197 lines
20 KiB
XML
197 lines
20 KiB
XML
<document id="006303236785BB06757EC80A5B497C2F" ID-DOI="10.5281/zenodo.4619638" ID-ISSN="2345-7600" ID-RefBankID="FF8BFFD82661FFFFFFA5FF89FFB0FFE1" ID-Zenodo-Dep="4619638" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616096590396" checkinUser="carolina" docAuthor="Daugeron, Christophe & Grootaert, Patrick" docDate="2005" docId="7E7087CC0140FF8EFC08FF583BEE507D" docLanguage="en" docName="RafflesBZool.53.2.211-220.pdf" docOrigin="Raffles Bulletin of Zoology 53 (2)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Empis (Coptophlebia) verruca Daugeron & Grootaert 2005, new species" docType="treatment" docVersion="4" lastPageNumber="216" masterDocId="8249FFB40145FF8BFF97FF993F0D5848" masterDocTitle="Empidine Dance Flies From Singapore And Southern Malaysia (Diptera: Empididae: Empidinae)" masterLastPageNumber="220" masterPageNumber="211" pageNumber="216" updateTime="1698891084545" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="5690E6724AA1209C87EDE36ABC243282">
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<mods:title id="FA2B32E0AEBD07CF2A27C9D3061D5228">Empidine Dance Flies From Singapore And Southern Malaysia (Diptera: Empididae: Empidinae)</mods:title>
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<mods:namePart id="41E15481FE962ED202E63E6B4AAF5429">Daugeron, Christophe</mods:namePart>
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<mods:roleTerm id="C98538646A615F813747B6CB6F444A71">Author</mods:roleTerm>
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<mods:namePart id="41A7F77F0BC926F388A649AD0C9A09E3">Grootaert, Patrick</mods:namePart>
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<mods:typeOfResource id="36A4D1F69390685060622F7AA0E3291F">text</mods:typeOfResource>
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<mods:title id="6853919234EE770D789D920F85C98DA3">Raffles Bulletin of Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="2E9104379E1AEADBC0CE5BBDB58B1674">
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<mods:date id="D4D5CDF75DCC8283996F736176453F5A">2005</mods:date>
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<mods:number id="65780F4432CF14F82CFBB5916F297B98">2005-12-31</mods:number>
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<mods:detail id="4C930B95D0BCC1D679721041951BFD21" type="volume">
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<mods:number id="589A046FE1D039A811219AC706E64984">53</mods:number>
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<mods:detail id="55564E7D03CA18091CB93E622D48A1B7" type="issue">
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<mods:number id="05ECB6B2F108FB1F780D56A288E70E87">2</mods:number>
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<mods:start id="420E561C20771496892555F950E1B0BF">211</mods:start>
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<mods:identifier id="DD624715F498F1F450EA3F6B21AC2828" type="DOI">10.5281/zenodo.4619638</mods:identifier>
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<mods:identifier id="32E2175518A56334D55A5FE58E9BDCAA" type="ISSN">2345-7600</mods:identifier>
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<treatment id="7E7087CC0140FF8EFC08FF583BEE507D" ID-DOI="http://doi.org/10.5281/zenodo.4619600" ID-Zenodo-Dep="4619600" LSID="urn:lsid:plazi:treatment:7E7087CC0140FF8EFC08FF583BEE507D" httpUri="http://treatment.plazi.org/id/7E7087CC0140FF8EFC08FF583BEE507D" lastPageNumber="216" pageId="5" pageNumber="216">
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<subSubSection id="BEC365510140FF8EFC08FF583A8E5890" box="[927,1411,192,217]" pageId="5" pageNumber="216" type="nomenclature">
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<paragraph id="F66636DA0140FF8EFC08FF583A8E5890" blockId="5.[927,1411,192,248]" box="[927,1411,192,217]" pageId="5" pageNumber="216">
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<heading id="AD2E81B60140FF8EFC08FF583A8E5890" bold="true" box="[927,1411,192,217]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="216" reason="2">
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<emphasis id="C4ADEAC80140FF8EFC08FF583A8E5890" bold="true" box="[927,1411,192,217]" pageId="5" pageNumber="216">
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<taxonomicName id="31D94D590140FF8EFC08FF583BE25891" ID-CoL="8BZ8Q" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[927,1263,192,217]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="216" phylum="Arthropoda" rank="species" species="verruca" status="sp. nov." subGenus="Coptophlebia">
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<emphasis id="C4ADEAC80140FF8EFC08FF583CE85891" bold="true" box="[927,997,193,217]" italics="true" pageId="5" pageNumber="216">Empis</emphasis>
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(
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<emphasis id="C4ADEAC80140FF8EFC62FF583B845891" bold="true" box="[1013,1161,193,217]" italics="true" pageId="5" pageNumber="216">Coptophlebia</emphasis>
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)
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<emphasis id="C4ADEAC80140FF8EFB0DFF583BE25891" bold="true" box="[1178,1263,193,217]" italics="true" pageId="5" pageNumber="216">verruca</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="DF9E57B30140FF8EFB69FF593A8E5890" box="[1278,1411,192,216]" pageId="5" pageNumber="216" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="BEC365510140FF8EFBB2FF783BBC58B0" box="[1061,1201,225,248]" pageId="5" pageNumber="216" type="description">
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<paragraph id="F66636DA0140FF8EFBB2FF783BBC58B0" blockId="5.[927,1411,192,248]" box="[1061,1201,225,248]" pageId="5" pageNumber="216">
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(
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<figureCitation id="6EE22A5F0140FF8EFBBAFF783B8058B0" box="[1069,1165,225,248]" captionStart="Fig" captionStartId="1.[113,149,2063,2084]" captionTargetBox="[117,1448,682,1971]" captionTargetId="figure-9@1.[102,1460,631,1988]" captionTargetPageId="1" captionText="Fig. 1. Male fore leg in frontal view: A, tibia and first tarsomere of E. (C.) neesoonensis; B, tibia and tarsus of E. (C.) producta; C, tibia and first tarsomere of E. (C.) tortuosa; D, tibia and first tarsomere of E. (C.) verruca." figureDoi="http://doi.org/10.5281/zenodo.4619640" httpUri="https://zenodo.org/record/4619640/files/figure.png" pageId="5" pageNumber="216">Figs. 1D</figureCitation>
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,
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<figureCitation id="6EE22A5F0140FF8EFB0CFF783BA558B0" box="[1179,1192,225,248]" captionStart="Fig" captionStartId="8.[137,173,2091,2112]" captionText="Fig. 8. Male hypopygium of E. (C.) verruca, lateral view. Scale = 4.2 mm." pageId="5" pageNumber="216">8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="BEC365510140FF8EFCBBFEB93C7F59B7" pageId="5" pageNumber="216" type="materials_examined">
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<paragraph id="F66636DA0140FF8EFCBBFEB93A3E591A" blockId="5.[812,1450,288,338]" pageId="5" pageNumber="216">
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<emphasis id="C4ADEAC80140FF8EFCBBFEB93B01597D" bold="true" box="[812,1036,288,309]" italics="true" pageId="5" pageNumber="216">Material examined. –</emphasis>
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<materialsCitation id="46B13C870140FF8EFB81FEB93A3E591A" collectingDate="2000-10-05" collectionCode="ZRC" collectorName="P. Grootaert" country="Singapore" location="Singapore" pageId="5" pageNumber="216" specimenCount="1" typeStatus="holotype">
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<typeStatus id="296288780140FF8EFB81FEB93B7F597D" box="[1046,1138,288,309]" pageId="5" pageNumber="216" type="holotype">Holotype</typeStatus>
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<emphasis id="C4ADEAC80140FF8EFBECFEB83B8E597E" box="[1147,1155,289,310]" italics="true" pageId="5" pageNumber="216">-</emphasis>
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male,
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<collectingCountry id="8ECE764A0140FF8EFB5BFEB93A3E597D" box="[1228,1331,288,309]" name="Singapore" pageId="5" pageNumber="216">Singapore</collectingCountry>
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, Nee Soon, sample 22057, coll.
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<collectorName id="5B2C530C0140FF8EFC63FEA43B63591A" box="[1012,1134,317,338]" pageId="5" pageNumber="216">P. Grootaert</collectorName>
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,
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<date id="8267101A0140FF8EFBEFFEA43BE8591A" box="[1144,1253,317,338]" pageId="5" pageNumber="216" value="2000-10-05">
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<collectingDate id="9223E9F20140FF8EFBEFFEA43BE8591A" box="[1144,1253,317,338]" pageId="5" pageNumber="216" value="2000-10-05">5 Oct.2000</collectingDate>
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</date>
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(
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<collectionCode id="90C8AE1F0140FF8EFB64FEA43A25591A" box="[1267,1320,317,338]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="216">ZRC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="F66636DA0140FF8EFCBBFEEE3C7F59B7" blockId="5.[812,1450,375,511]" pageId="5" pageNumber="216">
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<materialsCitation id="46B13C870140FF8EFCBBFEEE3BDB59C4" box="[812,1238,375,396]" collectionCode="ZRC" pageId="5" pageNumber="216" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<typeStatus id="296288780140FF8EFCBBFEEE3C8459C4" box="[812,905,375,396]" pageId="5" pageNumber="216" type="paratype">Paratypes</typeStatus>
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<emphasis id="C4ADEAC80140FF8EFC1AFEEE3C9459C4" box="[909,921,375,396]" italics="true" pageId="5" pageNumber="216">–</emphasis>
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<specimenCount id="E0DFFD530140FF8EFC09FEEE3CD059C4" box="[926,989,375,396]" pageId="5" pageNumber="216" type="male">1 male</specimenCount>
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with the same data (
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<collectionCode id="90C8AE1F0140FF8EFB0BFEEE3BC259C4" box="[1180,1231,375,396]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="216">ZRC</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="46B13C870140FF8EFB49FEEE3A59598D" collectingDate="2002-12-04" collectionCode="ZRC, MNHN" collectorName="P. Grootaert" country="Singapore" location="Singapore" pageId="5" pageNumber="216" specimenCount="7" specimenCount-female="1" specimenCount-male="6" typeStatus="paratype">
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<specimenCount id="E0DFFD530140FF8EFB49FEEE3A2A59C4" box="[1246,1319,375,396]" pageId="5" pageNumber="216" type="male">6 males</specimenCount>
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and
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<specimenCount id="E0DFFD530140FF8EFAC5FEEE3AAB59C4" box="[1362,1446,375,396]" pageId="5" pageNumber="216" type="female">1 female</specimenCount>
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,
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<collectingCountry id="8ECE764A0140FF8EFCBBFE0D3C9C59E1" box="[812,913,404,425]" name="Singapore" pageId="5" pageNumber="216">Singapore</collectingCountry>
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, Nee Soon, primary rain forest, sample 22054, coll.
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<collectorName id="5B2C530C0140FF8EFA01FE0D3C81598D" pageId="5" pageNumber="216">P. Grootaert</collectorName>
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,
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<date id="8267101A0140FF8EFC0EFE293B01598D" box="[921,1036,432,453]" pageId="5" pageNumber="216" value="2002-12-04">
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<collectingDate id="9223E9F20140FF8EFC0EFE293B01598D" box="[921,1036,432,453]" pageId="5" pageNumber="216" value="2002-12-04">4 Dec.2002</collectingDate>
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</date>
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(
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<collectionCode id="90C8AE1F0140FF8EFB8DFE293B40598D" box="[1050,1101,432,453]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="216">ZRC</collectionCode>
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, KBIN-IRScNB,
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<collectionCode id="90C8AE1F0140FF8EFB69FE293A40598D" box="[1278,1357,432,453]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="216">MNHN</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="46B13C870140FF8EFAF7FE293C6359B7" collectingDate="2002-12-03" collectionCode="ZRC" collectorName="P. Grootaert" country="Singapore" location="Singapore" pageId="5" pageNumber="216" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount id="E0DFFD530140FF8EFAF7FE293AAB598D" box="[1376,1446,432,453]" pageId="5" pageNumber="216" type="male">1 male</specimenCount>
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,
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<collectingCountry id="8ECE764A0140FF8EFCBBFE543C9D59AA" box="[812,912,461,482]" name="Singapore" pageId="5" pageNumber="216">Singapore</collectingCountry>
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, Nee Soon, sample 23102, coll.
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<collectorName id="5B2C530C0140FF8EFB29FE543A3959AA" box="[1214,1332,461,482]" pageId="5" pageNumber="216">P. Grootaert</collectorName>
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,
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<date id="8267101A0140FF8EFAABFE543AA759AA" box="[1340,1450,461,482]" pageId="5" pageNumber="216" value="2002-12-03">
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<collectingDate id="9223E9F20140FF8EFAABFE543AA759AA" box="[1340,1450,461,482]" pageId="5" pageNumber="216" value="2002-12-03">3 Dec.2002</collectingDate>
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</date>
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(
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<collectionCode id="90C8AE1F0140FF8EFCA4FE733C6A59B7" box="[819,871,490,511]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="216">ZRC</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="BEC365510140FF8EFCBBFDB03A4C5A2B" pageId="5" pageNumber="216" type="etymology">
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<paragraph id="F66636DA0140FF8EFCBBFDB03A4C5A2B" blockId="5.[812,1450,553,611]" pageId="5" pageNumber="216">
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<emphasis id="C4ADEAC80140FF8EFCBBFDB03CB35A09" bold="true" box="[812,958,553,577]" italics="true" pageId="5" pageNumber="216">Etymology. –</emphasis>
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From the Latin word
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<taxonomicName id="31D94D590140FF8EFB27FDB03A095A08" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[1200,1284,553,576]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="216" phylum="Arthropoda" rank="species" species="verruca">
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<emphasis id="C4ADEAC80140FF8EFB27FDB03A095A08" box="[1200,1284,553,576]" italics="true" pageId="5" pageNumber="216">verruca</emphasis>
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</taxonomicName>
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in reference to the presence of a tubercle on the male fore tibia.
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</paragraph>
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</subSubSection>
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<subSubSection id="BEC365510140FF8EFCBBFD083C725AA5" pageId="5" pageNumber="216" type="diagnosis">
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<paragraph id="F66636DA0140FF8EFCBBFD083C725AA5" blockId="5.[812,1450,657,749]" pageId="5" pageNumber="216">
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<emphasis id="C4ADEAC80140FF8EFCBBFD083CBE5AE0" box="[812,947,657,681]" italics="true" pageId="5" pageNumber="216">
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<emphasis id="C4ADEAC80140FF8EFCBBFD083CAC5AE1" bold="true" box="[812,929,657,681]" italics="true" pageId="5" pageNumber="216">Diagnosis.</emphasis>
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–
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</emphasis>
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Medium-sized brownish species with fore tibia deformed and bearing an anterodorsal tubercle at basal quarter.
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</paragraph>
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</subSubSection>
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<subSubSection id="BEC365510140FF8EFCBBFC823C725FE3" pageId="5" pageNumber="216" type="description">
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<paragraph id="F66636DA0140FF8EFCBBFC823C855E3B" blockId="5.[812,1451,795,1651]" pageId="5" pageNumber="216">
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||
<emphasis id="C4ADEAC80140FF8EFCBBFC823B505B7B" box="[812,1117,795,819]" italics="true" pageId="5" pageNumber="216">
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<emphasis id="C4ADEAC80140FF8EFCBBFC823B055B7B" bold="true" box="[812,1032,795,819]" italics="true" pageId="5" pageNumber="216">Description. – Male</emphasis>
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– Head
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</emphasis>
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. Occiput dark brown blackish, antenna blackish, second and third flagellomeres aristiform. Palpus brown, labrum length almost twice head height.
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<emphasis id="C4ADEAC80140FF8EFCBBFC1D3C745BD3" box="[812,889,900,923]" italics="true" pageId="5" pageNumber="216">Thorax</emphasis>
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dark brown. Proepisternum with 1 fine, short bristle, proepisternum with 1 rather long bristle. Dorsocentrals uniserial, strong, long.
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<emphasis id="C4ADEAC80140FF8EFBA3FC503B6A5BA8" box="[1076,1127,969,992]" italics="true" pageId="5" pageNumber="216">Legs</emphasis>
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(
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<figureCitation id="6EE22A5F0140FF8EFBEDFC503BDF5BA8" box="[1146,1234,969,992]" captionStart="Fig" captionStartId="1.[113,149,2063,2084]" captionTargetBox="[117,1448,682,1971]" captionTargetId="figure-9@1.[102,1460,631,1988]" captionTargetPageId="1" captionText="Fig. 1. Male fore leg in frontal view: A, tibia and first tarsomere of E. (C.) neesoonensis; B, tibia and tarsus of E. (C.) producta; C, tibia and first tarsomere of E. (C.) tortuosa; D, tibia and first tarsomere of E. (C.) verruca." figureDoi="http://doi.org/10.5281/zenodo.4619640" httpUri="https://zenodo.org/record/4619640/files/figure.png" pageId="5" pageNumber="216">Fig. 1D</figureCitation>
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) brown. Fore tibia deformed at base with a characteristic anterodorsal tubercle at basal quarter, 1 rather strong dorsal bristle at middle, longer than tibia depth; first fore tarsomere with 1 strong, long anterolateral bristle at base, 1 strong, long dorsal bristle apically. Mid tibia with 3 strong dorsal bristles (at basal quarter, near middle, at tip), the last two long, 1 strong ventral bristle at basal third; first mid tarsomere with 2 strong, long dorsal and ventral bristles basally, 1 strong, very long posterodorsal bristle apically, following three mid tarsomeres with 1 strong, long dorsal bristle apically. Hind tibia with row of strong, long dorsal bristles; first hind tarsomere with pair of strong, long dorsal bristles apically, following four tarsomeres with pair of distinct dorsal bristles apically.
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<emphasis id="C4ADEAC80140FF8EFAE6FA153AA45DEB" box="[1393,1449,1420,1443]" italics="true" pageId="5" pageNumber="216">Wing</emphasis>
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(length =
|
||
<quantity id="31219B3F0140FF8EFC06FA373CEF5D8D" box="[913,994,1454,1477]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="216" unit="mm" value="2.6">2.6 mm</quantity>
|
||
) feebly tinged with brown, A1 abbreviated. Halter brown.
|
||
<emphasis id="C4ADEAC80140FF8EFC54FA483B2A5DA0" box="[963,1063,1489,1512]" italics="true" pageId="5" pageNumber="216">Abdomen</emphasis>
|
||
brown. Sternites with pair of distinct bristles posteriorly.
|
||
<emphasis id="C4ADEAC80140FF8EFB89FA6D3BBD5E43" box="[1054,1200,1524,1547]" italics="true" pageId="5" pageNumber="216">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="6EE22A5F0140FF8EFB52FA6D3A1D5E43" box="[1221,1296,1524,1547]" captionStart="Fig" captionStartId="8.[137,173,2091,2112]" captionText="Fig. 8. Male hypopygium of E. (C.) verruca, lateral view. Scale = 4.2 mm." pageId="5" pageNumber="216">Fig. 8</figureCitation>
|
||
). Epandrium distinctly triangular with 3 strong, long bristles at tip. Hypandrium long, pointed at tip. Phallus somewhat narrowed apically.
|
||
</paragraph>
|
||
<paragraph id="F66636DA0140FF8EFCBBF9383C725FE3" blockId="5.[812,1451,1697,1963]" pageId="5" pageNumber="216">
|
||
<emphasis id="C4ADEAC80140FF8EFCBBF9383C8D5EF1" bold="true" box="[812,896,1697,1721]" italics="true" pageId="5" pageNumber="216">Female</emphasis>
|
||
similar to male except for the following characters: wing more distinctly brown. Fore femur distinctly paler than coxa and tibia; fore tibia with dorsal pennation as long as tibia depth; first fore tarsomere with short dorsal pennation except apically; mid femur, hind femur and tibia with long dorsal and ventral pennation; mid tibia with short dorsal pennation; first hind tarsomere with short dorsal pennation basally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BEC365510140FF8EFCBBF8403BEE507D" pageId="5" pageNumber="216" type="discussion">
|
||
<paragraph id="F66636DA0140FF8EFCBBF8403BEE507D" blockId="5.[812,1450,2009,2101]" pageId="5" pageNumber="216">
|
||
<emphasis id="C4ADEAC80140FF8EFCBBF8403CA65FB9" bold="true" box="[812,939,2009,2033]" italics="true" pageId="5" pageNumber="216">Remarks. –</emphasis>
|
||
This species belongs to the
|
||
<taxonomicName id="31D94D590140FF8EFB75F8403A475FB8" authorityName="Melander" authorityYear="1946" box="[1250,1354,2009,2032]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="216" phylum="Arthropoda" rank="species" species="hyalea">
|
||
<emphasis id="C4ADEAC80140FF8EFB75F8403A475FB8" box="[1250,1354,2009,2032]" italics="true" pageId="5" pageNumber="216">E. hyalea</emphasis>
|
||
</taxonomicName>
|
||
clade. It can be distinguished from all other species by the presence of a characteristic tubercle on fore tibia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |