year="2010">Glasby and Hutchings 2010</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="BAB09F761F76C57449113C93616D3A26" colspan="1" rowspan="1">
|
||
Paratypes 1938.5. 7.55 and the type description (
|
||
<bibRefCitation id="6D0A16651D1E0654E5CC1833F954B9DF" DOI="10.11646/zootaxa.4852.3.2" author="Hutchings" etAl="et al." firstAuthor="Hutchings" issue="3" journalOrPublisher="Zootaxa" pagination="285-308" refId="B17" refString="Hutchings P, Lavesque N, Priscilla L, Daffe G, Malathi E, Glasby CJ (2020) A new species of Marphysa (Annelida: Eunicida: Eunicidae) from India, with notes on previously described or reported species from the region. Zootaxa 4852 (3): 285–308. https://doi.org/10.11646/zootaxa.4852.3.2" title="A new species of Marphysa (Annelida: Eunicida: Eunicidae) from India, with notes on previously described or reported species from the region." volume="4852" year="2020">Hutchings et al. 2020</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="8A75824A5C8961CBDEFEB923083D5266" colspan="1" rowspan="1">
|
||
Holotype NTM W 23040 (
|
||
<bibRefCitation id="345AEEB90302857408070493F5AE2096" DOI="10.11646/zootaxa.2352.1.2" author="Glasby and Hutchings" firstAuthor="Glasby" issue="1" journalOrPublisher="Zootaxa" pagination="29-45" refId="B13" refString="Glasby CJ, Hutchings P (2010) A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from Northern Australia and a review of similar taxa from the Indo-West Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352 (1): 29–45. https://doi.org/10.11646/zootaxa.2352.1.2" title="A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from Northern Australia and a review of similar taxa from the Indo-West Pacific, including the genus Nauphanta Kinberg, 1865." volume="2352" year="2010">Glasby and Hutchings 2010</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="7A262B237751ADA12AC28163FB13F7C6" colspan="1" rowspan="1">
|
||
Holotype NL-ENNORE _ 01 (ZSI); Paratypes: ZSI-HQ / GNC / AN 6072 / 1 (
|
||
<bibRefCitation id="A41ED2DB8C798CB7DD6F3AADCC6FB7E0" DOI="10.11646/zootaxa.4852.3.2" author="Hutchings" etAl="et al." firstAuthor="Hutchings" issue="3" journalOrPublisher="Zootaxa" pagination="285-308" refId="B17" refString="Hutchings P, Lavesque N, Priscilla L, Daffe G, Malathi E, Glasby CJ (2020) A new species of Marphysa (Annelida: Eunicida: Eunicidae) from India, with notes on previously described or reported species from the region. Zootaxa 4852 (3): 285–308. https://doi.org/10.11646/zootaxa.4852.3.2" title="A new species of Marphysa (Annelida: Eunicida: Eunicidae) from India, with notes on previously described or reported species from the region." volume="4852" year="2020">Hutchings et al. 2020</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="1E40537FEDFC7FAB7A0C3DDA6FBB61A0" colspan="1" rowspan="1">Holotype: UMTAnn 2179 (this study)</td>
|
||
</tr>
|
||
<tr id="C24E3674E4FA5C5A86F882B26CAF44D5">
|
||
<td id="10F663F03A9AA0D4E13BDAE33EDF3FDD" colspan="1" rowspan="1">
|
||
Size (mm):
|
||
<abbrev id="ABBRID0E4FAK" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
,
|
||
<abbrev id="ABBRID0EBGAK" xlink_title="width at chaetiger 10">W 10</abbrev>
|
||
</td>
|
||
<td id="8A2A369DA7739821A0AEE7DA9E56762A" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EIGAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="57B769576D806861B7214F76A7D722BB" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EQGAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="FABB882939AB0027E8A584E7427C61D8" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EYGAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="05105F8758C550E2F5D01D04DDC336D3" colspan="1" rowspan="1">6 (4–9), 2.5 (2–3.9)</td>
|
||
<td id="21BA7C120EBA5F034389AF560D38366A" colspan="1" rowspan="1">4.5 (2.25–6.3), 2.85 (1.2–3.75)</td>
|
||
</tr>
|
||
<tr id="AC30402B08D4C344C3196C01D628B17B">
|
||
<td id="384E653E71EBBDE88539FFE2DE00AD6E" colspan="1" rowspan="1">Prostomium: shape</td>
|
||
<td id="3BFCDD8EFB10506DCA83834BC0653205" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="B8F7235536C3DF0F2B723756C54A79D4" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="2CF23B5BFBF4F6A20504A0D70C19736D" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="1DCF1B6644E3E7708FFE0622B31DE4B5" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="FCC7781B94FA6E40DF2C761F0B2EE61F" colspan="1" rowspan="1">Bilobed</td>
|
||
</tr>
|
||
<tr id="F99CB19987EC0A28E3FC3C1866B0D2CF">
|
||
<td id="2B374A578E5FE1987117662FB6DB5D1B" colspan="1" rowspan="1">Palps: reaching</td>
|
||
<td id="10FF66B085B624C47B1EC4F01C2B5745" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0E4HAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="ABD03ADCEDA554FF705DA3AB321535F8" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EFIAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="9DA566082AF8E5972D75C068BC05620F" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ENIAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="B23B8BD1B597A923A6A8151FD12FAD85" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EVIAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="BCD72FF720816E5A6BDA4013EBC5CD8D" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0E4IAK" xlink_title="first peristomial ring">PR-I</abbrev>
|
||
</td>
|
||
</tr>
|
||
<tr id="0C7B2B0B1CE385FB5CDBBAA6623BFEDD">
|
||
<td id="BBADC3A0B1A3589C44B5963FCA79EDFE" colspan="1" rowspan="1">Lateral antennae: reaching</td>
|
||
<td id="CC08C18ADB8633D161FEA0E0BAC7B0AB" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EJJAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="65435DE58D494E82E2474317ECCA8C1C" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ERJAK" xlink_title="first peristomial ring">PR-I</abbrev>
|
||
</td>
|
||
<td id="155E5BDC24608B3ACD0638DDCAC4FD5D" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EZJAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="AD6DCA085C0BF9F25D9DB1A80598A4E1" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EBKAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="61AEACFB6D60F825C91AEF95AB0D5AC4" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EJKAK" xlink_title="first peristomial ring">PR-I</abbrev>
|
||
</td>
|
||
</tr>
|
||
<tr id="E53DBD33674AE99B5F3E01441AC25C30">
|
||
<td id="7C4BB93BF9BAB35FAAD7B846AAAC935D" colspan="1" rowspan="1">Median antennae: reaching</td>
|
||
<td id="A9D0E29D274BE77BAAC7668F35DE3164" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EVKAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="734A5140DB1BEF6425DDE461F4192386" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0E4KAK" xlink_title="first peristomial ring">PR-I</abbrev>
|
||
</td>
|
||
<td id="B7FA849E3DE7FC3E8F29116CB615A973" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EFLAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="CA43D608BB36434585061BD82CA6E9C0" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ENLAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="4AC44BDC73F52AE0BB1A1FC5CAAC3E29" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EVLAK" xlink_title="first peristomial ring">PR-I</abbrev>
|
||
</td>
|
||
</tr>
|
||
<tr id="72DAAAD80F6D35390FA18C17FFE6D391">
|
||
<td id="13045AAD4AC610BA2E657234DB2EDA31" colspan="1" rowspan="1">Peduncle in prostomial appendages</td>
|
||
<td id="AF0DA5BE4B0065C696C055B7F6A333C9" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EBMAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="6FCFB1857DEEEDE6B1FAC8BD8457F2AB" colspan="1" rowspan="1">Absent</td>
|
||
<td id="171E22DCB7817F48CDFA5E26F92203E2" colspan="1" rowspan="1">Present</td>
|
||
<td id="87B7BB12CD0C65B1892573FF70B642D9" colspan="1" rowspan="1">Absent</td>
|
||
<td id="B12B4E0CE508470127C9372BA2E9A765" colspan="1" rowspan="1">Absent</td>
|
||
</tr>
|
||
<tr id="92D86A2E86072532FFA763AF8DD728B7">
|
||
<td id="41672C89683F9C13B6C27BEED30BD7FF" colspan="1" rowspan="1">Eyes *</td>
|
||
<td id="1F320F53411B435E75BE57FC2E24D80D" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EZMAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="BCC37042A8D8C341974FE776627AB459" colspan="1" rowspan="1">Present</td>
|
||
<td id="AE6E34B48E685B368FD5CB203148BE1D" colspan="1" rowspan="1">Absent</td>
|
||
<td id="7BBD47448D36E88F13DCCABB3A46950A" colspan="1" rowspan="1">Present</td>
|
||
<td id="694B882F065E0C1A5B9C4653E965EE9A" colspan="1" rowspan="1">Absent</td>
|
||
</tr>
|
||
<tr id="4BE6969A33A4FC02B67954877FAF923F">
|
||
<td id="BBFD03D51657518AF63B6CDCA45C5860" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EONAK" xlink_title="maxillary formula, roman numerals refer to number of maxilla">MF</abbrev>
|
||
: MII, MIII, MIV *
|
||
</td>
|
||
<td id="7F0C350E6B882233A8BFDBD66337001F" colspan="1" rowspan="1">
|
||
6,
|
||
<abbrev id="ABBRID0EWNAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="D77C5BD929C5EDA18B8B0AE8154348D4" colspan="1" rowspan="1">5 + 6, 12–13 + 0, 4 + 8</td>
|
||
<td id="7335693D8FB65B6AFD6C600ADBDA2BF8" colspan="1" rowspan="1">5 + 6, 7 + 0, 4 + 9</td>
|
||
<td id="DE4BCC433A0B73887698B5C278F5B538" colspan="1" rowspan="1">8 + 9, 10 + 0, 7 + 11</td>
|
||
<td id="25C4972E9690C1A7856BE3434FE0D5A5" colspan="1" rowspan="1">6 (5–6) + 7 (6–7), 7 (7–8) + 0, 4 + 10 (9–10)</td>
|
||
</tr>
|
||
<tr id="CACD632BA3C892DC8A56AB2368E08FA5">
|
||
<td id="9699C5E5D11D03B2891211052D7C21F4" colspan="1" rowspan="1">Branchiae: shape</td>
|
||
<td id="6D8CFC3B2ED1E8FD87B5DA8E3F38E5DD" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
<td id="AA528BA8FB0D2E12100D851EE75C9E46" colspan="1" rowspan="1">Pectinate</td>
|
||
<td id="0CCC911C05C414BD0098D6B1160D9936" colspan="1" rowspan="1">Pectinate</td>
|
||
<td id="98192531BC25E0F8A1EAEEF78D77EBFF" colspan="1" rowspan="1">Pectinate</td>
|
||
<td id="0AF42705B5F4FC335F2A72E37008C809" colspan="1" rowspan="1">Pectinate</td>
|
||
</tr>
|
||
<tr id="470A082017B8851DD1584E0D9DCFF307">
|
||
<td id="F753164B39332A46B2F2B7DF6B56ABEA" colspan="1" rowspan="1">Branchiae: start chaetiger; last chaetiger before pygidium</td>
|
||
<td id="9915E9C45A6809BF3BF20390D48E91D8" colspan="1" rowspan="1">7–60; end ~ 10 last chaetiger</td>
|
||
<td id="0B1D0C3CA563AAA2AAD12C25ADB92349" colspan="1" rowspan="1">22–52; end ~ 20 last chaetiger</td>
|
||
<td id="13B3CDA27BAB76A9DE4DB2B0768229F8" colspan="1" rowspan="1">31; end ~ 10 last chaetiger</td>
|
||
<td id="8551AD73A5B302D2359D179821C22140" colspan="1" rowspan="1">48–50, end ~ 10 last chaetiger</td>
|
||
<td id="393DE6339BD2964BEEE9820C9C4FDA8E" colspan="1" rowspan="1">20 (11–65); end ~ 13 last chaetiger</td>
|
||
</tr>
|
||
<tr id="2CFCFB32CEFA04CE1252FA995972C6AD">
|
||
<td id="A82D792BDC84B6CC03CB4503C4F2ADF2" colspan="1" rowspan="1">Branchial filaments: numbers</td>
|
||
<td id="5BA90757098D0B3BBB8346E6447552A5" colspan="1" rowspan="1">20</td>
|
||
<td id="6E782D1F526DB05C7676516B0B1E786F" colspan="1" rowspan="1">9</td>
|
||
<td id="AE2D3E24DB9C8D683E54A49BAD2416CD" colspan="1" rowspan="1">9</td>
|
||
<td id="921B5C759F7BBA6C96F00997AEEFFB10" colspan="1" rowspan="1">9</td>
|
||
<td id="0F27F311F8989C9D34C973255C10FA9E" colspan="1" rowspan="1">8</td>
|
||
</tr>
|
||
<tr id="01890AFE9E2B11D3337DA812C5C93521">
|
||
<td id="AA4C55C83CFC00D1B9224D019AEF8EC0" colspan="1" rowspan="1">Dorsal cirri: shaped</td>
|
||
<td id="8495CA33A41E734EBC2D2D49F60D1566" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EGQAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="40CA27F2080BCD55BC0D52779FD90A88" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EOQAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="CD39C16A6167FD121D095362CABDA31E" colspan="1" rowspan="1">Conical</td>
|
||
<td id="AFE3CC0C9CA18D6D82CD83996F9C6117" colspan="1" rowspan="1">Digitiform</td>
|
||
<td id="FA7A5DABD1694E7407E015DCBF074344" colspan="1" rowspan="1">Digitiform</td>
|
||
</tr>
|
||
<tr id="A86719D9442672993521890FAF07204C">
|
||
<td id="70F6C072468AE3AA8F0A2E645DAAD185" colspan="1" rowspan="1">Prechaetal lobe: shaped</td>
|
||
<td id="5D7EB43A4FB42E50C98B33A2B2A0E06C" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EDRAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="80EE7774B39D666A8FF790364171258D" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ELRAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="D51CC0AF8A2F303ADF589D18F1231BB8" colspan="1" rowspan="1">Ridge</td>
|
||
<td id="1BCC5435FCAE3FAEB5132C19AF8A0E19" colspan="1" rowspan="1">Transverse fold</td>
|
||
<td id="EBE6D6F2F6A6F16B89CD5E8A5D467B9B" colspan="1" rowspan="1">Transverse fold</td>
|
||
</tr>
|
||
<tr id="AF5D1F79F45B7310FD33552E3BADF09A">
|
||
<td id="4B3468EB648ACD7D504409260C616435" colspan="1" rowspan="1">Chaetal lobe: shaped</td>
|
||
<td id="78B4FDE422C065D69CF79A1546A805B4" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EASAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="37D720FEE5C78FB43E21FCB0037E5FDA" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EISAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="D8FFE3DE9D85E0026A82B2F9AFC94B4E" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EQSAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="33E222338187A2B8CCB0702661BA9CD4" colspan="1" rowspan="1">Rounded, conical</td>
|
||
<td id="FC01B73C3B1F5CFA787B81A7C76A5C09" colspan="1" rowspan="1">Rounded</td>
|
||
</tr>
|
||
<tr id="D6D88EE41D63AB7396016BA3A063AC97">
|
||
<td id="2C5557067CB6B40BCAC321FBC081161D" colspan="1" rowspan="1">Aciculae: shape; colour</td>
|
||
<td id="E37E8C1E5E0CACB8A6F25C7AEB317EA4" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ECTAK" xlink_title="no information available">NIA</abbrev>
|
||
; black
|
||
</td>
|
||
<td id="C59A3249D2EBD51D8D157E18A42D7640" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EKTAK" xlink_title="no information available">NIA</abbrev>
|
||
; black
|
||
</td>
|
||
<td id="6BBEBE43F6D75B38F5F76B44A8E64CFA" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ESTAK" xlink_title="no information available">NIA</abbrev>
|
||
; oblique
|
||
</td>
|
||
<td id="732B2F2E7405D70C97B24172D33AA328" colspan="1" rowspan="1">Blunt, black with paler tips</td>
|
||
<td id="1C93CD0E2CFACD65ECAE0DB3EBC1C8D7" colspan="1" rowspan="1">Blunt; black and translucent at distal end</td>
|
||
</tr>
|
||
<tr id="B99D87764F6C14259F7AD760DDB95C55">
|
||
<td id="813EF50A0BFA6EE610A4DCC1163A5C9A" colspan="1" rowspan="1">
|
||
Subacicular limbate chaetae: (
|
||
<abbrev id="ABBRID0EBUAK" xlink_title="present/absent">p / a</abbrev>
|
||
); distribution
|
||
</td>
|
||
<td id="203B2A666C9D40C28396ED4C2F8DE3D9" colspan="1" rowspan="1">
|
||
Present;
|
||
<abbrev id="ABBRID0EJUAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="688608BB14739817F3B1BA57643976A5" colspan="1" rowspan="1">Present; all chaetigers</td>
|
||
<td id="97E3767A2D6484EA0DD374E6999BC77A" colspan="1" rowspan="1">Present; posterior chaetigers</td>
|
||
<td id="A3C394C15509CF220EE96967878CCCFA" colspan="1" rowspan="1">Present; all chaetigers</td>
|
||
<td id="629D44652E7598FD3958EC29C07E1806" colspan="1" rowspan="1">Present; in median and posterior chaetigers</td>
|
||
</tr>
|
||
<tr id="1F841E198C78A5E98CA354C7BB0E3BE1">
|
||
<td id="F4F4E88E28684747A1E343927945BD9A" colspan="1" rowspan="1">Pectinate chaetae: number of type *</td>
|
||
<td id="5AD0F945D71F2D73CB58B8855F0CBA10" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EAVAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="8B287FC095946EC7E8FC34FDF694CB02" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EIVAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="B679AB481750CA86B3149C6D41AF9D10" colspan="1" rowspan="1">2</td>
|
||
<td id="66C03AB1534C175BAD96D6263D84D818" colspan="1" rowspan="1">2</td>
|
||
<td id="876E7BFEFCA5E985A925A9211F837B47" colspan="1" rowspan="1">5</td>
|
||
</tr>
|
||
<tr id="06FBE9E7F398E4DEBAAEE5F5BE2C1F52">
|
||
<td id="277253AB5CD36A05FDA77B631F977DC0" colspan="1" rowspan="1">Subacicular hook: shape; colour *</td>
|
||
<td id="B5FD748F07F01835EBF7F3A55F5671BC" colspan="1" rowspan="1">
|
||
Unidentate, strongly hooded;
|
||
<abbrev id="ABBRID0E4VAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="315C59312700FB40CD969F1045645EFA" colspan="1" rowspan="1">
|
||
Bidentate;
|
||
<abbrev id="ABBRID0EEWAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="EDFF8A8250B41E95FA07748DA5B04F50" colspan="1" rowspan="1">Bidentate, close-fitting hood; dark brown</td>
|
||
<td id="1310DB5D130D4D4AA52150DED8BF7926" colspan="1" rowspan="1">
|
||
Bidentate;
|
||
<abbrev id="ABBRID0EOWAK" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="55F761E27FC9660116EB7A921C6643F4" colspan="1" rowspan="1">Bidentate; light brown and translucent at distal end</td>
|
||
</tr>
|
||
<tr id="05F93368B7816EAA08158F42DA52F0EC">
|
||
<td id="BA2C0DFCCA300E4F89FB66D02D04BBDB" colspan="1" rowspan="1">Subacicular hook: start chaetiger *</td>
|
||
<td id="49D94CDE6F95A75A678683D30981C2BA" colspan="1" rowspan="1">50</td>
|
||
<td id="FA4225FAE9295CC15F4594CB293EA424" colspan="1" rowspan="1">26–35</td>
|
||
<td id="1C021C558AF47E8BA5D22737368BB821" colspan="1" rowspan="1">40</td>
|
||
<td id="6C0127DBB0A25AEEC7EB07F85624EDF9" colspan="1" rowspan="1">33–72</td>
|
||
<td id="0D8D6346364D566DC2731C2DC2AD3990" colspan="1" rowspan="1">22 (22–46)</td>
|
||
</tr>
|
||
<tr id="886E06B7CAE480090A60D0572B8E957E">
|
||
<td id="6710F3D99AF6F9A4C0ED4AFE5C6F1A8A" colspan="1" rowspan="1">Subacicular hook: distribution</td>
|
||
<td id="4FAAE8C9D421F82A411CC53CC476D32C" colspan="1" rowspan="1">Continuous</td>
|
||
<td id="7881C546397800080F00A75789D78C58" colspan="1" rowspan="1">Continuous</td>
|
||
<td id="A02B3D570120AB742136AD59BBC4DBF9" colspan="1" rowspan="1">Continuous</td>
|
||
<td id="521125170E5E16569254237A1761DEE8" colspan="1" rowspan="1">Continuous</td>
|
||
<td id="1B1A68FC9C5A23A51FA453397D262244" colspan="1" rowspan="1">Continuous</td>
|
||
</tr>
|
||
</tbody>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="E4081B73B4D2B21D9F5AE082D16556BA">
|
||
<taxonomicName id="10DAC4562A8DE16FB03BAB5213B2130D" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="835265C80F2D29ACAC8BB56B8E51337C" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D8AA0C15FE856A86F41E35DC1BA9B442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="156635DC77C5240C22A01EE7B3946C53" authorityName="Pillai" authorityYear="1958" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="borradailei">
|
||
<emphasis id="5C83866B35B4762F9B6412934C3D019F" italics="true">M. borradailei</emphasis>
|
||
</taxonomicName>
|
||
by the number of branchial filaments, shape of the subacicular hooks, chaetiger where the branchiae and subacicular hook occur, and the shape of postchaetal lobe in the anterior region.
|
||
<taxonomicName id="2C477E842A6650904A4693461787092D" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="DABFC6A5CB9FEB3746984CC5CF0DD679" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0723FCB2F3CB761AFB6132F50A11803A" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(TL: 52 (20–91) mm) has a maximum of eight branchial filaments whereas
|
||
<taxonomicName id="555C046EA2D375443BD69A9E1465BDC6" authorityName="Pillai" authorityYear="1958" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="borradailei">
|
||
<emphasis id="5EAAE4BDB2F0D80E01745BE44D7C6F28" italics="true">M. borradailei</emphasis>
|
||
</taxonomicName>
|
||
(TL:
|
||
<quantity id="7E8B2B638450754F4DA776D6C9B110BA" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="8.0" metricValueMin="1.0" unit="mm" value="4.5" valueMax="8.0" valueMin="1.0">1–8 mm</quantity>
|
||
) has up to 20 branchial filaments. The subacicular hook of
|
||
<taxonomicName id="F78610B68A1FB405B062F2B9A897C467" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="FAF684A2F7438894842DFB90DE8E0C5E" italics="true">M. ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8D711A31E04A1C694B8C110C78914075" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is bidentate and occurs from chaetiger 22 (22–46) onwards while
|
||
<taxonomicName id="0CE9964756BBCC9E479F0A52E29EB661" authorityName="Pillai" authorityYear="1958" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="borradailei">
|
||
<emphasis id="211036023350B66DEF095CEA6AA98308" italics="true">M. borradailei</emphasis>
|
||
</taxonomicName>
|
||
has a strongly hooded unidentate hook that occur from chaetiger 50 onwards.
|
||
<taxonomicName id="26B34569F2620A399F0D2F0C969AF0DC" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="1DD71CD50431C782E75A15658EEF832D" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2999694AE43565080421F6F3190399FA" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has rounded postchaetal lobe in anterior region, while
|
||
<taxonomicName id="9EEDF074F3E2A76C4446CEDA4C7ADD5D" authorityName="Pillai" authorityYear="1958" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="borradailei">
|
||
<emphasis id="37D6E090BA8343E9F00321FD015FC632" italics="true">M. borradailei</emphasis>
|
||
</taxonomicName>
|
||
has sub-conical shaped postchaetal lobes in the anterior region. The original description of
|
||
<taxonomicName id="F5B01BFE8E32F48D258368CCD3DFE7BC" authorityName="Pillai" authorityYear="1958" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="borradailei">
|
||
<emphasis id="888F1BFAA59463F04C8029EC3EBD1F5B" italics="true">M. borradailei</emphasis>
|
||
</taxonomicName>
|
||
makes it challenging to undertake a detailed morphological comparison and additional material from the type locality (
|
||
<collectingCountry id="A942AEA793F11DA66ACA9D4670BB73F6" name="Sri Lanka">Sri Lanka</collectingCountry>
|
||
) needs to be collected and redescribed.
|
||
</paragraph>
|
||
<paragraph id="08DB2EAE2B4709E2FF0D011DE6B975D2">
|
||
The new species can also be differentiated from
|
||
<taxonomicName id="3AE544C51AE6136CE91406EF88F7DFE5" authorityName="Southern" authorityYear="1921" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="gravelyi">
|
||
<emphasis id="14A37E460E3FDF892FF4745CAA2156A7" italics="true">M. gravelyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3DA0333C60D41AACEBE4282539586557" authorityName="Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby" authorityYear="2020" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="madrasi">
|
||
<emphasis id="0215A6170C4B9FA0B66438DEF00780D5" italics="true">M. madrasi</emphasis>
|
||
</taxonomicName>
|
||
by the absence of eyes, number of types of pectinate chaetae, number of branchial filaments, chaetiger where subacicular hooks begin and the length of the pygidial cirri.
|
||
<taxonomicName id="BF07A86ED9D654C57668CB72438E9E16" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="A9936AB96B4712C02AF70E050EE9C224" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6BB64FFF194E363BEE027E4616297E57" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has no eyes, while both
|
||
<taxonomicName id="6A46633BC4ADA37B5A1B550B4AEB9810" authorityName="Southern" authorityYear="1921" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="gravelyi">
|
||
<emphasis id="38D6E1CEFFDA6503B55D3CDC1154D5DD" italics="true">M. gravelyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="CCF5A85D43D9961FF6C16AA74CDD0D8D" authorityName="Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby" authorityYear="2020" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="madrasi">
|
||
<emphasis id="16C842D299EE0428797014B86AEE1A0E" italics="true">M. madrasi</emphasis>
|
||
</taxonomicName>
|
||
have a pair of eyes.
|
||
<taxonomicName id="33923AB5BD25950D7965A3B6AA3F38D4" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="D252439F50587D81D28514E76A460806" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2A38C8D092747ADFFE5FF69AA9FC17AE" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has
|
||
<specimenCount id="3561725D22E1AE91FD8F0797C2202B8D" count="5" type="generic" typeStatus="types">five types</specimenCount>
|
||
of pectinate chaetae (types 1, 3, 4, 5, 7), whereas
|
||
<taxonomicName id="40E64D19F6472EAC0D4DCE328E6A08D6" authorityName="Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby" authorityYear="2020" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="madrasi">
|
||
<emphasis id="1105D48BA27E06D9B7BCB9F0F2C36B94" italics="true">M. madrasi</emphasis>
|
||
</taxonomicName>
|
||
has only two (types 4, 5). While all these species have bidentate hooks, they begin on chaetiger 22 (22–46) in
|
||
<taxonomicName id="F905CB47EE0D8EBE3716D4A7332C2077" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="9B00B102DA0076884FDB8BA393FAC692" italics="true">M. ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="39A5CBB4BABAD67A4C63F0DE746C4CCA" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<abbrev id="ABBRID0EZ4AK" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
: 4.5 (2.25–6.3) mm),
|
||
<quantity id="7F7D25435258CFD8F22EEAB800E2FA00" metricMagnitude="-1" metricUnit="m" metricValue="7.747" metricValueMax="8.89" metricValueMin="6.604" unit="in" value="30.5" valueMax="35.0" valueMin="26.0">26–35 in</quantity>
|
||
<taxonomicName id="BBEA5CD1864727D94DBFA89F67D810D6" authorityName="Southern" authorityYear="1921" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="gravelyi">
|
||
<emphasis id="61833AC8C09064EF0EFA102EF0911B24" italics="true">M. gravelyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="F143DAE4D5DBDF5A19A4CC58406C6E98" metricMagnitude="0" metricUnit="m" metricValue="1.3335" metricValueMax="1.8288" metricValueMin="0.8382" unit="in" value="52.5" valueMax="72.0" valueMin="33.0">33–72 in</quantity>
|
||
<taxonomicName id="41908113A18ECBDFECD727C944CB341E" authorityName="Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby" authorityYear="2020" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="madrasi">
|
||
<emphasis id="700F658BCFC9078EEA7B11B4E3ED2577" italics="true">M. madrasi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<abbrev id="ABBRID0ET5AK" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
: 6 (4–9) mm).
|
||
<taxonomicName id="FD2BEBD183104843B305458D84019444" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="506CA1B818A181A8371D71329797CE6D" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="09F7FE719F5D16CA5D164C32976054C1" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has short and long pairs of pygidial cirri attached to the pygidium, whereas
|
||
<taxonomicName id="FB4C2505B903A57DEFD1C2A3D05D7911" authorityName="Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby" authorityYear="2020" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="madrasi">
|
||
<emphasis id="94DAD4120C350E314EBD2C7EEA9487E9" italics="true">M. madrasi</emphasis>
|
||
</taxonomicName>
|
||
only has one pair of short pygidial cirri.
|
||
</paragraph>
|
||
<paragraph id="9414AC3ACDB9ABCAC9E10933E0B4EABE">
|
||
<taxonomicName id="52E70E92A9B152CB2E3BD3898C23CA46" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="286E0BFADC243F4B1A6BC5D9353F4773" italics="true">Marphysa ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BC9C788ABA55BCF85E378A374C29B70F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="BE49494657714016AED99DAC740F6F77" authorityName="Glasby & Hutchings" authorityYear="2010" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="fauchaldi">
|
||
<emphasis id="713CE9BE01A876B8AFAB66592EFDDD00" italics="true">M. fauchaldi</emphasis>
|
||
</taxonomicName>
|
||
by the absence of peduncle in prostomial appendages, the chaetiger on which the branchiae and subacicular hook occur and the distribution of subacicular limbate chaetae. Subacicular hooks and branchiae of
|
||
<taxonomicName id="A71DFD59AA4EDBCD4F79DAA5CC915ACB" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="63984CCE9B83E7706528847D0B586840" italics="true">M. ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A3E85DB7AA1FD21DA4FB81494B44EF25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(TL: 52 (20–91) mm) have a wide range variation of chaetiger where they begin; from chaetiger 22 (22–46) and 20 (11–65), respectively compared to
|
||
<taxonomicName id="B3A844695D695E289A4E3C1E7A75A4FB" authorityName="Glasby & Hutchings" authorityYear="2010" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="fauchaldi">
|
||
<emphasis id="7C0B94F20ECE687BF191F168F97E0813" italics="true">M. fauchaldi</emphasis>
|
||
</taxonomicName>
|
||
(TL: 190 (78–155) mm); they begin from chaetiger 40 (31–50) and 31 (22–32), respectively. The subacicular limbate chaetae in
|
||
<taxonomicName id="17F56D2497926541A563EAD39744F205" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="ibaiensis" status="sp. nov.">
|
||
<emphasis id="16C226A4943BE561B5BE073E6F365456" italics="true">M. ibaiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CDC5E4FF6EB1DF4D96FED6D969A45135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
occur from mid-chaetigers onwards whereas in
|
||
<taxonomicName id="ADAC77B7B61B39E92FE1719C943E780D" authorityName="Glasby & Hutchings" authorityYear="2010" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="fauchaldi">
|
||
<emphasis id="E2672BBEF05D9FD8FA823C2EE4AFE94D" italics="true">M. fauchaldi</emphasis>
|
||
</taxonomicName>
|
||
, they are restricted to posterior chaetigers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |