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<document id="9A1486849D21A22BD206F1C71EB24DB1" ID="10.11646/zootaxa.4188.1.1" ID-DOI="10.11646/zootaxa.4188.1.1" ID-GBIF-Dataset="03a7cb89-747a-411a-855c-f46ad7df47b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="165406" ID-ZooBank="AA85050E-7653-44BE-9330-AC617BFE6DF8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478593795583" checkinUser="plazi" docAuthor="Ng, Heok Hee &amp; Kottelat, Maurice" docDate="2016" docId="03E0BE0EFF865430FF58FAADFCD6523E" docLanguage="en" docName="zootaxa.4188.1.1.pdf" docOrigin="Zootaxa 4188 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Glyptothorax keluk Ng &amp; Kottelat, 2016, new species" docType="treatment" docVersion="11" lastPageNumber="35" masterDocId="FFD9C676FF995413FFCFFFF2FFE65323" masterDocTitle="The Glyptothorax of Sundaland: a revisionary study (Teleostei: Sisoridae)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="32" updateTime="1698392689045" updateUser="plazi">
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<mods:title id="5F8D521EA1FC34821C6B53EFF88DF00C">The Glyptothorax of Sundaland: a revisionary study (Teleostei: Sisoridae)</mods:title>
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<mods:namePart id="29DC83971775A506725A1F09236B0852">Ng, Heok Hee</mods:namePart>
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<mods:namePart id="0A8FD3136B6BE1F403B702CA0B268CBD">Kottelat, Maurice</mods:namePart>
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<mods:title id="706A8EC167AA22ECEE4574DBC5A3562F">Zootaxa</mods:title>
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<mods:date id="7C35B531B106A5A710F186591A772534">2016</mods:date>
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<mods:number id="15B6CCD2389A6E4C829BC85E62373FA3">4188</mods:number>
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<mods:identifier id="BA91068AC4865DC23C0F259B0C698DA5" type="DOI">10.11646/zootaxa.4188.1.1</mods:identifier>
<mods:identifier id="DAB914F484ED2110C4B4E241516EF102" type="GBIF-Dataset">03a7cb89-747a-411a-855c-f46ad7df47b8</mods:identifier>
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<treatment id="03E0BE0EFF865430FF58FAADFCD6523E" ID-DOI="http://doi.org/10.5281/zenodo.6063754" ID-GBIF-Taxon="124624016" ID-Zenodo-Dep="6063754" LSID="urn:lsid:plazi:treatment:03E0BE0EFF865430FF58FAADFCD6523E" httpUri="http://treatment.plazi.org/id/03E0BE0EFF865430FF58FAADFCD6523E" lastPageId="35" lastPageNumber="35" pageId="31" pageNumber="32">
<subSubSection id="C3535C93FF86540CFF58FAADFF1F56B8" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BF60F18FF86540CFF58FAADFDFE5659" blockId="31.[151,536,1375,1435]" box="[151,536,1375,1402]" pageId="31" pageNumber="32">
<heading id="D0BEB874FF86540CFF58FAADFDFE5659" bold="true" box="[151,536,1375,1402]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="B93DD30AFF86540CFF58FAADFDFE5659" bold="true" box="[151,536,1375,1402]" pageId="31" pageNumber="32">
<taxonomicName id="4C49749BFF86540CFF58FAADFE67565A" box="[151,385,1375,1401]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="keluk" status="sp. nov.">
<emphasis id="B93DD30AFF86540CFF58FAADFE67565A" bold="true" box="[151,385,1375,1401]" italics="true" pageId="31" pageNumber="32">Glyptothorax keluk</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E6E71FF86540CFE47FA92FDFE5659" box="[392,536,1376,1402]" pageId="31" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF60F18FF86540CFF58FA70FF1F56B8" blockId="31.[151,536,1375,1435]" box="[151,249,1410,1435]" pageId="31" pageNumber="32">
(
<figureCitation id="1372139DFF86540CFF50FA70FF1656B8" box="[159,240,1410,1435]" captionStart="FIGURE 18" captionStartId="33.[151,250,1654,1676]" captionTargetBox="[151,1435,193,1631]" captionTargetId="figure@33.[151,1436,193,1632]" captionTargetPageId="33" captionText="FIGURE 18. Glyptothorax keluk, MZB 17217, holotype, 52.5 mm SL; Sumatra: Sumatera Selatan, Air Lintang. Dorsal, lateral and ventral views." httpUri="https://zenodo.org/record/266208/files/figure.png" pageId="31" pageNumber="32">Fig. 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF86540CFF58FA38FAA15508" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BF60F18FF86540CFF58FA38FD4C5524" blockId="31.[151,1437,1482,2011]" pageId="31" pageNumber="32">
<materialsCitation id="3B210545FF86540CFF58FA38FC7956C0" ID-GBIF-Occurrence="1324948219" box="[151,927,1482,1507]" collectionCode="MZB" location="Sumatra" pageId="31" pageNumber="32" specimenCode="MZB 17217, 52.5" specimenCount="2" stateProvince="Sumatra" typeStatus="holotype">
<emphasis id="B93DD30AFF86540CFF58FA38FEA856C0" bold="true" box="[151,334,1482,1507]" pageId="31" pageNumber="32">
<typeStatus id="54F2B1BAFF86540CFF58FA38FF3456C0" box="[151,210,1482,1507]" pageId="31" pageNumber="32">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54F2B1BAFF86540CFE92FA38FE2C56C0" box="[349,458,1482,1507]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBEFA763FF86540CFE11FA39FD5E56C0" box="[478,696,1482,1507]" pageId="31" pageNumber="32">
<collectionCode id="ED5897DDFF86540CFE11FA39FDFC56C1" box="[478,538,1483,1506]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="32">MZB</collectionCode>
17217,
<quantity id="4CB1A2FDFF86540CFD48FA38FD5E56C0" box="[647,696,1482,1507]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" pageId="31" pageNumber="32" unit="mm" value="52.5">52.5</quantity>
</specimenCode>
mm SL;
<collectingRegion id="498DC1FAFF86540CFCF2FA39FC7956C0" box="[829,927,1483,1507]" country="Indonesia" name="Sumatera Utara" pageId="31" pageNumber="32">Sumatra</collectingRegion>
</materialsCitation>
:
<materialsCitation id="3B210545FF86540CFC7DFA39FD4C5524" ID-GBIF-Occurrence="1324948226" collectingDate="2006-08-04" collectorName="L. M. Page" latitude="-4.072222" location="Air Lintang" longLatPrecision="21" longitude="103.32361" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Sumatera Selatan" typeStatus="holotype">
<collectingRegion id="498DC1FAFF86540CFC7DFA39FB6756C0" box="[946,1153,1482,1507]" country="Indonesia" name="Sumatera Selatan" pageId="31" pageNumber="32">Sumatera Selatan</collectingRegion>
,
<location id="8E9659C3FF86540CFB5AFA38FAC356C0" LSID="urn:lsid:plazi:treatment:03E0BE0EFF865430FF58FAADFCD6523E:8E9659C3FF86540CFB5AFA38FAC356C0" box="[1173,1317,1482,1507]" latitude="-4.072222" longLatPrecision="21" longitude="103.32361" name="Air Lintang" pageId="31" pageNumber="32" stateProvince="Sumatera Selatan">Air Lintang</location>
,
<geoCoordinate id="EE7D69DFFF86540CFAF6FA39FA7A56C0" box="[1337,1436,1483,1507]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="15" value="-4.072222">4°4'20&quot;S</geoCoordinate>
<geoCoordinate id="EE7D69DFFF86540CFF58FA1CFECF5524" box="[151,297,1518,1543]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="15" value="103.32361">103°19'25&quot;E</geoCoordinate>
;
<collectorName id="26BC6ACEFF86540CFEFBFA1DFE565524" box="[308,432,1519,1543]" pageId="31" pageNumber="32">L. M. Page</collectorName>
<emphasis id="B93DD30AFF86540CFE77FA1DFE165525" box="[440,496,1518,1542]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
,
<date id="FFF729D8FF86540CFE31FA1DFD415524" box="[510,679,1518,1543]" pageId="31" pageNumber="32" value="2006-08-04">
<collectingDate id="EFB3D030FF86540CFE31FA1DFD415524" box="[510,679,1518,1543]" pageId="31" pageNumber="32" value="2006-08-04">4 August 2006</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF60F18FF86540CFF08F9E1FAA15508" blockId="31.[151,1437,1482,2011]" box="[199,1351,1554,1579]" pageId="31" pageNumber="32">
Paratypes: UF 166629 (3),
<quantity id="4CB1A2FDFF86540CFE38F9E0FD7C5509" box="[503,666,1554,1579]" metricMagnitude="-2" metricUnit="m" metricValue="4.6899999999999995" metricValueMax="5.83" metricValueMin="3.55" pageId="31" pageNumber="32" unit="mm" value="46.9" valueMax="58.3" valueMin="35.5">35.558.3 mm</quantity>
SL;
<materialsCitation id="3B210545FF86540CFD1FF9E1FAA25508" ID-GBIF-Occurrence="1324948228" box="[720,1348,1554,1579]" collectionCode="ZRC" pageId="31" pageNumber="32" specimenCount="1">
<collectionCode id="ED5897DDFF86540CFD1FF9E1FCE15508" box="[720,775,1555,1579]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="31" pageNumber="32">ZRC</collectionCode>
54229 (2),
<quantity id="4CB1A2FDFF86540CFC46F9E1FBCA5509" box="[905,1068,1555,1579]" metricMagnitude="-2" metricUnit="m" metricValue="3.925" metricValueMax="4.13" metricValueMin="3.72" pageId="31" pageNumber="32" unit="mm" value="39.25" valueMax="41.3" valueMin="37.2">37.241.3 mm</quantity>
SL; data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF86540CFF08F9C4FC5454B0" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BF60F18FF86540CFF08F9C4FC5454B0" blockId="31.[151,1437,1482,2011]" pageId="31" pageNumber="32">
<emphasis id="B93DD30AFF86540CFF08F9C4FEA7556C" bold="true" box="[199,321,1590,1615]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
<taxonomicName id="4C49749BFF86540CFE85F9C4FDC4556D" box="[330,546,1590,1614]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="keluk">
<emphasis id="B93DD30AFF86540CFE85F9C4FDC4556D" box="[330,546,1590,1614]" italics="true" pageId="31" pageNumber="32">Glyptothorax keluk</emphasis>
</taxonomicName>
differs from Sundaic congeners except
<taxonomicName id="4C49749BFF86540CFC25F9C5FB6E556D" box="[1002,1160,1590,1614]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="platypogon">
<emphasis id="B93DD30AFF86540CFC25F9C5FB6E556D" box="[1002,1160,1590,1614]" italics="true" pageId="31" pageNumber="32">G. platypogon</emphasis>
</taxonomicName>
in having a convex (vs. straight) dorsoposterior margin of the adipose fin (
<figureCitation id="1372139DFF86540CFD1BF9A8FCFC5550" box="[724,794,1626,1651]" captionStart="FIGURE 7" captionStartId="11.[151,250,1642,1664]" captionTargetBox="[150,1435,193,1619]" captionTargetId="figure@11.[150,1436,193,1621]" captionTargetPageId="11" captionText="FIGURE 7. Lateral views of caudal region of: a. Glyptothorax amnestus, ZRC 38708, 54.1 mm SL; b. G. keluk, MZB 17217, holotype, 52.5 mm SL. Illustrations not to scale." httpUri="https://zenodo.org/record/165413/files/figure.png" pageId="31" pageNumber="32">Fig. 7</figureCitation>
). It can be distinguished from
<taxonomicName id="4C49749BFF86540CFBB8F9A9FAF35550" box="[1143,1301,1627,1651]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="platypogon">
<emphasis id="B93DD30AFF86540CFBB8F9A9FAF35550" box="[1143,1301,1627,1651]" italics="true" pageId="31" pageNumber="32">G. platypogon</emphasis>
</taxonomicName>
in having a longer pectoral-fin spine (19.422.9% SL vs. 13.617.9) and a longer caudal fin (26.930.7% SL vs. 20.127.0). The following unique combination of characters further distinguishes
<taxonomicName id="4C49749BFF86540CFC18F951FBD15598" box="[983,1079,1699,1723]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="keluk">
<emphasis id="B93DD30AFF86540CFC18F951FBD15598" box="[983,1079,1699,1723]" italics="true" pageId="31" pageNumber="32">G. keluk</emphasis>
</taxonomicName>
from Sundaic congeners: eye diameter 711% HL; head width 18.820.4% SL; anteromedial striae in thoracic adhesive apparatus absent; predorsal length 36.441.3% SL; dorsal-fin spine length 16.020.1% SL; margin of dorsal fin concave; dorsal-toadipose distance 18.422.5% SL; body depth at anus 14.015.8% SL; adipose-fin base length 13.617.2% SL; post-adipose distance 15.216.7% SL; caudal peduncle depth 7.78.8% SL; premaxillary toothband partially exposed when mouth is closed; anteromedial striae absent in thoracic adhesive apparatus; and uniformly dark body with neither contrasting markings nor prominent pale midlateral stripe.
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF865431FF08F86CFE2550CE" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32" type="description">
<paragraph id="8BF60F18FF865433FF08F86CFD57523F" blockId="31.[151,1437,1482,2011]" lastBlockId="32.[151,1436,151,285]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<emphasis id="B93DD30AFF86540CFF08F86CFEBF5494" bold="true" box="[199,345,1950,1975]" pageId="31" pageNumber="32">Description.</emphasis>
Morphometric data in
<tableCitation id="C6CB3AA3FF86540CFDA1F86CFD2F5494" box="[622,713,1950,1975]" captionStart="TABLE 6" captionStartId="32.[151,239,328,351]" captionTargetBox="[159,1284,378,1718]" captionTargetPageId="32" captionText="TABLE 6. Morphometric data for Glyptothorax keluk (n = 6)." httpUri="http://table.plazi.org/id/DF365F90FFB95433FF58FEBAFCC6527C" pageId="31" pageNumber="32" tableUuid="DF365F90FFB95433FF58FEBAFCC6527C">Table 6</tableCitation>
. Head depressed; body slender, subcylindrical. Dorsal profile rising evenly from tip of snout to origin of dorsal fin, then sloping gently ventrally from origin of dorsal fin to end of caudal peduncle. Ventral profile straight to anal-fin base, then sloping gently dorsally from anal-fin base to end of caudal peduncle. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Skin tuberculate, with tubercles of even size on sides of body. Lateral line complete and midlateral. Vertebrae 17+18=35* (1), 18+17=35 (2) or 18+18=36 (1).
</paragraph>
<caption id="DF365F90FFB95433FF58FEBAFCC6527C" ID-Table-UUID="DF365F90FFB95433FF58FEBAFCC6527C" box="[151,800,327,351]" httpUri="http://table.plazi.org/id/DF365F90FFB95433FF58FEBAFCC6527C" pageId="32" pageNumber="33" targetBox="[159,1284,378,1718]" targetIsTable="true" targetPageId="32">
<paragraph id="8BF60F18FFB95433FF58FEBAFCC6527C" blockId="32.[151,800,327,351]" box="[151,800,327,351]" pageId="32" pageNumber="33">
<emphasis id="B93DD30AFFB95433FF58FEBAFEEC527C" bold="true" box="[151,266,328,351]" pageId="32" pageNumber="33">TABLE 6.</emphasis>
Morphometric data for
<taxonomicName id="4C49749BFFB95433FDC5FEB5FD31527C" box="[522,727,327,351]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="keluk">
<emphasis id="B93DD30AFFB95433FDC5FEB5FD31527C" box="[522,727,327,351]" italics="true" pageId="32" pageNumber="33">Glyptothorax keluk</emphasis>
</taxonomicName>
(n=6).
</paragraph>
</caption>
<paragraph id="8BF60F18FFB95433FD49FE88FB055595" pageId="32" pageNumber="33">
<table id="F949FDB8FFB9ABECFF50FE88FAE25595" box="[159,1284,378,1718]" gridcols="4" gridrows="12" pageId="32" pageNumber="33">
<tr id="35790D5AFFB9ABECFF50FE88FAE2528C" box="[159,1284,378,431]" gridrow="0" pageId="32" pageNumber="33" rowspan-0="1">
<th id="76A86426FFB9ABECFD49FE88FCE4528C" box="[646,770,378,431]" gridcol="1" gridrow="0" pageId="32" pageNumber="33">Holotype MZB 17217</th>
<th id="76A86426FFB9ABECFC5EFE88FC11528C" box="[913,1015,378,431]" gridcol="2" gridrow="0" pageId="32" pageNumber="33">Range</th>
<th id="76A86426FFB9ABECFB53FE88FAE2528C" box="[1180,1284,378,431]" gridcol="3" gridrow="0" pageId="32" pageNumber="33">Mean±SD</th>
</tr>
<tr id="35790D5AFFB9ABECFF50FE3AFAE25124" box="[159,1284,456,519]" gridrow="1" pageId="32" pageNumber="33" rowspan-3="1">
<th id="76A86426FFB9ABECFF50FE3AFE3B5124" box="[159,477,456,519]" gridcol="0" gridrow="1" pageId="32" pageNumber="33">Standard length (mm) %SL</th>
<td id="76A86426FFB9ABECFD49FE3AFCE45124" box="[646,770,456,519]" gridcol="1" gridrow="1" pageId="32" pageNumber="33">52.5</td>
<td id="76A86426FFB9ABECFC5EFE3AFC115124" box="[913,1015,456,519]" gridcol="2" gridrow="1" pageId="32" pageNumber="33">35.558.3</td>
</tr>
<tr id="35790D5AFFB9ABECFF50FDE9FAE251A0" box="[159,1284,539,643]" gridrow="2" pageId="32" pageNumber="33">
<th id="76A86426FFB9ABECFF50FDE9FE3B51A0" box="[159,477,539,643]" gridcol="0" gridrow="2" pageId="32" pageNumber="33">Predorsal length Preanal length Prepelvic length</th>
<td id="76A86426FFB9ABECFD49FDE9FCE451A0" box="[646,770,539,643]" gridcol="1" gridrow="2" pageId="32" pageNumber="33">41.3 66.3 51.8</td>
<td id="76A86426FFB9ABECFC5EFDE9FC1151A0" box="[913,1015,539,643]" gridcol="2" gridrow="2" pageId="32" pageNumber="33">36.441.3 64.668.8 48.052.4</td>
<td id="76A86426FFB9ABECFB53FDE9FAE251A0" box="[1180,1284,539,643]" gridcol="3" gridrow="2" pageId="32" pageNumber="33">38.3±1.90 66.6±1.51 50.7±1.74</td>
</tr>
<tr id="35790D5AFFB9ABECFF50FD65FAE251DC" box="[159,1284,663,767]" gridrow="3" pageId="32" pageNumber="33">
<th id="76A86426FFB9ABECFF50FD65FE3B51DC" box="[159,477,663,767]" gridcol="0" gridrow="3" pageId="32" pageNumber="33">Prepectoral length Length of dorsal-fin base Dorsal-fin spine length</th>
<td id="76A86426FFB9ABECFD49FD65FCE451DC" box="[646,770,663,767]" gridcol="1" gridrow="3" pageId="32" pageNumber="33">22.1 15.0 18.7</td>
<td id="76A86426FFB9ABECFC5EFD65FC1151DC" box="[913,1015,663,767]" gridcol="2" gridrow="3" pageId="32" pageNumber="33">21.224.7 13.015.0 16.020.1</td>
<td id="76A86426FFB9ABECFB53FD65FAE251DC" box="[1180,1284,663,767]" gridcol="3" gridrow="3" pageId="32" pageNumber="33">22.8±1.37 13.9±0.79 18.1±1.50</td>
</tr>
<tr id="35790D5AFFB9ABECFF50FCE1FAE25058" box="[159,1284,787,891]" gridrow="4" pageId="32" pageNumber="33">
<th id="76A86426FFB9ABECFF50FCE1FE3B5058" box="[159,477,787,891]" gridcol="0" gridrow="4" pageId="32" pageNumber="33">Length of anal-fin base Pelvic-fin length Pectoral-fin length</th>
<td id="76A86426FFB9ABECFD49FCE1FCE45058" box="[646,770,787,891]" gridcol="1" gridrow="4" pageId="32" pageNumber="33">15.4 15.0</td>
<td id="76A86426FFB9ABECFC5EFCE1FC115058" box="[913,1015,787,891]" gridcol="2" gridrow="4" pageId="32" pageNumber="33">14.417.7 13.215.8 22.027.5</td>
<td id="76A86426FFB9ABECFB53FCE1FAE25058" box="[1180,1284,787,891]" gridcol="3" gridrow="4" pageId="32" pageNumber="33">15.6±1.26 14.9±1.02 23.8±2.53</td>
</tr>
<tr id="35790D5AFFB9ABECFF50FC7DFAE250D4" box="[159,1284,911,1015]" gridrow="5" pageId="32" pageNumber="33">
<th id="76A86426FFB9ABECFF50FC7DFE3B50D4" box="[159,477,911,1015]" gridcol="0" gridrow="5" pageId="32" pageNumber="33">Pectoral-fin spine length Caudal-fin length Length of adipose-fin base</th>
<td id="76A86426FFB9ABECFD49FC7DFCE450D4" box="[646,770,911,1015]" gridcol="1" gridrow="5" pageId="32" pageNumber="33">22.9 30.7 16.8</td>
<td id="76A86426FFB9ABECFC5EFC7DFC1150D4" box="[913,1015,911,1015]" gridcol="2" gridrow="5" pageId="32" pageNumber="33">19.422.9 26.930.7 13.617.2</td>
<td id="76A86426FFB9ABECFB53FC7DFAE250D4" box="[1180,1284,911,1015]" gridcol="3" gridrow="5" pageId="32" pageNumber="33">21.0±1.79 29.2±1.45 15.9±1.44</td>
</tr>
<tr id="35790D5AFFB9ABECFF50FBF9FAE25750" box="[159,1284,1035,1139]" gridrow="6" pageId="32" pageNumber="33">
<th id="76A86426FFB9ABECFF50FBF9FE3B5750" box="[159,477,1035,1139]" gridcol="0" gridrow="6" pageId="32" pageNumber="33">Dorsal to adipose distance Post-adipose distance Length of caudal peduncle</th>
<td id="76A86426FFB9ABECFD49FBF9FCE45750" box="[646,770,1035,1139]" gridcol="1" gridrow="6" pageId="32" pageNumber="33">22.5 16.4 18.5</td>
<td id="76A86426FFB9ABECFC5EFBF9FC115750" box="[913,1015,1035,1139]" gridcol="2" gridrow="6" pageId="32" pageNumber="33">18.422.5 15.216.7 17.619.6</td>
<td id="76A86426FFB9ABECFB53FBF9FAE25750" box="[1180,1284,1035,1139]" gridcol="3" gridrow="6" pageId="32" pageNumber="33">20.9±2.13 16.1±0.68 18.5±0.73</td>
</tr>
<tr id="35790D5AFFB9ABECFF50FB75FAE257CC" box="[159,1284,1159,1263]" gridrow="7" pageId="32" pageNumber="33">
<th id="76A86426FFB9ABECFF50FB75FE3B57CC" box="[159,477,1159,1263]" gridcol="0" gridrow="7" pageId="32" pageNumber="33">Depth of caudal peduncle Body depth at anus Body depth at dorsal-fin origin</th>
<td id="76A86426FFB9ABECFD49FB75FCE457CC" box="[646,770,1159,1263]" gridcol="1" gridrow="7" pageId="32" pageNumber="33">8.4 15.0 20.8</td>
<td id="76A86426FFB9ABECFC5EFB75FC1157CC" box="[913,1015,1159,1263]" gridcol="2" gridrow="7" pageId="32" pageNumber="33">7.78.8 14.015.8 18.220.8</td>
<td id="76A86426FFB9ABECFB53FB75FAE257CC" box="[1180,1284,1159,1263]" gridcol="3" gridrow="7" pageId="32" pageNumber="33">8.3±0.45 15.0±0.71 19.4±1.30</td>
</tr>
<tr id="35790D5AFFB9ABECFF50FAF1FAE25648" box="[159,1284,1283,1387]" gridrow="8" pageId="32" pageNumber="33">
<th id="76A86426FFB9ABECFF50FAF1FE3B5648" box="[159,477,1283,1387]" gridcol="0" gridrow="8" pageId="32" pageNumber="33">Head length Head width Head depth</th>
<td id="76A86426FFB9ABECFD49FAF1FCE45648" box="[646,770,1283,1387]" gridcol="1" gridrow="8" pageId="32" pageNumber="33">30.5 20.4 15.4</td>
<td id="76A86426FFB9ABECFC5EFAF1FC115648" box="[913,1015,1283,1387]" gridcol="2" gridrow="8" pageId="32" pageNumber="33">26.630.5 18.820.4 13.918.2</td>
<td id="76A86426FFB9ABECFB53FAF1FAE25648" box="[1180,1284,1283,1387]" gridcol="3" gridrow="8" pageId="32" pageNumber="33">28.6±1.53 19.7±0.58 15.5±1.62</td>
</tr>
<tr id="35790D5AFFB9ABECFF50FA8DFAE256C4" box="[159,1284,1407,1511]" gridrow="9" pageId="32" pageNumber="33">
<th id="76A86426FFB9ABECFF50FA8DFE3B56C4" box="[159,477,1407,1511]" gridcol="0" gridrow="9" pageId="32" pageNumber="33">%HL Snout length Interorbital distance</th>
<td id="76A86426FFB9ABECFD49FA8DFCE456C4" box="[646,770,1407,1511]" gridcol="1" gridrow="9" pageId="32" pageNumber="33">51 24</td>
<td id="76A86426FFB9ABECFC5EFA8DFC1156C4" box="[913,1015,1407,1511]" gridcol="2" gridrow="9" pageId="32" pageNumber="33">4552 2431</td>
<td id="76A86426FFB9ABECFB53FA8DFAE256C4" box="[1180,1284,1407,1511]" gridcol="3" gridrow="9" pageId="32" pageNumber="33">49±3.0 27±2.6</td>
</tr>
<tr id="35790D5AFFB9ABECFF50FA09FAE25540" box="[159,1284,1531,1635]" gridrow="10" pageId="32" pageNumber="33">
<th id="76A86426FFB9ABECFF50FA09FE3B5540" box="[159,477,1531,1635]" gridcol="0" gridrow="10" pageId="32" pageNumber="33">Eye diameter Nasal barbel length Maxillary barbel length</th>
<td id="76A86426FFB9ABECFD49FA09FCE45540" box="[646,770,1531,1635]" gridcol="1" gridrow="10" pageId="32" pageNumber="33">9 24 84</td>
<td id="76A86426FFB9ABECFC5EFA09FC115540" box="[913,1015,1531,1635]" gridcol="2" gridrow="10" pageId="32" pageNumber="33">711 2130 7999</td>
<td id="76A86426FFB9ABECFB53FA09FAE25540" box="[1180,1284,1531,1635]" gridcol="3" gridrow="10" pageId="32" pageNumber="33">10±1.6 25±4.0 89±7.8</td>
</tr>
<tr id="35790D5AFFB9ABECFF50F985FAE25595" box="[159,1284,1655,1718]" gridrow="11" pageId="32" pageNumber="33">
<th id="76A86426FFB9ABECFF50F985FE3B5595" box="[159,477,1655,1718]" gridcol="0" gridrow="11" pageId="32" pageNumber="33">Inner mandibular barbel length Outer mandibular barbel length</th>
<td id="76A86426FFB9ABECFD49F985FCE45595" box="[646,770,1655,1718]" gridcol="1" gridrow="11" pageId="32" pageNumber="33">35 49</td>
<td id="76A86426FFB9ABECFC5EF985FC115595" box="[913,1015,1655,1718]" gridcol="2" gridrow="11" pageId="32" pageNumber="33">3036 4354</td>
<td id="76A86426FFB9ABECFB53F985FAE25595" box="[1180,1284,1655,1718]" gridcol="3" gridrow="11" pageId="32" pageNumber="33">33±2.4 48±4.3</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF60F18FFB95433FF08F905FEC25483" blockId="32.[151,1436,1783,1952]" pageId="32" pageNumber="33">Head depressed and broad, triangular when viewed laterally. Snout prominent. Anterior and posterior nares large and separated only by base of nasal barbel. Gill opening broad, extending from ventral margin of posttemporal to isthmus. First branchial arch with 2+6* (3), 2+7 (2), 2+8 (1) or 2+9 (1) rakers. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eye ovoid, horizontal axis longest; located entirely in dorsal half of head.</paragraph>
<caption id="DF365F90FFB85432FF58F984FEB65589" httpUri="https://zenodo.org/record/266208/files/figure.png" pageId="33" pageNumber="34" targetBox="[151,1435,193,1631]" targetPageId="33">
<paragraph id="8BF60F18FFB85432FF58F984FEB65589" blockId="33.[151,1435,1654,1706]" pageId="33" pageNumber="34">
<emphasis id="B93DD30AFFB85432FF58F984FEF955A8" bold="true" box="[151,287,1654,1676]" pageId="33" pageNumber="34">FIGURE 18.</emphasis>
<taxonomicName id="4C49749BFFB85432FEEAF984FE0155AF" box="[293,487,1654,1676]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="keluk">
<emphasis id="B93DD30AFFB85432FEEAF984FE0155AF" box="[293,487,1654,1676]" italics="true" pageId="33" pageNumber="34">Glyptothorax keluk</emphasis>
</taxonomicName>
, MZB 17217, holotype, 52.5 mm SL; Sumatra: Sumatera Selatan, Air Lintang. Dorsal, lateral and ventral views.
</paragraph>
</caption>
<paragraph id="8BF60F18FFB85432FF08F920FD9E5475" blockId="33.[151,1437,1746,2022]" pageId="33" pageNumber="34">Barbels in four pairs. Maxillary barbel long and slender, extending to middle of pectoral-fin base. Nasal barbel slender, extending to midway between its base and anterior orbital margin. Inner mandibular-barbel extending to midway between its base and that of pectoral spine. Outer mandibular barbel extending to two-thirds of distance between its base and that of pectoral spine.</paragraph>
<paragraph id="8BF60F18FFB85432FF08F890FEBA54C5" blockId="33.[151,1437,1746,2022]" pageId="33" pageNumber="34">Mouth inferior, premaxillary tooth band partially (approximately two-thirds) exposed when mouth is closed. Oral teeth small and villiform, in irregular rows on all tooth-bearing surfaces. Premaxillary teeth appearing in single broad semilunate band. Dentary teeth in a single crescentic band, consisting of two separate halves tightly bound at midline.</paragraph>
<paragraph id="8BF60F18FFBB5431FF08FF65FE9A53DB" blockId="34.[151,1436,151,2013]" pageId="34" pageNumber="35">
Thoracic adhesive apparatus consisting of keratinized striae in an elongate ovate field extending from isthmus to just beyond posterior limit of pectoral-fin base (
<figureCitation id="1372139DFFBB5431FD16FF4EFCFB53F6" box="[729,797,188,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1784,1806]" captionTargetBox="[228,1361,193,1761]" captionTargetId="figure@6.[228,1362,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. Thoracic adhesive apparatus of: a. Glyptothorax amnestus, ZRC 54231, paratype, 51.2 mm SL; b. G. decussatus, MZB 17216, holotype, 74.5 mm SL; c. G. exodon, BMNH 1982.3. 29.190, paratype, 63.2 mm SL; d. G. famelicus, USNM 193015, holotype, 73.3 mm SL; e. G. f u s c u s, ZRC 42123, 54.3 mm SL; f. G. keluk, MZB 17217, holotype, 52.5 mm SL; g. G. ketambe, MZB 8694, holotype, 100.0 mm SL; h. G. major, ZRC 38761, 59.1 mm SL; i. G. nieuwenhuisi, MZB 16475, 61.0 mm SL; j. G. pictus, MZB 17218, holotype, 54.2 mm SL; k. G. platypogon, ZRC 48376, 56.3 mm SL; l. G. platypogonides, UF 166626, 52.4 mm SL; m. G. plectilis, ZRC 46420, paratype, 44.3 mm SL; n. G. prashadi, CMK 25369, 50.7 mm SL; o. G. robustus, ZRC 48377, 59.9 mm SL; p. G. schmidti, CMK 4536, 74.6 mm SL; q. G. stibaros, MZB 17215, holotype, 55.3 mm SL. Illustrations not to scale." httpUri="https://zenodo.org/record/266202/files/figure.png" pageId="34" pageNumber="35">Fig. 3</figureCitation>
f). Anteromedial striae absent. Narrow, v-shaped medial pit on posterior half.
</paragraph>
<paragraph id="8BF60F18FFBB5431FF08FEF6FD29511E" blockId="34.[151,1436,151,2013]" pageId="34" pageNumber="35">
Dorsal fin located above anterior third of body, with I,6 (6) rays; fin margin concave; spine short and straight, smooth on anterior and posterior margins. Adipose fin with anterior margin straight or slightly concave and posterior margin convex. Caudal fin strongly forked, with lower lobe slightly longer than upper lobe and i,7,8,i (6) principal rays. Procurrent rays symmetrical and extending only slightly anterior to fin base. Anal-fin base vertically opposite adipose-fin base. Anal fin with straight anterior margin and straight or slightly concave posterior margin; with iv,7,1 (1), iv,8 (2) or iv,8,i* (3) rays. Pelvic-fin origin at vertical through posterior limit of dorsal-fin base. Pelvic fin with slightly convex margin and i,5 (6) rays; tip of adpressed fin not reaching anal-fin origin. Pectoral fin with I,7,i* (3), I,8 (2) or I,8,i (1) rays; posterior fin margin slightly concave; anterior spine margin smooth, posterior margin with 79 (
<typeStatus id="54F2B1BAFFBB5431FE06FDD6FDD6511E" box="[457,560,548,573]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
=8) serrations.
</paragraph>
<paragraph id="8BF60F18FFBB5431FF08FDB5FE2550CE" blockId="34.[151,1436,151,2013]" pageId="34" pageNumber="35">
<emphasis id="B93DD30AFFBB5431FF08FDB5FEA95143" bold="true" box="[199,335,583,608]" pageId="34" pageNumber="35">Coloration.</emphasis>
In 70% ethanol: dorsal and lateral surfaces of head and body dark brownish gray, fading to light gray or beige on ventral surfaces. A thin, light brown mid-dorsal stripe extending from base of last dorsal-fin ray to origin of adipose fin; stripe absent in 2 of 6 individuals examined. Laterosensory pores rimmed in beige, imparting appearance of a diffuse light brown or light gray midlateral line. Dorsal and ventral surfaces of caudal peduncle slightly paler in 2 of 6 individuals examined. All fins with rays proximally gray to grayish brown, becoming hyaline more distally, and diffuse melanophores on fin membranes. Pectoral and pelvic fins with gray or grayish brown on base of fin rays and hyaline posterior margin. Anal fin with gray or grayish brown base; gray or grayish brown spot on anterior third of fin present in 3 of 6 individuals examined. Adipose fin gray or grayish brown with hyaline distal margin. Base of caudal fin with dark crescent in 5 of 6 individuals examined. Each caudal-fin lobe with irregular, elongate gray or grayish brown blotch on about half of outer fin rays; most of inner rays of lobes hyaline. Maxillary and nasal barbels gray or grayish brown dorsally, light brown or light gray ventrally. Mandibular barbels beige or light gray.
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FFBB5431FF08FC05FA765733" box="[199,1424,1015,1040]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BF60F18FFBB5431FF08FC05FA765733" blockId="34.[151,1436,151,2013]" box="[199,1424,1015,1040]" pageId="34" pageNumber="35">
<emphasis id="B93DD30AFFBB5431FF08FC05FE865733" bold="true" box="[199,352,1015,1040]" pageId="34" pageNumber="35">Distribution.</emphasis>
<taxonomicName id="4C49749BFFBB5431FEA8FC0AFDD85733" box="[359,574,1016,1040]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="keluk">
<emphasis id="B93DD30AFFBB5431FEA8FC0AFDD85733" box="[359,574,1016,1040]" italics="true" pageId="34" pageNumber="35">Glyptothorax keluk</emphasis>
</taxonomicName>
is known only from the Musi River drainage in southern
<collectingRegion id="498DC1FAFFBB5431FB0CFC0AFAC65733" box="[1219,1312,1016,1040]" country="Indonesia" name="Sumatera Utara" pageId="34" pageNumber="35">Sumatra</collectingRegion>
(
<figureCitation id="1372139DFFBB5431FAFFFC05FA655733" box="[1328,1411,1015,1040]" captionStart="FIGURE 19" captionStartId="35.[151,250,1403,1425]" captionTargetBox="[151,1436,339,1381]" captionTargetId="figure@35.[151,1436,339,1381]" captionTargetPageId="35" captionText="FIGURE 19. Map showing collection localities of Glyptothorax keluk, G. ketambe and G. major." httpUri="https://zenodo.org/record/165424/files/figure.png" pageId="34" pageNumber="35">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FFBB5430FF08FBEEFCD6523E" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="etymology">
<paragraph id="8BF60F18FFBB5431FF08FBEEFBE3577B" blockId="34.[151,1436,151,2013]" pageId="34" pageNumber="35">
<emphasis id="B93DD30AFFBB5431FF08FBEEFEA85716" bold="true" box="[199,334,1052,1077]" pageId="34" pageNumber="35">Etymology.</emphasis>
The specific epithet comes from the Malay
<taxonomicName id="4C49749BFFBB5431FCAAFBEEFC405716" box="[869,934,1052,1077]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="keluk">keluk</taxonomicName>
, meaning curve, and is used as a noun in apposition in reference to the convex dorsoposterior margin of the adipose fin.
</paragraph>
<paragraph id="8BF60F18FFBB5431FF08FB96FB4D5557" blockId="34.[151,1436,151,2013]" pageId="34" pageNumber="35">
<emphasis id="B93DD30AFFBB5431FF08FB96FE8A575E" bold="true" box="[199,364,1124,1149]" pageId="34" pageNumber="35">Comparisons.</emphasis>
<taxonomicName id="4C49749BFFBB5431FEBAFB96FDA8575F" box="[373,590,1124,1148]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="keluk">
<emphasis id="B93DD30AFFBB5431FEBAFB96FDA8575F" box="[373,590,1124,1148]" italics="true" pageId="34" pageNumber="35">Glyptothorax keluk</emphasis>
</taxonomicName>
superficially resembles
<taxonomicName id="4C49749BFFBB5431FCABFB96FC0B575F" box="[868,1005,1124,1148]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="amnestus">
<emphasis id="B93DD30AFFBB5431FCABFB96FC0B575F" box="[868,1005,1124,1148]" italics="true" pageId="34" pageNumber="35">G. amnestus</emphasis>
</taxonomicName>
in having a uniformly dark body, but can be additionally distinguished from it in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus and having a more slender snout when viewed dorsally (
<figureCitation id="1372139DFFBB5431FCB3FB5EFC5957E6" box="[892,959,1196,1221]" captionStart="FIGURE 6" captionStartId="10.[151,250,1949,1971]" captionTargetBox="[151,1436,193,1948]" captionTargetId="figure@10.[151,1436,193,1997]" captionTargetPageId="10" captionText="FIGURE 6. Dorsal views of head of: a. Glyptothorax amnestus, ZRC 38708, 54.1 mm SL; b. G. keluk, MZB 17217, holotype, 52.5 mm SL. Illustrations not to scale." httpUri="https://zenodo.org/record/165412/files/figure.png" pageId="34" pageNumber="35">Fig. 6</figureCitation>
) and a more slender caudal peduncle (7.7 8.8% SL vs. 9.811.2). There are six other congeners known from
<collectingRegion id="498DC1FAFFBB5431FC4CFB22FC0357CB" box="[899,997,1232,1256]" country="Indonesia" name="Sumatera Utara" pageId="34" pageNumber="35">Sumatra</collectingRegion>
:
<taxonomicName id="4C49749BFFBB5431FC20FB22FB9C57CB" box="[1007,1146,1232,1256]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFFBB5431FC20FB22FB9C57CB" box="[1007,1146,1232,1256]" italics="true" pageId="34" pageNumber="35">G. famelicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFFBB5431FB48FB22FB0A57CB" box="[1159,1260,1232,1256]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="fuscus">
<emphasis id="B93DD30AFFBB5431FB48FB22FB0A57CB" box="[1159,1260,1232,1256]" italics="true" pageId="34" pageNumber="35">G. fuscus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFFBB5431FB35FB22FA9057CB" box="[1274,1398,1232,1256]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="ketambe">
<emphasis id="B93DD30AFFBB5431FB35FB22FA9057CB" box="[1274,1398,1232,1256]" italics="true" pageId="34" pageNumber="35">G. ketambe</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFFBB5431FA4BFB22FEA4562F" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="platypogonides">
<emphasis id="B93DD30AFFBB5431FA4BFB22FEA4562F" italics="true" pageId="34" pageNumber="35">G. platypogonides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFFBB5431FE80FB06FE20562F" box="[335,454,1268,1292]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="plectilis">
<emphasis id="B93DD30AFFBB5431FE80FB06FE20562F" box="[335,454,1268,1292]" italics="true" pageId="34" pageNumber="35">G. plectilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49749BFFBB5431FE33FB06FD9C562F" box="[508,634,1268,1292]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B93DD30AFFBB5431FE33FB06FD9C562F" box="[508,634,1268,1292]" italics="true" pageId="34" pageNumber="35">G. schmidti</emphasis>
</taxonomicName>
.
<taxonomicName id="4C49749BFFBB5431FD48FB06FCB8562F" box="[647,862,1268,1292]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="keluk">
<emphasis id="B93DD30AFFBB5431FD48FB06FCB8562F" box="[647,862,1268,1292]" italics="true" pageId="34" pageNumber="35">Glyptothorax keluk</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C49749BFFBB5431FB8BFB06FB28562F" box="[1092,1230,1268,1292]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFFBB5431FB8BFB06FB28562F" box="[1092,1230,1268,1292]" italics="true" pageId="34" pageNumber="35">G. famelicus</emphasis>
</taxonomicName>
and
<emphasis id="B93DD30AFFBB5431FACBFB06FA9A562F" box="[1284,1404,1268,1292]" italics="true" pageId="34" pageNumber="35">G. ke t a m b e</emphasis>
in lacking (vs. having) a prominent pale midlateral stripe on the body, a greater predorsal length (36.441.3% SL vs. 32.135.9), a longer dorsal-fin spine (16.020.1% SL vs. 8.914.8), a longer pectoral-fin spine (19.422.9% SL vs. 14.016.6) and a shorter post-adipose distance (15.216.7% SL vs. 19.122.0), and from
<emphasis id="B93DD30AFFBB5431FB68FA92FAED565B" box="[1191,1291,1376,1400]" italics="true" pageId="34" pageNumber="35">G. f u s cu s</emphasis>
in having a concave (vs. straight) margin of the dorsal fin and a more slender body (depth at anus 14.015.8% SL vs. 15.6 20.7). It is further distinguished from
<taxonomicName id="4C49749BFFBB5431FDF4FA5AFD5456E3" box="[571,690,1448,1472]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="plectilis">
<emphasis id="B93DD30AFFBB5431FDF4FA5AFD5456E3" box="[571,690,1448,1472]" italics="true" pageId="34" pageNumber="35">G. plectilis</emphasis>
</taxonomicName>
in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus, from
<taxonomicName id="4C49749BFFBB5431FE82FA3EFDF056C7" box="[333,534,1484,1508]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="platypogonides">
<emphasis id="B93DD30AFFBB5431FE82FA3EFDF056C7" box="[333,534,1484,1508]" italics="true" pageId="34" pageNumber="35">G. platypogonides</emphasis>
</taxonomicName>
in having a uniform (vs. mottled) coloration, a smaller eye (diameter 711% HL vs. 1114), a longer adipose-fin base (13.617.2% SL vs. 10.713.7) and a shorter post-adipose distance (15.2 16.7% SL vs. 17.621.5), and from
<taxonomicName id="4C49749BFFBB5431FDFAF9E6FD53550F" box="[565,693,1556,1580]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B93DD30AFFBB5431FDFAF9E6FD53550F" box="[565,693,1556,1580]" italics="true" pageId="34" pageNumber="35">G. schmidti</emphasis>
</taxonomicName>
in lacking (vs. having) a prominent pale midlateral stripe on the body, having a longer dorsal-fin spine (16.020.1% SL vs. 8.914.9), a shorter dorsal-to-adipose distance (18.4 22.5% SL vs. 23.928.1) and a shorter post-adipose distance (15.216.7% SL vs. 18.922.0).
</paragraph>
<paragraph id="8BF60F18FFBB5430FF08F972FCD6523E" blockId="34.[151,1436,151,2013]" lastBlockId="35.[151,1436,151,285]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
Among the remaining Sundaic congeners,
<emphasis id="B93DD30AFFBB5431FD62F972FCE055BB" box="[685,774,1664,1688]" italics="true" pageId="34" pageNumber="35">G. k el u k</emphasis>
further differs from
<taxonomicName id="4C49749BFFBB5431FC37F972FB7755BB" box="[1016,1169,1664,1688]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="decussatus">
<emphasis id="B93DD30AFFBB5431FC37F972FB7755BB" box="[1016,1169,1664,1688]" italics="true" pageId="34" pageNumber="35">G. decussatus</emphasis>
</taxonomicName>
in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus, having a more slender body (depth at anus 14.015.8% SL vs. 17.819.7) and a more slender caudal peduncle (depth 7.78.8% SL vs. 10.310.9), from
<taxonomicName id="4C49749BFFBB5431FB76F93AFACC55C3" box="[1209,1322,1736,1760]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFFBB5431FB76F93AFACC55C3" box="[1209,1322,1736,1760]" italics="true" pageId="34" pageNumber="35">G. exodon</emphasis>
</taxonomicName>
in having the premaxillary toothband approximately two-thirds (vs. almost entirely) exposed when the mouth is closed, a wider head (18.820.4% SL vs. 16.018.1), a shorter post-adipose distance (15.216.7% SL vs. 21.524.0) and a deeper caudal peduncle (7.78.8% SL vs. 6.87.3), and from
<taxonomicName id="4C49749BFFBB5431FC8DF8C6FC45546F" box="[834,931,1844,1868]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="major">
<emphasis id="B93DD30AFFBB5431FC8DF8C6FC45546F" box="[834,931,1844,1868]" italics="true" pageId="34" pageNumber="35">G. major</emphasis>
</taxonomicName>
in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus. It is further distinguished from
<taxonomicName id="4C49749BFFBB5431FCA7F8AAFBFA5453" box="[872,1052,1880,1904]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="nieuwenhuisi">
<emphasis id="B93DD30AFFBB5431FCA7F8AAFBFA5453" box="[872,1052,1880,1904]" italics="true" pageId="34" pageNumber="35">G. nieuwenhuisi</emphasis>
</taxonomicName>
in lacking (vs. having) dark spots on the body, from
<taxonomicName id="4C49749BFFBB5431FEADF88EFE2554B7" box="[354,451,1916,1940]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B93DD30AFFBB5431FEADF88EFE2554B7" box="[354,451,1916,1940]" italics="true" pageId="34" pageNumber="35">G. pictus</emphasis>
</taxonomicName>
in having a uniformly dark body without contrasting markings (vs. paler body with dark vertical bars at adipose-fin base and caudal-fin base), and from
<taxonomicName id="4C49749BFFBB5431FC4BF852FBEC549B" box="[900,1034,1952,1976]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="prashadi">
<emphasis id="B93DD30AFFBB5431FC4BF852FBEC549B" box="[900,1034,1952,1976]" italics="true" pageId="34" pageNumber="35">G. prashadi</emphasis>
</taxonomicName>
in having a concave (vs. straight) margin of the dorsal fin, a more slender body (depth at anus 14.015.8% SL vs. 15.818.8).
<taxonomicName id="4C49749BFFBB5431FB0DF836FA7A54FF" box="[1218,1436,1988,2012]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="keluk">
<emphasis id="B93DD30AFFBB5431FB0DF836FA7A54FF" box="[1218,1436,1988,2012]" italics="true" pageId="34" pageNumber="35">Glyptothorax keluk</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C49749BFFBA5430FEB1FF6AFE195393" box="[382,511,152,176]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B93DD30AFFBA5430FEB1FF6AFE195393" box="[382,511,152,176]" italics="true" pageId="35" pageNumber="36">G. robustus</emphasis>
</taxonomicName>
in having (vs. lacking) a medial pit in the thoracic adhesive apparatus, a longer pectoral-fin spine (19.422.9% SL vs. 16.819.6) and a shorter post-adipose distance (15.216.7% SL vs. 17.0 19.6), and from
<taxonomicName id="4C49749BFFBA5430FE9EFF12FE2C53DB" box="[337,458,224,248]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="stibaros">
<emphasis id="B93DD30AFFBA5430FE9EFF12FE2C53DB" box="[337,458,224,248]" italics="true" pageId="35" pageNumber="36">G. stibaros</emphasis>
</taxonomicName>
in having a smooth (vs. serrated) posterior margin of the dorsal-fin spine and a more slender body (depth at anus 14.015.8% SL vs. 15.620.6).
</paragraph>
</subSubSection>
</treatment>
</document>