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<document id="CFC2F0749C8E926A68E6C52A76FB3307" ID-DOI="10.2478/if-2017-0019" ID-ISSN="2533-4069" ID-Zenodo-Dep="5385228" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630301147497" checkinUser="diego" docAuthor="Morales, Jorge &amp; Pickford, Martin" docDate="2017" docId="03FB2F7EF40FFFDAFF7DFCA45F03FCCB" docLanguage="en" docName="19_Morales_new-DOI-1.pdf" docOrigin="Fossil Imprint 73 (3 - 4)" docSource="http://dx.doi.org/10.2478/if-2017-0019" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Dissopsalis pyroclasticus SAVAGE 1965" docType="treatment" docVersion="6" lastPageNumber="335" masterDocId="FFC25706F40CFFD9FF91FF905A43FFD7" masterDocTitle="New Hyaenodonts (Ferae, Mammalia) From The Early Miocene Of Napak (Uganda), Koru (Kenya) And Grillental (Namibia)" masterLastPageNumber="359" masterPageNumber="332" pageNumber="335" updateTime="1699220586648" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="14A2B658B2C74F2A41176BF8D6835918">New Hyaenodonts (Ferae, Mammalia) From The Early Miocene Of Napak (Uganda), Koru (Kenya) And Grillental (Namibia)</mods:title>
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<mods:namePart id="E58605088B7EEF3E5D699419CCD80A2C">Morales, Jorge</mods:namePart>
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<taxonomicName id="4C52E5EBF40FFFDAFF7DFCA458D7FC9B" authority="SAVAGE, 1965" authorityName="SAVAGE" authorityYear="1965" box="[236,660,820,844]" class="Mammalia" family="Hyaenodontidae" genus="Dissopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="3" pageNumber="335" phylum="Chordata" rank="species" species="pyroclasticus">
<emphasis id="B926427AF40FFFDAFF7DFCA458D7FC9B" bold="true" box="[236,660,820,844]" pageId="3" pageNumber="335">
<emphasis id="B926427AF40FFFDAFF7DFCA45BBEFC9B" bold="true" box="[236,509,820,844]" italics="true" pageId="3" pageNumber="335">Dissopsalis pyroclasticus</emphasis>
<bibRefCitation id="EFC3E399F40FFFDAFD95FCA55811FC9D" author="Savage, R. J. G." box="[516,594,821,844]" pageId="3" pageNumber="335" pagination="239 - 316" refId="ref17738" refString="Savage, R. J. G. (1965): The Miocene Carnivora of East Africa. Fossil Mammals of Africa: 19. - Bulletin of the British Museum of Natural History (Geology), 10: 239 - 316." type="journal article" year="1965">SAVAGE</bibRefCitation>
, 1965
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<paragraph id="8BED9E68F40FFFDAFF37FCF4588CFC4C" blockId="3.[129,748,868,923]" pageId="3" pageNumber="335">H o l o t y p e. Right mandible containing p4, m1 m3 and alveoli of canine and p1 p3 (NHMUK M 19082).</paragraph>
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<subSubSection id="C348CDE3F40FFFDAFCE0FF085F03FCCB" pageId="3" pageNumber="335" type="description">
<paragraph id="8BED9E68F40FFFDAFCE0FF085F38FF38" blockId="3.[843,1482,152,239]" pageId="3" pageNumber="335">
T y p e l o c a l i t y. Maboko,
<collectingCountry id="F345DEF8F40FFFDAFB25FF085EB9FF78" box="[1204,1274,152,175]" name="Kenya" pageId="3" pageNumber="335">Kenya</collectingCountry>
(erroneously given as Kaboor (error for Kabua = alternative name for Maboko)), Northern Frontier District,
<collectingCountry id="F345DEF8F40FFFDAFBFDFF485EF1FF38" box="[1132,1202,216,239]" name="Kenya" pageId="3" pageNumber="335">Kenya</collectingCountry>
by
<bibRefCitation id="EFC3E399F40FFFDAFB48FF485F34FF38" author="Savage, R. J. G." box="[1241,1399,216,239]" pageId="3" pageNumber="335" pagination="239 - 316" refId="ref17738" refString="Savage, R. J. G. (1965): The Miocene Carnivora of East Africa. Fossil Mammals of Africa: 19. - Bulletin of the British Museum of Natural History (Geology), 10: 239 - 316." type="journal article" year="1965">Savage (1965)</bibRefCitation>
.
</paragraph>
<paragraph id="8BED9E68F40FFFDAFCE0FE975F6FFEC9" blockId="3.[881,1324,263,286]" box="[881,1324,263,286]" pageId="3" pageNumber="335">A g e. Basal Middle Miocene, ca. 15 Ma.</paragraph>
<paragraph id="8BED9E68F40FFFDAFCE0FEA65EABFE9A" blockId="3.[881,1256,310,333]" box="[881,1256,310,333]" pageId="3" pageNumber="335">
D i a g n o s i s. See
<bibRefCitation id="EFC3E399F40FFFDAFBD7FEA65EA7FE9A" author="Savage, R. J. G." box="[1094,1252,310,333]" pageId="3" pageNumber="335" pagination="239 - 316" refId="ref17738" refString="Savage, R. J. G. (1965): The Miocene Carnivora of East Africa. Fossil Mammals of Africa: 19. - Bulletin of the British Museum of Natural History (Geology), 10: 239 - 316." type="journal article" year="1965">Savage (1965)</bibRefCitation>
.
</paragraph>
<paragraph id="8BED9E68F40FFFDAFCE0FEF55F03FCCB" blockId="3.[843,1482,357,796]" pageId="3" pageNumber="335">
D i s c u s s i o n.
<bibRefCitation id="EFC3E399F40FFFDAFBB9FEF55EECFEAB" author="Barry, J. C." box="[1064,1199,357,380]" pageId="3" pageNumber="335" pagination="25 - 45" refId="ref15277" refString="Barry, J. C. (1988): Dissopsalis, a middle and late Miocene proviverrine creodont (Mammalia) from Pakistan and Kenya. - Journal of Vertebrate Paleontology, 8: 25 - 45. https: // doi. org / 10.1080 / 02724634.1988.10011682" type="journal article" year="1988">Barry (1988)</bibRefCitation>
cited several quite obvious differences between the lower dentitions of
<taxonomicName id="4C52E5EBF40FFFDAFAC0FE155E74FE6B" authority="PILGRIM, 1910" authorityName="PILGRIM" authorityYear="1910" class="Mammalia" family="Hyaenodontidae" genus="Dissopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="3" pageNumber="335" phylum="Chordata" rank="species" species="carnifex">
<emphasis id="B926427AF40FFFDAFAC0FE1559E2FE6B" italics="true" pageId="3" pageNumber="335">Dissopsalis carnifex</emphasis>
<bibRefCitation id="EFC3E399F40FFFDAFC37FE355E74FE6B" author="Pilgrim, G. E." box="[934,1079,421,444]" pageId="3" pageNumber="335" pagination="63 - 71" refId="ref17409" refString="Pilgrim, G. E. (1910): Notices of new Mammalian Genera and species from the Tertiaries of India. - Records of the Geological Survey of India, 40: 63 - 71." type="journal article" year="1910">PILGRIM, 1910</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C52E5EBF40FFFDAFBF9FE355F33FE6B" authorityName="SAVAGE" authorityYear="1965" box="[1128,1392,421,444]" class="Mammalia" family="Hyaenodontidae" genus="Dissopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="3" pageNumber="335" phylum="Chordata" rank="species" species="pyroclasticus">
<emphasis id="B926427AF40FFFDAFBF9FE355F33FE6B" box="[1128,1392,421,444]" italics="true" pageId="3" pageNumber="335">Dissopsalis pyroclasticus</emphasis>
</taxonomicName>
. Among these were the more pronounced metaconid in the molars of
<taxonomicName id="4C52E5EBF40FFFDAFCDAFE7559B5FE2B" authorityName="SAVAGE" authorityYear="1965" box="[843,1014,485,508]" class="Mammalia" family="Hyaenodontidae" genus="Dissopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="3" pageNumber="335" phylum="Chordata" rank="species" species="pyroclasticus">
<emphasis id="B926427AF40FFFDAFCDAFE7559B5FE2B" box="[843,1014,485,508]" italics="true" pageId="3" pageNumber="335">D. pyroclasticus</emphasis>
</taxonomicName>
, the differences in length of the p4 and m3, and the taller crown and the diminutive talonid of the m3 of
<taxonomicName id="4C52E5EBF40FFFDAFA21FD955996FDEB" authorityName="SAVAGE" authorityYear="1965" class="Mammalia" family="Hyaenodontidae" genus="Dissopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="3" pageNumber="335" phylum="Chordata" rank="species" species="pyroclasticus">
<emphasis id="B926427AF40FFFDAFA21FD955996FDEB" italics="true" pageId="3" pageNumber="335">D. pyroclasticus</emphasis>
</taxonomicName>
. To these differences could be added the greater size of the m2 with respect to the m
<quantity id="4CAA338DF40FFFDAFB70FDD55F48FD8B" box="[1249,1291,581,604]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="335" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C52E5EBF40FFFDAFA84FDD55F80FD8B" authorityName="SAVAGE" authorityYear="1965" box="[1301,1475,581,604]" class="Mammalia" family="Hyaenodontidae" genus="Dissopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="3" pageNumber="335" phylum="Chordata" rank="species" species="pyroclasticus">
<emphasis id="B926427AF40FFFDAFA84FDD55F80FD8B" box="[1301,1475,581,604]" italics="true" pageId="3" pageNumber="335">D. pyroclasticus</emphasis>
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. These differences, although important, do not exclude the possibility of the two species being classified in the same genus.
<bibRefCitation id="EFC3E399F40FFFDAFC09FD355E2BFD6B" author="Borths, M. R. &amp; Holdroyd, P. A. &amp; Seiffert, E. R." box="[920,1128,677,700]" pageId="3" pageNumber="335" refId="ref15479" refString="Borths, M. R. Holdroyd, P. A., Seiffert, E. R. (2016): Hyainailourine and Teratodontine cranial material from the late Eocene of Egypt and the application of parsimony and Bayesian methods to the phylogeny and biogeography of Hyaenodonta (Placentalia, Mammalia). - PeerJ, 4: e 2639 (87 pp.). https: // doi. org / 10.7717 / peerj. 2639" type="book" year="2016">Borths et al. (2016</bibRefCitation>
: supplementary tab. 2) included, but did not describe, additional upper and lower dentitions in this species, the study of which would, without doubt, help to clarify the relationship between the two species.
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