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<document id="ADE4506BC8662EF197B86FEEE39C3831" ID-DOI="10.5281/zenodo.4619638" ID-ISSN="2345-7600" ID-RefBankID="FF8BFFD82661FFFFFFA5FF89FFB0FFE1" ID-Zenodo-Dep="4619638" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616096590396" checkinUser="carolina" docAuthor="Daugeron, Christophe &amp; Grootaert, Patrick" docDate="2005" docId="7E7087CC0147FF88FC38FF583B445930" docLanguage="en" docName="RafflesBZool.53.2.211-220.pdf" docOrigin="Raffles Bulletin of Zoology 53 (2)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Empis (Coptophlebia) producta Daugeron &amp; Grootaert 2005, new species" docType="treatment" docVersion="3" lastPageNumber="214" masterDocId="8249FFB40145FF8BFF97FF993F0D5848" masterDocTitle="Empidine Dance Flies From Singapore And Southern Malaysia (Diptera: Empididae: Empidinae)" masterLastPageNumber="220" masterPageNumber="211" pageNumber="213" updateTime="1698891084545" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9D83039A109766D44CF2B2142F23C536">Empidine Dance Flies From Singapore And Southern Malaysia (Diptera: Empididae: Empidinae)</mods:title>
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<mods:namePart id="19E20BEAC375FE2F111F7D0BBC688B00">Daugeron, Christophe</mods:namePart>
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<mods:namePart id="E680F79BB33FEBC0D8600AAC2DD8C5AB">Grootaert, Patrick</mods:namePart>
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<mods:title id="A1B875E17BD74069100709F4891F6A99">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="AE1EE217428F2C581C8AFF61044286E4">
<mods:date id="38F56D8AD39995B9B85C8022D97DAF5C">2005</mods:date>
<mods:detail id="E220CE48D9E53EC26EE19EB30E3548DB" type="pubDate">
<mods:number id="802A6E279C89C5AC351CA3A6081CC347">2005-12-31</mods:number>
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<mods:number id="A4E1DD9FD56A4CA350F7F4351B986E65">53</mods:number>
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<treatment id="7E7087CC0147FF88FC38FF583B445930" LSID="urn:lsid:plazi:treatment:7E7087CC0147FF88FC38FF583B445930" httpUri="http://treatment.plazi.org/id/7E7087CC0147FF88FC38FF583B445930" lastPageId="3" lastPageNumber="214" pageId="2" pageNumber="213">
<subSubSection id="BEC365510147FF89FC38FF583AAD5890" box="[943,1440,192,217]" pageId="2" pageNumber="213" type="nomenclature">
<paragraph id="F66636DA0147FF89FC38FF583AAD5890" blockId="2.[943,1440,192,248]" box="[943,1440,192,217]" pageId="2" pageNumber="213">
<heading id="AD2E81B60147FF89FC38FF583AAD5890" bold="true" box="[943,1440,192,217]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="213" reason="2">
<emphasis id="C4ADEAC80147FF89FC38FF583AAD5890" bold="true" box="[943,1440,192,217]" pageId="2" pageNumber="213">
<taxonomicName id="31D94D590147FF89FC38FF583A035891" ID-CoL="8BZ7P" authority="Daugeron &amp; Grootaert, 2005" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[943,1294,192,217]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="213" phylum="Arthropoda" rank="species" species="producta" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="C4ADEAC80147FF89FC38FF583CF85891" bold="true" box="[943,1013,193,217]" italics="true" pageId="2" pageNumber="213">Empis</emphasis>
(
<emphasis id="C4ADEAC80147FF89FB91FF583B975891" bold="true" box="[1030,1178,193,217]" italics="true" pageId="2" pageNumber="213">Coptophlebia</emphasis>
)
<emphasis id="C4ADEAC80147FF89FB3CFF583A035891" bold="true" box="[1195,1294,193,217]" italics="true" pageId="2" pageNumber="213">producta</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="DF9E57B30147FF89FA8BFF593AAD5890" box="[1308,1440,192,216]" pageId="2" pageNumber="213" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BEC365510147FF89FBB7FF783BE858B0" box="[1056,1253,225,248]" pageId="2" pageNumber="213" type="description">
<paragraph id="F66636DA0147FF89FBB7FF783BE858B0" blockId="2.[943,1440,192,248]" box="[1056,1253,225,248]" pageId="2" pageNumber="213">
(
<figureCitation id="6EE22A5F0147FF89FBBFFF783B8558B0" box="[1064,1160,225,248]" captionStart="Fig" captionStartId="1.[113,149,2063,2084]" captionTargetBox="[117,1448,682,1971]" captionTargetId="figure-9@1.[102,1460,631,1988]" captionTargetPageId="1" captionText="Fig. 1. Male fore leg in frontal view: A, tibia and first tarsomere of E. (C.) neesoonensis; B, tibia and tarsus of E. (C.) producta; C, tibia and first tarsomere of E. (C.) tortuosa; D, tibia and first tarsomere of E. (C.) verruca." figureDoi="http://doi.org/10.5281/zenodo.4619640" httpUri="https://zenodo.org/record/4619640/files/figure.png" pageId="2" pageNumber="213">Figs. 1B</figureCitation>
,
<figureCitation id="6EE22A5F0147FF89FB02FF783BA958B0" box="[1173,1188,225,248]" captionStart="Fig" captionStartId="3.[112,148,2081,2102]" captionTargetBox="[117,772,1073,2005]" captionTargetId="figure-20@3.[96,795,1048,2030]" captionTargetPageId="3" captionText="Fig. 3. Habitus of E. (C.) producta, male." figureDoi="http://doi.org/10.5281/zenodo.4619644" httpUri="https://zenodo.org/record/4619644/files/figure.png" pageId="2" pageNumber="213">3</figureCitation>
,
<figureCitation id="6EE22A5F0147FF89FB25FF783BCC58B0" box="[1202,1217,225,248]" captionStart="Fig" captionStartId="4.[137,173,2093,2114]" captionTargetBox="[146,1483,720,2009]" captionTargetId="figure-25@4.[108,1518,662,2069]" captionTargetPageId="4" captionText="Fig. 4. Male hypopygium of E. (C.) producta, lateral view. Scale = 4.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4619646" httpUri="https://zenodo.org/record/4619646/files/figure.png" pageId="2" pageNumber="213">4</figureCitation>
,
<figureCitation id="6EE22A5F0147FF89FB59FF783BD158B0" box="[1230,1244,225,248]" captionStart="Fig" captionStartId="5.[112,148,2062,2083]" captionTargetBox="[112,753,1070,2038]" captionTargetPageId="5" captionText="Fig. 5. Photographs showing the typical coastal habitat of E. (C.) producta: mangroves of Singapore (Pulau Ubin)." figureDoi="http://doi.org/10.5281/zenodo.4619648" httpUri="https://zenodo.org/record/4619648/files/figure.png" pageId="2" pageNumber="213">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BEC365510147FF89FCD4FEB93C965A54" pageId="2" pageNumber="213" type="materials_examined">
<paragraph id="F66636DA0147FF89FCD4FEB93C845927" blockId="2.[835,1474,288,367]" pageId="2" pageNumber="213">
<emphasis id="C4ADEAC80147FF89FCD4FEB93B19597D" bold="true" box="[835,1044,288,309]" italics="true" pageId="2" pageNumber="213">Material examined. </emphasis>
<materialsCitation id="46B13C870147FF89FB8EFEB93C845927" collectingDate="2002-12-11" collectionCode="ZRC" collectorName="P. Grootaert" country="Singapore" location="Singapore" pageId="2" pageNumber="213" specimenCount="1" typeStatus="holotype">
<typeStatus id="296288780147FF89FB8EFEB93B7F597D" box="[1049,1138,288,309]" pageId="2" pageNumber="213" type="holotype">Holotype</typeStatus>
<emphasis id="C4ADEAC80147FF89FBE0FEB83B72597E" box="[1143,1151,289,310]" italics="true" pageId="2" pageNumber="213">-</emphasis>
male,
<collectingCountry id="8ECE764A0147FF89FB2AFEB93A2C597D" box="[1213,1313,288,309]" name="Singapore" pageId="2" pageNumber="213">Singapore</collectingCountry>
, Pulau [= Island] Ubin, Chek Jawa, sample 22059, coll.
<collectorName id="5B2C530C0147FF89FB57FEA43A37591A" box="[1216,1338,317,338]" pageId="2" pageNumber="213">P. Grootaert</collectorName>
,
<date id="8267101A0147FF89FAD2FEA43ACF591A" box="[1349,1474,317,338]" pageId="2" pageNumber="213" value="2002-12-11">
<collectingDate id="9223E9F20147FF89FAD2FEA43ACF591A" box="[1349,1474,317,338]" pageId="2" pageNumber="213" value="2002-12-11">11 Dec.2002</collectingDate>
</date>
(
<collectionCode id="90C8AE1F0147FF89FCDDFEC33C735927" box="[842,894,346,367]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="213">ZRC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F66636DA0147FF89FCD4FE0D3C965A54" blockId="2.[835,1474,404,540]" pageId="2" pageNumber="213">
<materialsCitation id="46B13C870147FF89FCD4FE0D3BA959E1" box="[835,1188,404,425]" pageId="2" pageNumber="213" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="296288780147FF89FCD4FE0D3CAF59E1" box="[835,930,404,425]" pageId="2" pageNumber="213" type="paratype">Paratypes</typeStatus>
<emphasis id="C4ADEAC80147FF89FC3EFE0D3CB859E1" box="[937,949,404,425]" italics="true" pageId="2" pageNumber="213"></emphasis>
<specimenCount id="E0DFFD530147FF89FC2CFE0D3B0A59E1" box="[955,1031,404,425]" pageId="2" pageNumber="213" type="male">3 males</specimenCount>
with same data
</materialsCitation>
,
<materialsCitation id="46B13C870147FF89FB38FE0D3CEC59AA" collectingDate="2002-12-11" collectionCode="ZRC" collectorName="P. Grootaert" country="Singapore" location="Pulau" municipality="Ubin" pageId="2" pageNumber="213" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="E0DFFD530147FF89FB38FE0D3BFE59E1" box="[1199,1267,404,425]" pageId="2" pageNumber="213" type="male">1 male</specimenCount>
,
<collectingCountry id="8ECE764A0147FF89FB69FE0D3A6E59E1" box="[1278,1379,404,425]" name="Singapore" pageId="2" pageNumber="213">Singapore</collectingCountry>
,
<location id="F30660010147FF89FAF9FE0D3AA859E1" LSID="urn:lsid:plazi:treatment:7E7087CC0147FF88FC38FF583B445930:F30660010147FF89FAF9FE0D3AA859E1" box="[1390,1445,404,425]" country="Singapore" municipality="Ubin" name="Pulau" pageId="2" pageNumber="213">Pulau</location>
[= Island]
<collectingMunicipality id="1602ACA00147FF89FC05FE293CCB598D" box="[914,966,432,453]" pageId="2" pageNumber="213">Ubin</collectingMunicipality>
,
<date id="8267101A0147FF89FC43FE293B54598D" box="[980,1113,432,453]" pageId="2" pageNumber="213" value="2002-12-11">
<collectingDate id="9223E9F20147FF89FC43FE293B54598D" box="[980,1113,432,453]" pageId="2" pageNumber="213" value="2002-12-11">11 Dec.2002</collectingDate>
</date>
, mangrove, sample 23119, coll.
<collectorName id="5B2C530C0147FF89FA39FE293C9059AA" pageId="2" pageNumber="213">P. Grootaert</collectorName>
(
<collectionCode id="90C8AE1F0147FF89FC3FFE543CD659AA" box="[936,987,461,482]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="213">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="46B13C870147FF89FC7EFE543C955A54" collectingDate="2003-11-19" collectionCode="ZRC, MNHN" collectorName="P. Grootaert" country="Singapore" location="Pulau" municipality="Ubin" pageId="2" pageNumber="213" specimenCount="35" specimenCount-male="35" typeStatus="paratype">
<specimenCount id="E0DFFD530147FF89FC7EFE543B3359AA" box="[1001,1086,461,482]" pageId="2" pageNumber="213" type="male">35 males</specimenCount>
,
<collectingCountry id="8ECE764A0147FF89FBD2FE543BAB59AA" box="[1093,1190,461,482]" name="Singapore" pageId="2" pageNumber="213">Singapore</collectingCountry>
,
<location id="F30660010147FF89FB39FE543BEE59AA" LSID="urn:lsid:plazi:treatment:7E7087CC0147FF88FC38FF583B445930:F30660010147FF89FB39FE543BEE59AA" box="[1198,1251,461,482]" country="Singapore" municipality="Ubin" name="Pulau" pageId="2" pageNumber="213">Pulau</location>
[= Island]
<collectingMunicipality id="1602ACA00147FF89FAD1FE543A7A59AA" box="[1350,1399,461,482]" pageId="2" pageNumber="213">Ubin</collectingMunicipality>
, sample 23080, coll.
<collectorName id="5B2C530C0147FF89FC52FE733B4E59B7" box="[965,1091,490,511]" pageId="2" pageNumber="213">P. Grootaert</collectorName>
,
<date id="8267101A0147FF89FBC7FE733BDE59B7" box="[1104,1235,490,511]" pageId="2" pageNumber="213" value="2003-11-19">
<collectingDate id="9223E9F20147FF89FBC7FE733BDE59B7" box="[1104,1235,490,511]" pageId="2" pageNumber="213" value="2003-11-19">19 Nov.2003</collectingDate>
</date>
(
<collectionCode id="90C8AE1F0147FF89FB73FE733A1A59B7" box="[1252,1303,490,511]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="213">ZRC</collectionCode>
, KBIN-IRScNB,
<collectionCode id="90C8AE1F0147FF89FCD4FD9E3C9C5A54" box="[835,913,519,540]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="213">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BEC365510147FF89FCD4FDDD3CB35AD3" pageId="2" pageNumber="213" type="etymology">
<paragraph id="F66636DA0147FF89FCD4FDDD3CB35AD3" blockId="2.[835,1474,580,667]" pageId="2" pageNumber="213">
<emphasis id="C4ADEAC80147FF89FCD4FDDD3CDF5A14" bold="true" box="[835,978,580,604]" italics="true" pageId="2" pageNumber="213">Etymology. </emphasis>
From the Latin word
<emphasis id="C4ADEAC80147FF89FB2FFDDD3A2E5A13" box="[1208,1315,580,603]" italics="true" pageId="2" pageNumber="213">productus</emphasis>
in reference to the dorsal part of the epandrium that is considerably produced posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="BEC365510147FF89FCD4FD5D3CE25B33" pageId="2" pageNumber="213" type="diagnosis">
<paragraph id="F66636DA0147FF89FCD4FD5D3CE25B33" blockId="2.[835,1474,708,891]" pageId="2" pageNumber="213">
<emphasis id="C4ADEAC80147FF89FCD4FD5D3CC75A94" bold="true" box="[835,970,708,732]" italics="true" pageId="2" pageNumber="213">Diagnosis. </emphasis>
Medium-sized blackish species with all bristles black, fore tibia deformed with row of anterodorsal spinelike bristles on basal third, first fore tarsomere elongated, following three tarsomeres swollen with numerous long, fine dorsal bristles, wing with visible veins, epandrium produced posterodorsally.
</paragraph>
</subSubSection>
<subSubSection id="BEC365510147FF88FCD4FC3D3E3A5B90" lastPageId="3" lastPageNumber="214" pageId="2" pageNumber="213" type="description">
<paragraph id="F66636DA0147FF89FCD4FC3D3ACF5B93" blockId="2.[835,1474,932,987]" pageId="2" pageNumber="213">
<emphasis id="C4ADEAC80147FF89FCD4FC3D3B795BF3" box="[835,1140,932,956]" italics="true" pageId="2" pageNumber="213">
<emphasis id="C4ADEAC80147FF89FCD4FC3D3B3E5BF4" bold="true" box="[835,1075,932,956]" italics="true" pageId="2" pageNumber="213">Description. Male </emphasis>
Head
</emphasis>
. Occiput dark brown blackish, antenna black, first flagellomere somewhat conical, second
</paragraph>
<caption id="A2A666520147FF89FF1FF7B43C92500A" ID-DOI="http://doi.org/10.5281/zenodo.4619642" ID-Zenodo-Dep="4619642" box="[136,927,2093,2114]" httpUri="https://zenodo.org/record/4619642/files/figure.png" pageId="2" pageNumber="213" startId="2.[136,172,2093,2114]" targetBox="[144,1464,1104,2005]" targetPageId="2">
<paragraph id="F66636DA0147FF89FF1FF7B43C92500A" blockId="2.[136,927,2093,2114]" box="[136,927,2093,2114]" pageId="2" pageNumber="213">
Fig. 2. Male hypopygium of
<taxonomicName id="31D94D590147FF89FE32F7B43D62500A" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[421,623,2093,2114]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="213" phylum="Arthropoda" rank="species" species="neesoonensis" subGenus="Coptophlebia">
<emphasis id="C4ADEAC80147FF89FE32F7B43EB6500A" box="[421,443,2093,2114]" italics="true" pageId="2" pageNumber="213">E.</emphasis>
(
<emphasis id="C4ADEAC80147FF89FE5EF7B43EED500A" box="[457,480,2093,2114]" italics="true" pageId="2" pageNumber="213">C.</emphasis>
)
<emphasis id="C4ADEAC80147FF89FE79F7B43D62500A" box="[494,623,2093,2114]" italics="true" pageId="2" pageNumber="213">neesoonensis</emphasis>
</taxonomicName>
, lateral view. Scale = 4.2 mm.
</paragraph>
</caption>
<paragraph id="F66636DA0146FF88FFE6FF583E525BD0" blockId="3.[113,753,193,920]" pageId="3" pageNumber="214">
and third flagellomeres aristiform. Palpus brown, labrum length 1.5x head height.
<emphasis id="C4ADEAC80146FF88FE07FF7B3ED258B1" box="[400,479,226,249]" italics="true" pageId="3" pageNumber="214">Thorax</emphasis>
blackish. Dorsocentrals uniserial ending with 2 strong, long bristles in prescutellar depression.
<emphasis id="C4ADEAC80146FF88FF66FEBB3E295971" box="[241,292,290,313]" italics="true" pageId="3" pageNumber="214">Legs</emphasis>
(
<figureCitation id="6EE22A5F0146FF88FEA5FEB83E885970" box="[306,389,289,312]" captionStart="Fig" captionStartId="1.[113,149,2063,2084]" captionTargetBox="[117,1448,682,1971]" captionTargetId="figure-9@1.[102,1460,631,1988]" captionTargetPageId="1" captionText="Fig. 1. Male fore leg in frontal view: A, tibia and first tarsomere of E. (C.) neesoonensis; B, tibia and tarsus of E. (C.) producta; C, tibia and first tarsomere of E. (C.) tortuosa; D, tibia and first tarsomere of E. (C.) verruca." figureDoi="http://doi.org/10.5281/zenodo.4619640" httpUri="https://zenodo.org/record/4619640/files/figure.png" pageId="3" pageNumber="214">Fig. 1B</figureCitation>
). Fore tibia deformed with row of anterodorsal spine-like bristles on basal third ending in 2-4 longer bristles (well visible in anterior view); first fore tarsomere elongated, almost as long as the following four tarsomeres together, with 1 distinct anterolateral and posterodorsal bristles at base and at tip respectively; following three tarsomeres swollen with numerous long rather fine dorsal bristles, 1 strong, long ventral bristle. Mid tibia with 3 strong, long ventral and dorsal bristles; first mid tarsomere with 1 strong, long posterodorsal bristle at base and at tip. Hind tibia and tarsus with several strong, long postero- and anterodorsal bristles, hind tarsus somewhat thick.
<emphasis id="C4ADEAC80146FF88FD21FD1B3DFD5AD1" box="[694,752,642,665]" italics="true" pageId="3" pageNumber="214">Wing</emphasis>
(length =
<quantity id="31219B3F0146FF88FF4CFD383E3D5AF0" box="[219,304,673,696]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="214" unit="mm" value="2.4">2.4 mm</quantity>
) feebly tinged with brown, all veins well visible except M1, A1 abbreviated. Halter dark brown.
<emphasis id="C4ADEAC80146FF88FFE6FD7B3FDA5AB1" box="[113,215,738,761]" italics="true" pageId="3" pageNumber="214">Abdomen</emphasis>
dark brown, paler, with distinct short bristles at base.
<emphasis id="C4ADEAC80146FF88FF3AFC9B3E3B5B51" box="[173,310,770,793]" italics="true" pageId="3" pageNumber="214">Hypopygium</emphasis>
(
<figureCitation id="6EE22A5F0146FF88FED4FC983E895B50" box="[323,388,769,792]" captionStart="Fig" captionStartId="4.[137,173,2093,2114]" captionTargetBox="[146,1483,720,2009]" captionTargetId="figure-25@4.[108,1518,662,2069]" captionTargetPageId="4" captionText="Fig. 4. Male hypopygium of E. (C.) producta, lateral view. Scale = 4.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4619646" httpUri="https://zenodo.org/record/4619646/files/figure.png" pageId="3" pageNumber="214">Fig. 4</figureCitation>
). Epandrium rather truncate at tip, projected posterodorsally, with row of very strong, long bristles on the dorsal margin, several shorter, finer bristles at tip. Hypandrium somewhat rounded at tip. Phallus rather long, narrowed at tip.
</paragraph>
<paragraph id="F66636DA0146FF88FFE6FC583E3A5B90" blockId="3.[113,311,961,985]" box="[113,311,961,985]" pageId="3" pageNumber="214">
<emphasis id="C4ADEAC80146FF88FFE6FC583FC85B91" bold="true" box="[113,197,961,985]" italics="true" pageId="3" pageNumber="214">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="BEC365510146FF88FCBBFF583B445930" pageId="3" pageNumber="214" type="discussion">
<paragraph id="F66636DA0146FF88FCBBFF583B445930" blockId="3.[812,1452,193,376]" pageId="3" pageNumber="214">
<emphasis id="C4ADEAC80146FF88FCBBFF583CA65891" bold="true" box="[812,939,193,217]" italics="true" pageId="3" pageNumber="214">Remarks. </emphasis>
This species belongs to the
<taxonomicName id="31D94D590146FF88FB75FF583A665890" authorityName="Meijere" authorityYear="1907" box="[1250,1387,193,216]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="214" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="C4ADEAC80146FF88FB75FF583A665890" box="[1250,1387,193,216]" italics="true" pageId="3" pageNumber="214">E. jacobsoni</emphasis>
</taxonomicName>
clade as defined recently (Daugeron &amp; Grootaert, 2005). Within this clade it is closely related to
<taxonomicName id="31D94D590146FF88FB10FE983A265950" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[1159,1323,257,280]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="214" phylum="Arthropoda" rank="species" species="padangensis">
<emphasis id="C4ADEAC80146FF88FB10FE983A265950" box="[1159,1323,257,280]" italics="true" pageId="3" pageNumber="214">E. padangensis</emphasis>
</taxonomicName>
(
<collectingCountry id="8ECE764A0146FF88FAAEFE983AA55950" box="[1337,1448,257,280]" name="Indonesia" pageId="3" pageNumber="214">Indonesia</collectingCountry>
:
<collectingRegion id="341DF8380146FF88FCBBFEB83C855970" box="[812,904,289,312]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="214">Sumatra</collectingRegion>
) with fore tarsus distinctly less rounded than in this species. Both species occur in the same kind of coastal habitat, i.e. in mangroves (
<figureCitation id="6EE22A5F0146FF88FC6EFEF83B365930" box="[1017,1083,353,376]" captionStart="Fig" captionStartId="5.[112,148,2062,2083]" captionTargetBox="[112,753,1070,2038]" captionTargetPageId="5" captionText="Fig. 5. Photographs showing the typical coastal habitat of E. (C.) producta: mangroves of Singapore (Pulau Ubin)." figureDoi="http://doi.org/10.5281/zenodo.4619648" httpUri="https://zenodo.org/record/4619648/files/figure.png" pageId="3" pageNumber="214">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>