139 lines
17 KiB
XML
139 lines
17 KiB
XML
<document id="7FCA2D8782B2CB93BF560AB210C57E2A" ID-DOI="10.2478/if-2017-0019" ID-ISSN="2533-4069" ID-Zenodo-Dep="5385228" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630301147497" checkinUser="diego" docAuthor="Morales, Jorge & Pickford, Martin" docDate="2017" docId="03FB2F7EF408FFDCFC15F80C582CF846" docLanguage="en" docName="19_Morales_new-DOI-1.pdf" docOrigin="Fossil Imprint 73 (3 - 4)" docSource="http://dx.doi.org/10.2478/if-2017-0019" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Buhakia hyaenoides" docType="treatment" docVersion="5" lastPageNumber="337" masterDocId="FFC25706F40CFFD9FF91FF905A43FFD7" masterDocTitle="New Hyaenodonts (Ferae, Mammalia) From The Early Miocene Of Napak (Uganda), Koru (Kenya) And Grillental (Namibia)" masterLastPageNumber="359" masterPageNumber="332" pageNumber="336" updateTime="1699220586648" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
|
||
<mods:mods id="9454D1B516BE70B117C57077B4505889" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="28BBE990406649FB7F6A3333B2B6AB34">
|
||
<mods:title id="9102558BAC39E114D30DC51D2F47425B">New Hyaenodonts (Ferae, Mammalia) From The Early Miocene Of Napak (Uganda), Koru (Kenya) And Grillental (Namibia)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="1BAD7D27017603634062AD275CB6239C" type="personal">
|
||
<mods:role id="F40AE041E966E52897FEDD5E136817EF">
|
||
<mods:roleTerm id="E8B45F9A38360347653E788A2E881D4F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="201A7CD778930C527782D8867BBACEB3">Morales, Jorge</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="81F2E6673AA4E786D94EBEC6DCA54339" type="personal">
|
||
<mods:role id="C1E000004E2146D36B3078E334BB5FAF">
|
||
<mods:roleTerm id="84C0BCA67E8D0F1B79836B4CE0DE17D1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0F550B717FED4838AD66E459C1DDA6E6">Pickford, Martin</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="4A594C7B79703484AA8B526955EB5994">text</mods:typeOfResource>
|
||
<mods:relatedItem id="AD17BFB402E8A5EC84AEF7C1F8731D1C" type="host">
|
||
<mods:titleInfo id="CD26A3867910A4C5F8CCFF6322FFAA53">
|
||
<mods:title id="E6DCC57C43E902F3FF8466047516CCFE">Fossil Imprint</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="AF999B5F09C28413DF2682DB5C3CFA75">
|
||
<mods:date id="27FB977ED0ABD9C25D621BFE707C24AD">2017</mods:date>
|
||
<mods:detail id="F19E4F0CBF62D1231195DA6FFD287910" type="pubDate">
|
||
<mods:number id="E10035CF8604B545222BA227B8C084B4">2017-12-31</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="219C26877609181AE89DCAA7621309F2" type="volume">
|
||
<mods:number id="4760ABC8F854E9FE10E674FBD427AD5A">73</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="1610013B5F6E0624CA95441D256F8285" type="issue">
|
||
<mods:number id="9C3711680B1D204F8ACA8D2B92516B58">3 - 4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="7CB6489272004EB0AFBBACEE461C2E9A" unit="page">
|
||
<mods:start id="E71DF140ECDA40E40FA1908C90DD24FF">332</mods:start>
|
||
<mods:end id="CC4CF37B0EB2934B1E21A772BBAF5E17">359</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="52153BDEEC019448DC8464D1788928AF">
|
||
<mods:url id="65E2C3FCF27A3F243A702F32DD9FFC8F">http://dx.doi.org/10.2478/if-2017-0019</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="938531C2DA412695E5DE1D03F7C0DDC2">journal article</mods:classification>
|
||
<mods:identifier id="F2C547C8C8888731CA7D2524427473B2" type="DOI">10.2478/if-2017-0019</mods:identifier>
|
||
<mods:identifier id="134AECD0F2F1C04AA9D27AC4BF9C38D4" type="ISSN">2533-4069</mods:identifier>
|
||
<mods:identifier id="4BF864D1F22CFFF946B0A42EFBE559AE" type="Zenodo-Dep">5385228</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03FB2F7EF408FFDCFC15F80C582CF846" LSID="urn:lsid:plazi:treatment:03FB2F7EF408FFDCFC15F80C582CF846" httpUri="http://treatment.plazi.org/id/03FB2F7EF408FFDCFC15F80C582CF846" lastPageId="5" lastPageNumber="337" pageId="4" pageNumber="336">
|
||
<subSubSection id="C348CDE3F408FFDDFC15F80C5F20F862" box="[900,1379,1948,1973]" pageId="4" pageNumber="336" type="nomenclature">
|
||
<paragraph id="8BED9E68F408FFDDFC15F80C5F20F862" blockId="4.[900,1379,1948,1973]" box="[900,1379,1948,1973]" pageId="4" pageNumber="336">
|
||
<heading id="D0A52904F408FFDDFC15F80C5F20F862" box="[900,1379,1948,1973]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="336" reason="2">
|
||
<taxonomicName id="4C52E5EBF408FFDDFC15F80C5F20F862" authority="(MORALES et al., 2003)" baseAuthorityName="MORALES" baseAuthorityYear="2003" box="[900,1379,1948,1973]" class="Mammalia" family="Teratodontidae" genus="Buhakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="336" phylum="Chordata" rank="species" species="hyaenoides">
|
||
<emphasis id="B926427AF408FFDDFC15F80C5F20F862" bold="true" box="[900,1379,1948,1973]" pageId="4" pageNumber="336">
|
||
<emphasis id="B926427AF408FFDDFC15F80C5E26F863" bold="true" box="[900,1125,1948,1972]" italics="true" pageId="4" pageNumber="336">Buhakia hyaenoides</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC3E399F408FFDDFBE2F80E5E99F864" author="Morales, J. & Pickford, M. & Fraile, S. & Salesa, M. J. & Soria, D." box="[1139,1242,1950,1973]" pageId="4" pageNumber="336" pagination="177 - 194" refId="ref16599" refString="Morales, J., Pickford, M., Fraile, S., Salesa, M. J., Soria, D. (2003): Creodonta and Carnivora from Arrisdrift, early Middle Miocene of Southern Namibia. - Memoir of the Geological Survey of Namibia, 19: 177 - 194." type="journal article" year="2003">MORALES</bibRefCitation>
|
||
et al., 2003)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348CDE3F408FFDCFBADF82E582CF846" lastPageId="5" lastPageNumber="337" pageId="4" pageNumber="336" type="description">
|
||
<paragraph id="8BED9E68F408FFDDFBADF82E5EE8F802" blockId="4.[1084,1195,1982,2005]" box="[1084,1195,1982,2005]" pageId="4" pageNumber="336">
|
||
<figureCitation id="136982EDF408FFDDFBADF82E5EE8F802" box="[1084,1195,1982,2005]" captionStart="Text-fig" captionStartId="5.[129,210,889,911]" captionTargetBox="[206,748,168,834]" captionTargetId="figure-855@5.[132,767,151,885]" captionTargetPageId="5" captionText="Text-fig. 2. Buhakia hyaenoides (MORALES et al., 2003) from Arrisdrift, Sperrgebiet (Namibia). GSN AD 241’99 (holotype), left m2. a: lingual view, b: buccal view, c: occlusal view (stereo pairs)." figureDoi="http://doi.org/10.5281/zenodo.5385232" httpUri="https://zenodo.org/record/5385232/files/figure.png" pageId="4" pageNumber="336">Text-fig. 2</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="8BED9E68F408FFDDFCA5F87D5F31F7D3" blockId="4.[820,1394,2029,2052]" box="[820,1394,2029,2052]" pageId="4" pageNumber="336">
|
||
2003
|
||
<taxonomicName id="4C52E5EBF408FFDDFC11F87D5E38F7D3" box="[896,1147,2029,2052]" class="Mammalia" family="Viverridae" genus="Africanictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="336" phylum="Chordata" rank="species" species="hyaenoides">
|
||
<emphasis id="B926427AF408FFDDFC11F87D5E38F7D3" box="[896,1147,2029,2052]" italics="true" pageId="4" pageNumber="336">Africanictis hyaenoides</emphasis>
|
||
</taxonomicName>
|
||
; Morales et al., p. 183.
|
||
</paragraph>
|
||
<paragraph id="8BED9E68F408FFDDFCCBF78C5FE7F7E4" blockId="4.[858,1444,2076,2099]" box="[858,1444,2076,2099]" pageId="4" pageNumber="336">
|
||
H o l o t y p e. Left m2 (GSN AD 241’99) (
|
||
<figureCitation id="136982EDF408FFDDFAB5F78C5FD4F7E4" box="[1316,1431,2076,2099]" captionStart="Text-fig" captionStartId="5.[129,210,889,911]" captionTargetBox="[206,748,168,834]" captionTargetId="figure-855@5.[132,767,151,885]" captionTargetPageId="5" captionText="Text-fig. 2. Buhakia hyaenoides (MORALES et al., 2003) from Arrisdrift, Sperrgebiet (Namibia). GSN AD 241’99 (holotype), left m2. a: lingual view, b: buccal view, c: occlusal view (stereo pairs)." figureDoi="http://doi.org/10.5281/zenodo.5385232" httpUri="https://zenodo.org/record/5385232/files/figure.png" pageId="4" pageNumber="336">Text-fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF2DCEE0F409FFDCFF10FCE95A81FC30" ID-DOI="http://doi.org/10.5281/zenodo.5385232" ID-Zenodo-Dep="5385232" httpUri="https://zenodo.org/record/5385232/files/figure.png" pageId="5" pageNumber="337" startId="5.[129,210,889,911]" targetBox="[206,748,168,834]" targetPageId="5">
|
||
<paragraph id="8BED9E68F409FFDCFF10FCE95A81FC30" blockId="5.[129,764,889,999]" pageId="5" pageNumber="337">
|
||
<emphasis id="B926427AF409FFDCFF10FCE95AB1FC58" bold="true" box="[129,242,889,911]" pageId="5" pageNumber="337">Text-fig. 2.</emphasis>
|
||
<taxonomicName id="4C52E5EBF409FFDCFF6DFCEA5883FC58" authority="(MORALES et al., 2003)" baseAuthorityName="MORALES" baseAuthorityYear="2003" box="[252,704,889,911]" class="Mammalia" family="Teratodontidae" genus="Buhakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="hyaenoides">
|
||
<emphasis id="B926427AF409FFDCFF6DFCEA587BFC5A" bold="true" box="[252,568,889,911]" pageId="5" pageNumber="337">
|
||
<emphasis id="B926427AF409FFDCFF6DFCEA5B89FC58" bold="true" box="[252,458,890,911]" italics="true" pageId="5" pageNumber="337">Buhakia hyaenoides</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC3E399F409FFDCFE4AFCE9587BFC5A" author="Morales, J. & Pickford, M. & Fraile, S. & Salesa, M. J. & Soria, D." box="[475,568,889,911]" pageId="5" pageNumber="337" pagination="177 - 194" refId="ref16599" refString="Morales, J., Pickford, M., Fraile, S., Salesa, M. J., Soria, D. (2003): Creodonta and Carnivora from Arrisdrift, early Middle Miocene of Southern Namibia. - Memoir of the Geological Survey of Namibia, 19: 177 - 194." type="journal article" year="2003">MORALES</bibRefCitation>
|
||
</emphasis>
|
||
et al., 2003)
|
||
</taxonomicName>
|
||
from Arrisdrift, Sperrgebiet (Namibia). GSN AD 241’99 (holotype), left m2. a: lingual view, b: buccal view, c: occlusal view (stereo pairs).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BED9E68F409FFDCFF37FBBE5814FB92" blockId="5.[166,599,1070,1093]" box="[166,599,1070,1093]" pageId="5" pageNumber="337">
|
||
T y p e l o c a l i t y. Arrisdrift,
|
||
<collectingCountry id="F345DEF8F409FFDCFE62FBBE5811FB92" box="[499,594,1070,1093]" name="Namibia" pageId="5" pageNumber="337">Namibia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BED9E68F409FFDCFF37FBCE5BA8FBA2" blockId="5.[166,491,1118,1141]" box="[166,491,1118,1141]" pageId="5" pageNumber="337">A g e. Basal Middle Miocene.</paragraph>
|
||
<paragraph id="8BED9E68F409FFDCFF37FB1E582CF846" blockId="5.[129,768,1166,1937]" pageId="5" pageNumber="337">
|
||
D i s c u s s i o n.
|
||
<bibRefCitation id="EFC3E399F409FFDCFECBFB1E586CFB72" author="Morales, J. & Pickford, M. & Fraile, S. & Salesa, M. J. & Soria, D." box="[346,559,1166,1189]" pageId="5" pageNumber="337" pagination="177 - 194" refId="ref16599" refString="Morales, J., Pickford, M., Fraile, S., Salesa, M. J., Soria, D. (2003): Creodonta and Carnivora from Arrisdrift, early Middle Miocene of Southern Namibia. - Memoir of the Geological Survey of Namibia, 19: 177 - 194." type="journal article" year="2003">Morales et al.(2003)</bibRefCitation>
|
||
defined
|
||
<taxonomicName id="4C52E5EBF409FFDCFD12FB1E5ABAFB12" class="Mammalia" family="Viverridae" genus="Africanictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="hyaenoides">
|
||
<emphasis id="B926427AF409FFDCFD12FB1E5ABAFB12" italics="true" pageId="5" pageNumber="337">Africanictis hyaenoides</emphasis>
|
||
</taxonomicName>
|
||
at the locality of Arrisdrift (
|
||
<collectingCountry id="F345DEF8F409FFDCFDD4FB3E58E7FB12" box="[581,676,1198,1221]" name="Namibia" pageId="5" pageNumber="337">Namibia</collectingCountry>
|
||
) on the basis of a suspected m1 with specialised features (loss of the metaconid, narrow, cutting trigonid and reduced talonid, although it possesses a hypoconid and entoconid) thereby differing from the
|
||
<typeStatus id="54E920CAF409FFDCFEC5FAA05BC1FA90" box="[340,386,1328,1351]" pageId="5" pageNumber="337">type</typeStatus>
|
||
species of the genus,
|
||
<taxonomicName id="4C52E5EBF409FFDCFD12FAA05BA7FABF" authority="(Morales et al. 1998 b)" baseAuthorityName="Morales" baseAuthorityYear="1998" class="Mammalia" family="Viverridae" genus="Africanictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="meini">
|
||
<emphasis id="B926427AF409FFDCFD12FAA05AFFFABF" italics="true" pageId="5" pageNumber="337">Africanictis meini</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC3E399F409FFDCFF47FAC15B9FFABF" author="Morales, J. & Pickford, M. & Soria, D. & Fraile, S." box="[214,476,1361,1384]" pageId="5" pageNumber="337" pagination="27 - 40" refId="ref16850" refString="Morales, J., Pickford, M., Soria, D., Fraile, S., (1998 b): New carnivores from the basal Middle Miocene of Arrisdrift, Namibia. - Eclogae Geologicae Helvetiae, 91: 27 - 40." type="journal article" year="1998">Morales et al. 1998b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Among the carnivores described by
|
||
<bibRefCitation id="EFC3E399F409FFDCFE9FFAE15B9BFA5F" author="Morlo, M. & Miller, E. R. & El-Barkooky, A. N." box="[270,472,1393,1416]" pageId="5" pageNumber="337" pagination="145 - 159" refId="ref16949" refString="Morlo, M., Miller, E. R., El-Barkooky, A. N. (2007): Creodonta and Carnivora from Wadi Moghra, Egypt. - Journal of Vertebrate Paleontology, 27: 145 - 159. https: // doi. org / 10.1671 / 0272 - 4634 (2007) 27 [145: CACF" type="journal article" year="2007">Morlo et al. (2007)</bibRefCitation>
|
||
from Wadi Moghra,
|
||
<collectingCountry id="F345DEF8F409FFDCFD28FAE158B8FA5F" box="[697,763,1393,1416]" name="Egypt" pageId="5" pageNumber="337">Egypt</collectingCountry>
|
||
, there was a new creodont,
|
||
<taxonomicName id="4C52E5EBF409FFDCFE37FA0258D3FA7E" box="[422,656,1426,1449]" class="Mammalia" family="Teratodontidae" genus="Buhakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="moghraensis">
|
||
<emphasis id="B926427AF409FFDCFE37FA0258D3FA7E" box="[422,656,1426,1449]" italics="true" pageId="5" pageNumber="337">Buhakia moghraensis</emphasis>
|
||
</taxonomicName>
|
||
, based on a subadult mandible of which the dentition was interpreted to be dp4, m1 and m2. In particular, the last molar in this specimen is morphologically quite close to the Arrisdrift species, which is only slightly larger (
|
||
<figureCitation id="136982EDF409FFDCFDD0F98458F9F9FC" box="[577,698,1556,1579]" captionStart="Text-fig" captionStartId="6.[125,206,988,1010]" captionTargetBox="[196,1388,197,904]" captionTargetId="figure-41@6.[124,1441,151,955]" captionTargetPageId="6" captionText="Text-fig. 3. Bivariate plots of the lower teeth (m3 – m2) of small to medium sized Miocene hyaenodonts from African localities. Data source: Pilgrim (1912, 1932), Colbert (1935), Savage (1965), Barry (1988), Morales et al. (1998a, 2003, 2007, 2008, 2010), Holroyd (1999), Morlo et al. (2007), Rasmussen et al. (2009), Borths et al. (2016), Borths and Seiffert (2017)." figureDoi="http://doi.org/10.5281/zenodo.5385234" httpUri="https://zenodo.org/record/5385234/files/figure.png" pageId="5" pageNumber="337">Text-fig. 3</figureCitation>
|
||
). The Arrisdrift molar is unworn, and has open roots, indicting its subadult status, similar in age to the individual of
|
||
<taxonomicName id="4C52E5EBF409FFDCFD79F9C55B4AF95A" class="Mammalia" family="Teratodontidae" genus="Buhakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="moghraensis">
|
||
<emphasis id="B926427AF409FFDCFD79F9C55B4AF95A" italics="true" pageId="5" pageNumber="337">B. moghraensis</emphasis>
|
||
</taxonomicName>
|
||
. The Arrisdrift molar has a basal cingulum on the buccal side of the paraconid which sweeps upwards along the anterior margin and swells such that occlusion with the talonid of the preceding molar (m1) is via this cingulum and the anteriormost extremity of the paraconid, a conformation visible in
|
||
<taxonomicName id="4C52E5EBF409FFDCFF76F8885B8DF8F8" box="[231,462,1816,1839]" class="Mammalia" family="Teratodontidae" genus="Buhakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="moghraensis">
|
||
<emphasis id="B926427AF409FFDCFF76F8885B8DF8F8" box="[231,462,1816,1839]" italics="true" pageId="5" pageNumber="337">Buhakia moghraensis</emphasis>
|
||
</taxonomicName>
|
||
and frequent in hyaenodonts in which the molars are imbricated. Other characters common to the two species are the loss of the metaconid and the presence of cutting talonid and hypoconid.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |