168 lines
20 KiB
XML
168 lines
20 KiB
XML
<document id="2DCD0B1E43300DC4C888A73F0C2F85C7" ID-CLB-Dataset="6414" ID-DOI="10.5252/g2013n1a2" ID-GBIF-Dataset="c36e980f-7d7d-4b7d-a253-d3d4dc73de11" ID-ISSN="1638-9395" ID-Zenodo-Dep="4537923" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613149088530" checkinUser="felipe" docAuthor="Löser, Hannes" docDate="2013" docId="03F487D7FFEFFFB5FD1F860EFA2F759D" docLanguage="en" docName="Geodiversitas.35.1.21-29.pdf" docOrigin="Geodiversitas 35 (1)" docSource="http://dx.doi.org/10.5252/g2013n1a2" docStyle="DocumentStyle:F8AA0CB5AB58FB1C1EB3E4696C1B2D36.3:Geodiversitas.2013-2014.journal_article" docStyleId="F8AA0CB5AB58FB1C1EB3E4696C1B2D36" docStyleName="Geodiversitas.2013-2014.journal_article" docStyleVersion="3" docTitle="Asteroseriidae Alloiteau 1952" docType="treatment" docVersion="7" lastPageNumber="23" masterDocId="FFCDFFAFFFEEFFB7FFC88044FE607730" masterDocTitle="Revision of the family Asteroseriidae (Anthozoa, Scleractinia, Cretaceous)" masterLastPageNumber="29" masterPageNumber="21" pageNumber="22" updateTime="1698929317586" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="3793045237DD1AD574110B3F7CD3F399">Revision of the family Asteroseriidae (Anthozoa, Scleractinia, Cretaceous)</mods:title>
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<mods:namePart id="49CE710152986DC16F80F6F491A52C56">Löser, Hannes</mods:namePart>
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<mods:title id="2ED4D961584B6808F5594BC09DAC133B">Geodiversitas</mods:title>
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<mods:date id="F0A950FC7400087667B720F7524D6CCD">2013</mods:date>
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<treatment id="03F487D7FFEFFFB5FD1F860EFA2F759D" ID-DOI="http://doi.org/10.5281/zenodo.4815115" ID-GBIF-Taxon="182399279" ID-Zenodo-Dep="4815115" LSID="urn:lsid:plazi:treatment:03F487D7FFEFFFB5FD1F860EFA2F759D" httpUri="http://treatment.plazi.org/id/03F487D7FFEFFFB5FD1F860EFA2F759D" lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22">
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<subSubSection id="C347654AFFEFFFB6FD1F860EFA527193" pageId="1" pageNumber="22" type="nomenclature">
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<paragraph id="8BE236C1FFEFFFB6FD1F860EFAE77155" blockId="1.[727,1159,1610,1637]" box="[727,1159,1610,1637]" pageId="1" pageNumber="22">
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<heading id="D0AA81ADFFEFFFB6FD1F860EFAE77155" box="[727,1159,1610,1637]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="22" reason="3">
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Family
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<taxonomicName id="4C5D4D42FFEFFFB6FCE1860FFAE77155" ID-CoL="862W4" authority="Alloiteau, 1952" authorityName="Alloiteau" authorityYear="1952" box="[809,1159,1610,1637]" class="Anthozoa" family="Asteroseriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="1" pageNumber="22" phylum="Cnidaria" rank="family">ASTEROSERIIDAE Alloiteau, 1952</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BE236C1FFEFFFB6FD6D86C8FA527193" blockId="1.[677,1074,1675,1699]" box="[677,1074,1675,1699]" pageId="1" pageNumber="22">RANGE. — Hauterivian to Cenomanian.</paragraph>
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</subSubSection>
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<subSubSection id="C347654AFFECFFB5FFB9809CFC7C751C" pageId="2" pageNumber="23" type="diagnosis">
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<paragraph id="8BE236C1FFECFFB5FFB9809CFC7C751C" blockId="2.[112,648,215,556]" pageId="2" pageNumber="23">
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DIAGNOSIS. — Small solitary patellate corallum, generally free-living, but often attached to a small piece of shell debris or a benthic foraminifer. Calicular outline circular.
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<collectingRegion id="4999F823FFECFFB5FF008163FE9D760F" box="[200,253,295,319]" country="Spain" name="Ceuta" pageId="2" pageNumber="23">Septa</collectingRegion>
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compact. Symmetry of septa radial and regularly hexameral with regular cycles. Septal cycles differ in length, but almost not in thickness.
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<collectingRegion id="4999F823FFECFFB5FDFF811FFC0C7643" box="[567,620,347,371]" country="Spain" name="Ceuta" pageId="2" pageNumber="23">Septa</collectingRegion>
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of younger cyles often and regularly connected to septa of preceeding cycles. Septal upper margin with regular large dentations, lateral face with granulae arranged in vertical rows. Pali or paliform lobes absent. Synapticulae present, in a regular pattern. Columella formed by septal fusion in the centre of the calice or parietal. Endotheca absent. Wall compact, formation unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C347654AFFECFFB5FFB98211FF6173B7" pageId="2" pageNumber="23" type="discussion">
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<paragraph id="8BE236C1FFECFFB5FFB98211FEB5755D" blockId="2.[112,647,597,749]" box="[113,213,597,623]" pageId="2" pageNumber="23">REMARKS</paragraph>
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<paragraph id="8BE236C1FFECFFB5FFB98230FC7375DD" blockId="2.[112,647,597,749]" pageId="2" pageNumber="23">Microstructures are hardly preserved in all studied specimens, but the coarsly ornamented upper margins of the septal distal border suggests that the septa are composed of large trabeculae.</paragraph>
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<paragraph id="8BE236C1FFECFFB5FFB98357FF3D741B" blockId="2.[112,648,787,1097]" box="[113,349,787,813]" pageId="2" pageNumber="23">SYSTEMATIC POSITION</paragraph>
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<paragraph id="8BE236C1FFECFFB5FFB98376FF5E7379" blockId="2.[112,648,787,1097]" pageId="2" pageNumber="23">
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The comparable large trabeculae, ornamentation of the lateral face and upper margin of the septa, and the pattern of the synapticulae clearly indicates that the family should be placed into the suborder
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<taxonomicName id="4C5D4D42FFECFFB5FF1183F4FF5D74FA" authorityName="Verrill" authorityYear="1870" box="[217,317,944,970]" class="Anthozoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="subOrder" subOrder="Fungiina">Fungiina</taxonomicName>
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. The existence of fulturae (Gill 1981) could only be proven in one species due to preservation problems and the scarcity of samples. The
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<taxonomicName id="4C5D4D42FFECFFB5FF56844BFF517319" authorityName="Alloiteau" authorityYear="1952" box="[158,305,1039,1065]" class="Anthozoa" family="Asteroseriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="family">Asteroseriidae</taxonomicName>
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are probably the ancestors of the modern
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<taxonomicName id="4C5D4D42FFECFFB5FF03846BFF5A7379" authorityName="Dana" authorityYear="1846" box="[203,314,1071,1097]" class="Anthozoa" family="Fungiidae" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="family">Fungiidae</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BE236C1FFECFFB5FFB98434FF6173B7" blockId="2.[111,647,1136,1573]" box="[113,257,1136,1162]" pageId="2" pageNumber="23">COMPARISON</paragraph>
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</subSubSection>
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<subSubSection id="C347654AFFECFFB5FFB984CAFF0E7152" pageId="2" pageNumber="23" type="reference_group">
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<paragraph id="8BE236C1FFECFFB5FFB984CAFC7F7115" blockId="2.[111,647,1136,1573]" pageId="2" pageNumber="23">
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The
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<taxonomicName id="4C5D4D42FFECFFB5FF5684CAFFAF7399" ID-CoL="84KHP" authority="Vaughan, 1905" authorityName="Vaughan" authorityYear="1905" box="[158,463,1166,1193]" class="Anthozoa" family="Micrabaciidae" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="family">Micrabaciidae Vaughan, 1905</taxonomicName>
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(suborder
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<taxonomicName id="4C5D4D42FFECFFB5FDF384CAFEEC73F8" authorityName="Verrill" authorityYear="1870" class="Anthozoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="subOrder" subOrder="Fungiina">Fungiina</taxonomicName>
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) have perforated septa and a perforated wall. The
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<taxonomicName id="4C5D4D42FFECFFB5FFB88489FFCB73D8" ID-CoL="8586P" authority="Alloiteau, 1952" authorityName="Alloiteau" authorityYear="1952" box="[112,427,1229,1256]" class="Anthozoa" family="Funginellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="family">Funginellidae Alloiteau, 1952</taxonomicName>
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(suborder
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<taxonomicName id="4C5D4D42FFECFFB5FDD78489FCE073D7" authorityName="Verrill" authorityYear="1870" box="[543,640,1229,1255]" class="Anthozoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="subOrder" subOrder="Fungiina">Fungiina</taxonomicName>
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, currently only encompassing the genera
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<taxonomicName id="4C5D4D42FFECFFB5FDD284A9FF7C7217" ID-CoL="043a1d93-5204-48a5-a347-e9e3270dff5a" authority="d'Orbigny, 1849" authorityName="d'Orbigny" authorityYear="1849" class="Anthozoa" family="Cunnolitidae" genus="Funginella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="genus">
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<emphasis id="B929EAD3FFECFFB5FDD284A9FCE67237" box="[538,646,1261,1287]" italics="true" pageId="2" pageNumber="23">Funginella</emphasis>
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d’Orbigny, 1849
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</taxonomicName>
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and
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<taxonomicName id="4C5D4D42FFECFFB5FE828549FC3C7217" authority="Beauvais, 1982" authorityName="Beauvais" authorityYear="1982" box="[330,604,1293,1319]" class="Anthozoa" family="Agariciidae" genus="Elasmogyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="genus">
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<emphasis id="B929EAD3FFECFFB5FE828549FFDA7217" box="[330,442,1293,1319]" italics="true" pageId="2" pageNumber="23">Elasmogyra</emphasis>
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<bibRefCitation id="EFCC4B30FFECFFB5FE098549FC3C7217" author="BEAUVAIS M." box="[449,604,1293,1319]" pageId="2" pageNumber="23" refId="ref4865" refString="BEAUVAIS M. 1982. - Revision Systematique des Madreporaires des couches de Gosau. Comptoir geologique, Paris, 5 vols." type="book" year="1982">Beauvais, 1982</bibRefCitation>
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</taxonomicName>
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) are characterised (based on the
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<typeStatus id="54E68863FFECFFB5FE4E8569FFD37277" box="[390,435,1325,1351]" pageId="2" pageNumber="23">type</typeStatus>
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material of
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<taxonomicName id="4C5D4D42FFECFFB5FDE18569FF047256" ID-CoL="863N7" authority="Michelin, 1847" authorityName="Michelin" authorityYear="1847" class="Anthozoa" family="Cunnolitidae" genus="Cyclolites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="species" species="haueri">
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<emphasis id="B929EAD3FFECFFB5FDE18569FED27256" italics="true" pageId="2" pageNumber="23">Cyclolites haueri</emphasis>
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Michelin, 1847
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</taxonomicName>
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,
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<typeStatus id="54E68863FFECFFB5FEA78509FFFC7257" box="[367,412,1357,1383]" pageId="2" pageNumber="23">type</typeStatus>
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species of
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<taxonomicName id="4C5D4D42FFECFFB5FDDB8508FC1D7256" ID-CoL="043a1d93-5204-48a5-a347-e9e3270dff5a" authorityName="d'Orbigny" authorityYear="1849" box="[531,637,1356,1382]" class="Anthozoa" family="Cunnolitidae" genus="Funginella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="genus">
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<emphasis id="B929EAD3FFECFFB5FDDB8508FC1D7256" box="[531,637,1356,1382]" italics="true" pageId="2" pageNumber="23">Funginella</emphasis>
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</taxonomicName>
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) by much larger coralla showing septa with larger trabeculae. The fine structure of the septa and presence or absence of synapticulae in these taxa are unknown. The members of the
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<taxonomicName id="4C5D4D42FFECFFB5FE8B858FFE977135" authority="Milne-Edwards & Haime, 1849" authorityName="Milne-Edwards & Haime" authorityYear="1849" class="Anthozoa" family="Cyclolitidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="family">Cyclolitidae Milne-Edwards & Haime, 1849</taxonomicName>
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(suborder
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<taxonomicName id="4C5D4D42FFECFFB5FEA885AFFF947136" authorityName="Morycowa & Roniewicz" authorityYear="1995" box="[352,500,1515,1542]" class="Anthozoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="subOrder" subOrder="Microsolenina">Microsolenina</taxonomicName>
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) are also larger and show pennulae and perforated septa.
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</paragraph>
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<paragraph id="8BE236C1FFECFFB5FFB9860FFF0E7152" blockId="2.[113,647,1611,1700]" box="[113,366,1611,1637]" pageId="2" pageNumber="23">GENERA OF THE FAMILY</paragraph>
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</subSubSection>
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<subSubSection id="C347654AFFECFFB5FFB9862DFA2F759D" pageId="2" pageNumber="23" type="description">
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<paragraph id="8BE236C1FFECFFB5FFB9862DFCE77193" blockId="2.[113,647,1611,1700]" pageId="2" pageNumber="23">
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The family was established for two solitary coral genera of the Cenomanian:
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<taxonomicName id="4C5D4D42FFECFFB5FE4386CDFCE77193" authority="de Fromentel, 1867" authorityName="de Fromentel" authorityYear="1867" box="[395,647,1673,1699]" class="Anthozoa" family="Agariciidae" genus="Asteroseris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="26" phylum="Cnidaria" rank="genus">
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<emphasis id="B929EAD3FFECFFB5FE4386CDFF8F7193" box="[395,495,1673,1699]" italics="true" pageId="2" pageNumber="23">Asteroseris</emphasis>
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de
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<bibRefCitation id="EFCC4B30FFECFFB5FDDC86CDFCE77193" author="FROMENTEL E. DE" box="[532,647,1673,1699]" pageId="2" pageNumber="23" pagination="289 - 336" refId="ref5022" refString="FROMENTEL E. DE 1867. - Zoophytes, terrain cretace (7). Paleontologie francaise (A. d'Orbigny ed.) 8: 289 - 336." type="journal article" year="1867">Fromentel,</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BE236C1FFECFFB5FD6B809CFA2F759D" blockId="2.[675,1212,216,685]" pageId="2" pageNumber="23">
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1867 and
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<taxonomicName id="4C5D4D42FFECFFB5FCC3809DFA2977C2" authority="de Fromentel, 1867" authorityName="de Fromentel" authorityYear="1867" box="[779,1097,216,242]" class="Magnoliopsida" family="Asteraceae" genus="Microseris" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="genus">
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||
<emphasis id="B929EAD3FFECFFB5FCC3809DFD1177C2" box="[779,881,217,242]" italics="true" pageId="2" pageNumber="23">Microseris</emphasis>
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||
de
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||
<bibRefCitation id="EFCC4B30FFECFFB5FC5E809CFA2977C2" author="FROMENTEL E. DE" box="[918,1097,216,242]" pageId="2" pageNumber="23" pagination="289 - 336" refId="ref5022" refString="FROMENTEL E. DE 1867. - Zoophytes, terrain cretace (7). Paleontologie francaise (A. d'Orbigny ed.) 8: 289 - 336." type="journal article" year="1867">Fromentel, 1867</bibRefCitation>
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</taxonomicName>
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. Both genera were monospecific when established. Included here is
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<taxonomicName id="4C5D4D42FFECFFB5FD32815CFA557601" authority="d'Orbigny, 1849" authorityName="d'Orbigny" authorityYear="1849" box="[762,1077,279,306]" class="Anthozoa" family="Asteroseriidae" genus="Actinoseris" higherTaxonomySource="CoL" kingdom="Plantae" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="genus">
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<emphasis id="B929EAD3FFECFFB5FD32815CFD0C7602" box="[762,876,280,306]" italics="true" pageId="2" pageNumber="23">Actinoseris</emphasis>
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d’Orbigny, 1849
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</taxonomicName>
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, which was considered by
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<bibRefCitation id="EFCC4B30FFECFFB5FCF78173FD897662" author="ALLOITEAU J." box="[831,1001,311,338]" pageId="2" pageNumber="23" pagination="1 - 2" refId="ref4776" refString="ALLOITEAU J. 1956. - Actinoseris cenomaniensis. Palaeontologia Universalis, N. S. 142: 1 - 2. Unpublished." type="book chapter" year="1956">Alloiteau (1956</bibRefCitation>
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, unpubl.) to belong to the
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<taxonomicName id="4C5D4D42FFECFFB5FD2D8112FAE97641" authority="Vaughan & Wells, 1943" authorityName="Vaughan & Wells" authorityYear="1943" box="[741,1161,342,369]" class="Anthozoa" family="Siderastraeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="family">Siderastraeidae Vaughan & Wells, 1943</taxonomicName>
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, but by
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<bibRefCitation id="EFCC4B30FFECFFB5FD0A8132FD0F76A1" author="ZLATARSKI V. N." box="[706,879,374,401]" pageId="2" pageNumber="23" pagination="1 - 28" refId="ref5874" refString="ZLATARSKI V. N. 1968. - [Paleobiological aspects of madreporarian corals from the Urgonian in the middle Predbalkan.] Blgarska Akademiya na naukite Geologicheski Institut, Sofia: 1 - 28." type="book chapter" year="1968">Zlatarski (1968)</bibRefCitation>
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to belong to the
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<taxonomicName id="4C5D4D42FFECFFB5FBD78132FAD576A0" authorityName="Alloiteau" authorityYear="1952" box="[1055,1205,374,400]" class="Anthozoa" family="Funginellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="family">Funginellidae</taxonomicName>
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.
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<taxonomicName id="4C5D4D42FFECFFB5FD6B81D2FD6F7680" authorityName="d'Orbigny" authorityYear="1849" box="[675,783,406,432]" class="Anthozoa" family="Asteroseriidae" genus="Actinoseris" higherTaxonomySource="CoL" kingdom="Plantae" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="genus">
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<emphasis id="B929EAD3FFECFFB5FD6B81D2FD6F7680" box="[675,783,406,432]" italics="true" pageId="2" pageNumber="23">Actinoseris</emphasis>
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</taxonomicName>
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was established with two species. More species were later assigned to all three genera. The genera differ by the formation of their columella and the distribution of synapticulae. In
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<taxonomicName id="4C5D4D42FFECFFB5FB9981B1FAD9753F" authorityName="de Fromentel" authorityYear="1867" box="[1105,1209,501,527]" class="Anthozoa" family="Agariciidae" genus="Asteroseris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="genus">
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<emphasis id="B929EAD3FFECFFB5FB9981B1FAD9753F" box="[1105,1209,501,527]" italics="true" pageId="2" pageNumber="23">Asteroseris</emphasis>
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</taxonomicName>
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septa of the first cycle are fused together in the centre of the calice, whereas in both other genera a trabecular columella exist. The distinction between
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<taxonomicName id="4C5D4D42FFECFFB5FD6B8230FD6D75BE" authorityName="d'Orbigny" authorityYear="1849" box="[675,781,628,654]" class="Anthozoa" family="Asteroseriidae" genus="Actinoseris" higherTaxonomySource="CoL" kingdom="Plantae" order="Scleractinia" pageId="2" pageNumber="23" phylum="Cnidaria" rank="genus">
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<emphasis id="B929EAD3FFECFFB5FD6B8230FD6D75BE" box="[675,781,628,654]" italics="true" pageId="2" pageNumber="23">Actinoseris</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C5D4D42FFECFFB5FCF68230FDC475BD" authorityName="de Fromentel" authorityYear="1867" box="[830,932,628,653]" class="Magnoliopsida" family="Asteraceae" genus="Microseris" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="genus">
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<emphasis id="B929EAD3FFECFFB5FCF68230FDC475BD" box="[830,932,628,653]" italics="true" pageId="2" pageNumber="23">Microseris</emphasis>
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</taxonomicName>
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is weak; both genera differ only in the distribution of synapticulae.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |