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<document id="4F5690E4F692993842C1D65B5C3D6BDF" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2021" docId="03B18797FFCC9D2E4BA503E0DBB2268D" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Benedictesmus timber Shear &amp; Marek 2021, n. sp." docType="treatment" docVersion="5" lastPageNumber="120" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="118" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7A895FABD49215363179DD81FEF98962" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EB7FD5823FF8C3A872E5C81840D2D832">
<mods:title id="B16208DC5D290F9C4A4B2BD930BF3890">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
</mods:titleInfo>
<mods:name id="87EA49BC29F7AFD8C4AF2D0BCF7159C6" type="personal">
<mods:role id="E94218FFE9BF11C9855D499B725159C7">
<mods:roleTerm id="0A5572959022438C8E3869CCE64E5EA9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D0B40717A109DA6ADF26644C1AAA1B0">Shear, William A.</mods:namePart>
<mods:affiliation id="5B35022C5DE22D4199988DBA4150E7DC">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
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<mods:role id="2EF7431D882C027E61BED2D1ADE073F7">
<mods:roleTerm id="CFC4B58381BBDBE1F0A0F0333609B24A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB716E12107CCEE36AB405E9094222D5">Marek, Paul E.</mods:namePart>
<mods:nameIdentifier id="6D366C5EAD72E3D124E9370B0EE459FB" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation id="AE89BA710E31CD017A5DB854D68E5B47">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
<mods:nameIdentifier id="3E23329FA78CD083EDCC9B718C5513FD" type="email">pmarek@vt.edu</mods:nameIdentifier>
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<mods:title id="20C8688F654DE140F79BDB0BAEE29A55">Zootaxa</mods:title>
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<mods:part id="94CDF96E42F6FA8A9010F933B567BE48">
<mods:date id="A113BE124BD4220A8240B8F29F772C55">2021</mods:date>
<mods:detail id="EEBBA81962DBD04719305AC04A73B809" type="pubDate">
<mods:number id="BA288160DF0B331F4F484471FB5E8DBA">2021-05-24</mods:number>
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<mods:detail id="C0AC4C1FBD11FBA4F966ADE2F10507C7" type="volume">
<mods:number id="2395D8243DC2E8A7EA99B1BD18FE3EAA">4975</mods:number>
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<mods:end id="6939DCE47F59689D5FEA3ED378CDA0A3">126</mods:end>
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<mods:classification id="C9C1F31D7419E0D95614AC17596C0E4C">journal article</mods:classification>
<mods:identifier id="7AAAC3BA41F8F578A4FD21E1ADAF903A" type="CLB-Dataset">6194</mods:identifier>
<mods:identifier id="8774C17CA373427481F248C263BBC42A" type="DOI">10.11646/zootaxa.4975.1.3</mods:identifier>
<mods:identifier id="964F67479467945A4FE45ADCB586F1D9" type="GBIF-Dataset">529d48db-4299-4e2d-9aaf-6c4178c08b18</mods:identifier>
<mods:identifier id="201C6775137A0FB744D52956F059651F" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B18797FFCC9D2E4BA503E0DBB2268D" ID-DOI="http://doi.org/10.5281/zenodo.4805067" ID-GBIF-Taxon="182644916" ID-Zenodo-Dep="4805067" LSID="urn:lsid:plazi:treatment:03B18797FFCC9D2E4BA503E0DBB2268D" httpUri="http://treatment.plazi.org/id/03B18797FFCC9D2E4BA503E0DBB2268D" lastPageId="39" lastPageNumber="120" pageId="37" pageNumber="118">
<subSubSection id="C302650AFFCC9D2C4BA503E0DBFD26B8" box="[151,702,1754,1781]" pageId="37" pageNumber="118" type="nomenclature">
<paragraph id="8BA73681FFCC9D2C4BA503E0DBFD26B8" blockId="37.[151,702,1754,1817]" box="[151,702,1754,1781]" pageId="37" pageNumber="118">
<heading id="D0EF81EDFFCC9D2C4BA503E0DBFD26B8" bold="true" box="[151,702,1754,1781]" fontSize="11" level="1" pageId="37" pageNumber="118" reason="1">
<emphasis id="B96CEA93FFCC9D2C4BA503E0DBFD26B8" bold="true" box="[151,702,1754,1781]" pageId="37" pageNumber="118">
<taxonomicName id="4C184D02FFCC9D2C4BA503E0DB2C26B8" authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2021" box="[151,623,1754,1781]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="37" pageNumber="118" phylum="Arthropoda" rank="species" species="timber" status="sp. nov.">
<emphasis id="B96CEA93FFCC9D2C4BA503E0D8E226B8" bold="true" box="[151,417,1754,1781]" italics="true" pageId="37" pageNumber="118">Benedictesmus timber</emphasis>
Shear &amp; Marek
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFCC9D2C494E03E1DBFD26B8" box="[636,702,1755,1781]" pageId="37" pageNumber="118" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFCC9D2C4BA503C4D8DC2754" box="[151,415,1790,1817]" pageId="37" pageNumber="118" type="description">
<paragraph id="8BA73681FFCC9D2C4BA503C4D8DC2754" blockId="37.[151,702,1754,1817]" box="[151,415,1790,1817]" pageId="37" pageNumber="118">
<figureCitation id="13232A04FFCC9D2C4BA503C4D8702754" box="[151,307,1790,1817]" captionStart="FIGURES 117–120" captionStartId="39.[151,264,1199,1223]" captionTargetBox="[151,1436,181,1176]" captionTargetId="figure-17@39.[151,1436,181,1176]" captionTargetPageId="39" captionText="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805069" httpUri="https://zenodo.org/record/4805069/files/figure.png" pageId="37" pageNumber="118">Figs 118–120</figureCitation>
,
<figureCitation id="13232A04FFCC9D2C4A0F03C4D8DC2754" box="[317,415,1790,1817]" captionStart="FIGURES 133–137" captionStartId="42.[151,264,1252,1276]" captionTargetBox="[160,1429,181,1218]" captionTargetId="figure-24@42.[151,1436,181,1228]" captionTargetPageId="42" captionText="FIGURES 133–137. Drawings of gonopods of polydesmid species. 133. Right gonopod of Benedictesmus aureus, n. sp., mesal view. 134. Right gonopod of Benedictesmus ellenae,n. sp., lateral view. 135, Right gonopod of Benedictesmus yaquina, n. sp., lateral view. Fig. 136, 137, Benedictesmus timber.n. sp. 136, Right gonopod, lateral view. 137, Tip of right gonopod, mesal view. Abbreviation: p, pulvillus." figureDoi="http://doi.org/10.5281/zenodo.4805079" httpUri="https://zenodo.org/record/4805079/files/figure.png" pageId="37" pageNumber="118">136, 137</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFCC9D2C4BA5027DD89D27E4" pageId="37" pageNumber="118" type="materials_examined">
<paragraph id="8BA73681FFCC9D2C4BA5027DD89D27E4" blockId="37.[151,1436,1862,2033]" pageId="37" pageNumber="118">
<emphasis id="B96CEA93FFCC9D2C4BA5027DD99A272C" bold="true" box="[151,217,1863,1889]" pageId="37" pageNumber="118">Type.</emphasis>
<materialsCitation id="3B703CDCFFCC9D2C4BD0027CDB82272C" ID-GBIF-Occurrence="3128519337" box="[226,705,1862,1889]" collectionCode="SPRR" pageId="37" pageNumber="118" specimenCount="1" typeStatus="holotype">
Male
<typeStatus id="54A38823FFCC9D2C4A16027CD8C5272D" box="[292,390,1862,1888]" pageId="37" pageNumber="118" type="holotype">holotype</typeStatus>
from
<quantity id="4CE09B64FFCC9D2C4AFC027DDB5B272D" box="[462,536,1862,1889]" metricMagnitude="2" metricUnit="m" metricValue="4.828032" pageId="37" pageNumber="118" unit="mi" value="0.3">0.3 mi</quantity>
west of
<collectionCode id="ED09AE44FFCC9D2C494F027DDB82272C" box="[637,705,1863,1889]" pageId="37" pageNumber="118">SPRR</collectionCode>
</materialsCitation>
<materialsCitation id="3B703CDCFFCC9D2C49F8027DD89927E4" ID-GBIF-Occurrence="3128519302" collectingDate="1971-11-27" collectionCode="CAS" collectorName="E. M. Benedict. The" location="Timber" pageId="37" pageNumber="118" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
overpass on SR26, 3 mi west of
<location id="8EC7605AFFCC9D2C4F73027CDDD7272D" LSID="urn:lsid:plazi:treatment:03B18797FFCC9D2E4BA503E0DBB2268D:8EC7605AFFCC9D2C4F73027CDDD7272D" box="[1089,1172,1862,1888]" name="Timber" pageId="37" pageNumber="118" stateProvince="Washington">Timber</location>
, 45.5251°, -123.2390°,
<collectingRegion id="49DCF863FFCC9D2C4BA50250D85F27C9" box="[151,284,1898,1924]" country="United States of America" name="Washington" pageId="37" pageNumber="118">Washington</collectingRegion>
Co.,
<collectingRegion id="49DCF863FFCC9D2C4A680251D8F227C8" box="[346,433,1899,1925]" country="United States of America" name="Oregon" pageId="37" pageNumber="118">Oregon</collectingRegion>
, collected
<date id="FFA61041FFCC9D2C49180251DA4027C8" box="[554,771,1898,1925]" pageId="37" pageNumber="118" value="1971-11-27">
<collectingDate id="EFE2E9A9FFCC9D2C49180251DA4027C8" box="[554,771,1898,1925]" pageId="37" pageNumber="118" value="1971-11-27">27 November 1971</collectingDate>
</date>
by
<collectorName id="26ED5357FFCC9D2C481D0251DD5727C9" box="[815,1044,1898,1925]" pageId="37" pageNumber="118">E. M. Benedict. The</collectorName>
<typeStatus id="54A38823FFCC9D2C4F290250DD3E27C9" box="[1051,1149,1898,1924]" pageId="37" pageNumber="118" type="holotype">holotype</typeStatus>
is mounted on SEM stub WS33-12, deposited in
<collectionCode id="ED09AE44FFCC9D2C4A9202B5D89927E4" box="[416,474,1935,1961]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="118" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFCC9D2C4BF50288DA4627BC" pageId="37" pageNumber="118" type="diagnosis">
<paragraph id="8BA73681FFCC9D2C4BF50288DA4627BC" blockId="37.[151,1436,1862,2033]" pageId="37" pageNumber="118">
<emphasis id="B96CEA93FFCC9D2C4BF50288D8012781" bold="true" box="[199,322,1970,1996]" pageId="37" pageNumber="118">Diagnosis.</emphasis>
The terminal zone of the gonopods is unique among
<taxonomicName id="4C184D02FFCC9D2C48980288DDE92781" box="[938,1194,1970,1996]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="37" pageNumber="118" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96CEA93FFCC9D2C48980288DD132781" box="[938,1104,1970,1996]" italics="true" pageId="37" pageNumber="118">Benedictesmus</emphasis>
species
</taxonomicName>
in that both terminal processes are bent or curled into hooks (
<figureCitation id="13232A04FFCC9D2C496B02ECDBBB27BC" box="[601,760,2006,2033]" captionStart="FIGURES 133–137" captionStartId="42.[151,264,1252,1276]" captionTargetBox="[160,1429,181,1218]" captionTargetId="figure-24@42.[151,1436,181,1228]" captionTargetPageId="42" captionText="FIGURES 133–137. Drawings of gonopods of polydesmid species. 133. Right gonopod of Benedictesmus aureus, n. sp., mesal view. 134. Right gonopod of Benedictesmus ellenae,n. sp., lateral view. 135, Right gonopod of Benedictesmus yaquina, n. sp., lateral view. Fig. 136, 137, Benedictesmus timber.n. sp. 136, Right gonopod, lateral view. 137, Tip of right gonopod, mesal view. Abbreviation: p, pulvillus." figureDoi="http://doi.org/10.5281/zenodo.4805079" httpUri="https://zenodo.org/record/4805079/files/figure.png" pageId="37" pageNumber="118">Figs 136, 137</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF676609FFCF9D2F4BA502ADDAA127BA" ID-DOI="http://doi.org/10.5281/zenodo.4805063" ID-Zenodo-Dep="4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="38" pageNumber="119" startId="38.[151,264,1943,1967]" targetBox="[170,1418,182,1917]" targetPageId="38">
<paragraph id="8BA73681FFCF9D2F4BA502ADDAA127BA" blockId="38.[151,1437,1943,2040]" pageId="38" pageNumber="119">
<emphasis id="B96CEA93FFCF9D2F4BA502ADD82127E2" bold="true" box="[151,354,1943,1967]" pageId="38" pageNumber="119">FIGURES 111–116</emphasis>
.
<emphasis id="B96CEA93FFCF9D2F4A5D02ADDA6B27E2" bold="true" box="[367,808,1943,1968]" pageId="38" pageNumber="119">
<taxonomicName id="4C184D02FFCF9D2F4A5D02ADDB2627E2" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[367,613,1943,1967]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="119" phylum="Arthropoda" rank="species" species="yaquina" status="n. gen., n. sp.">
<emphasis id="B96CEA93FFCF9D2F4A5D02ADDB2627E2" bold="true" box="[367,613,1943,1967]" italics="true" pageId="38" pageNumber="119">Benedictesmus yaquina</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFCF9D2F494002A2DBBC27FD" box="[626,767,1944,1968]" pageId="38" pageNumber="119" rank="species">n. gen., n. sp.</taxonomicNameLabel>
111
</emphasis>
. Male, lateral view.
<emphasis id="B96CEA93FFCF9D2F48C502ADDD5F27E2" bold="true" box="[1015,1052,1943,1967]" pageId="38" pageNumber="119">112</emphasis>
. Head and anterior rings, dorsal view.
<emphasis id="B96CEA93FFCF9D2F4BA50281D9FF279E" bold="true" box="[151,188,1979,2003]" pageId="38" pageNumber="119">113</emphasis>
. Midbody ring, dorsal view.
<emphasis id="B96CEA93FFCF9D2F4AD20281DB46279E" bold="true" box="[480,517,1979,2003]" pageId="38" pageNumber="119">114</emphasis>
. Pygidium, lateral view.
<emphasis id="B96CEA93FFCF9D2F48320281DA66279E" bold="true" box="[768,805,1979,2003]" pageId="38" pageNumber="119">115</emphasis>
. Gonopods
<emphasis id="B96CEA93FFCF9D2F48AD0286DA9D279E" box="[927,990,1980,2003]" italics="true" pageId="38" pageNumber="119">in situ</emphasis>
, ventral view.
<emphasis id="B96CEA93FFCF9D2F4F400281DDD4279E" bold="true" box="[1138,1175,1979,2003]" pageId="38" pageNumber="119">116</emphasis>
. Same, lateral view. SEM stub WS33-14. Abbreviations:
<emphasis id="B96CEA93FFCF9D2F4AE302DAD89C27B5" bold="true" box="[465,479,2016,2040]" pageId="38" pageNumber="119">p</emphasis>
, pulvillus;
<emphasis id="B96CEA93FFCF9D2F496302DADB2E27B5" bold="true" box="[593,621,2016,2040]" pageId="38" pageNumber="119">pp</emphasis>
, pulvillar process;
<emphasis id="B96CEA93FFCF9D2F480202DADA0727B5" bold="true" box="[816,836,2016,2040]" pageId="38" pageNumber="119">tz</emphasis>
, terminal zone.
</paragraph>
</caption>
<caption id="DF676609FFCE9D2E4BA50195D98D257E" ID-DOI="http://doi.org/10.5281/zenodo.4805069" ID-Zenodo-Dep="4805069" httpUri="https://zenodo.org/record/4805069/files/figure.png" pageId="39" pageNumber="120" startId="39.[151,264,1199,1223]" targetBox="[151,1436,181,1176]" targetPageId="39">
<paragraph id="8BA73681FFCE9D2E4BA50195D98D257E" blockId="39.[151,1437,1199,1331]" pageId="39" pageNumber="120">
<emphasis id="B96CEA93FFCE9D2E4BA50195D82E248A" bold="true" box="[151,365,1199,1223]" pageId="39" pageNumber="120">FIGURES 117–120.</emphasis>
Species of
<emphasis id="B96CEA93FFCE9D2E4AD10195DBC52485" bold="true" box="[483,646,1199,1224]" pageId="39" pageNumber="120">
<taxonomicName id="4C184D02FFCE9D2E4AD10195DB3C248A" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[483,639,1199,1223]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="120" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEA93FFCE9D2E4AD10195DB3C248A" bold="true" box="[483,639,1199,1223]" italics="true" pageId="39" pageNumber="120">Benedictesmus</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25F57E8FFCE9D2E49BF018ADB97248A" box="[653,724,1200,1224]" pageId="39" pageNumber="120" rank="genus">
<emphasis id="B96CEA93FFCE9D2E49BF018ADB8D2485" bold="true" box="[653,718,1200,1224]" pageId="39" pageNumber="120">n. gen</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B96CEA93FFCE9D2E49E90195DA43248A" bold="true" box="[731,768,1199,1223]" pageId="39" pageNumber="120">117</emphasis>
. Gonopod tips of
<emphasis id="B96CEA93FFCE9D2E48880195DDF4248A" box="[954,1207,1199,1223]" italics="true" pageId="39" pageNumber="120">
<taxonomicName id="4C184D02FFCE9D2E48880195DDF2248A" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[954,1201,1199,1223]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="120" phylum="Arthropoda" rank="species" species="yaquina" status="sp. nov.">
<emphasis id="B96CEA93FFCE9D2E48880195DDF2248A" bold="true" box="[954,1201,1199,1223]" italics="true" pageId="39" pageNumber="120">Benedictesmus yaquina</emphasis>
</taxonomicName>
,
</emphasis>
<emphasis id="B96CEA93FFCE9D2E4F8D018ADDBB2485" bold="true" box="[1215,1272,1200,1224]" pageId="39" pageNumber="120">
<taxonomicNameLabel id="A25F57E8FFCE9D2E4F8D018ADDBB2485" box="[1215,1272,1200,1224]" pageId="39" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, posterior view. SEM stub WS33-14. Figs
<emphasis id="B96CEA93FFCE9D2E4AAD01E9D8B524A6" bold="true" box="[415,502,1235,1259]" pageId="39" pageNumber="120">118–120</emphasis>
.
<emphasis id="B96CEA93FFCE9D2E493001E9DA1524A6" bold="true" box="[514,854,1235,1260]" pageId="39" pageNumber="120">
<emphasis id="B96CEA93FFCE9D2E493001E9DBAE24A6" bold="true" box="[514,749,1235,1259]" italics="true" pageId="39" pageNumber="120">
<taxonomicName id="4C184D02FFCE9D2E493001E9DBA424A6" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[514,743,1235,1259]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="120" phylum="Arthropoda" rank="species" species="timber" status="sp. nov.">Benedictesmus timber</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25F57E8FFCE9D2E49C101EEDA6824A1" box="[755,811,1236,1260]" pageId="39" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
118
</emphasis>
. Head and anterior rings, dorsal view.
<emphasis id="B96CEA93FFCE9D2E4FE801E9DDBC24A6" bold="true" box="[1242,1279,1235,1259]" pageId="39" pageNumber="120">119</emphasis>
. Midbody ring, dorsal view.
<emphasis id="B96CEA93FFCE9D2E4A2901CDD8022542" bold="true" box="[283,321,1271,1295]" pageId="39" pageNumber="120">120</emphasis>
. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations:
<emphasis id="B96CEA93FFCE9D2E488D01C2DA8E255D" bold="true" box="[959,973,1272,1296]" pageId="39" pageNumber="120">p</emphasis>
, pulvillus;
<emphasis id="B96CEA93FFCE9D2E4F7101C2DD1C255D" bold="true" box="[1091,1119,1272,1296]" pageId="39" pageNumber="120">pp</emphasis>
, pulvillar process;
<emphasis id="B96CEA93FFCE9D2E4E1401C2DC79255D" bold="true" box="[1318,1338,1272,1296]" pageId="39" pageNumber="120">tz</emphasis>
, terminal zone.
</paragraph>
</caption>
<subSubSection id="C302650AFFCE9D2E4BF50058DC1A2531" box="[199,1369,1378,1404]" pageId="39" pageNumber="120" type="etymology">
<paragraph id="8BA73681FFCE9D2E4BF50058DC1A2531" blockId="39.[151,1437,1378,1728]" box="[199,1369,1378,1404]" pageId="39" pageNumber="120">
<emphasis id="B96CEA93FFCE9D2E4BF50058D80D2531" bold="true" box="[199,334,1378,1404]" pageId="39" pageNumber="120">Etymology.</emphasis>
The species name is a noun in apposition referring to the
<typeStatus id="54A38823FFCE9D2E48E40058DD452531" box="[982,1030,1378,1404]" pageId="39" pageNumber="120">type</typeStatus>
locality near Timber,
<collectingRegion id="49DCF863FFCE9D2E4FCF0058DC162531" box="[1277,1365,1378,1404]" country="United States of America" name="Oregon" pageId="39" pageNumber="120">Oregon</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFCE9D2E4BF500BFDB1526D1" pageId="39" pageNumber="120" type="description">
<paragraph id="8BA73681FFCE9D2E4BF500BFDB1526D1" blockId="39.[151,1437,1378,1728]" pageId="39" pageNumber="120">
<emphasis id="B96CEA93FFCE9D2E4BF500BFD81A25D2" bold="true" box="[199,345,1413,1439]" pageId="39" pageNumber="120">Description.</emphasis>
Male
<typeStatus id="54A38823FFCE9D2E4A9500BCDB4E25ED" box="[423,525,1414,1440]" pageId="39" pageNumber="120" type="holotype">holotype</typeStatus>
. Nineteen rings. Length about 5.0 mm, greatest width
<quantity id="4CE09B64FFCE9D2E4FBB00BCDDB725ED" box="[1161,1268,1414,1441]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="39" pageNumber="120" unit="mm" value="0.52">0.52 mm</quantity>
. Head densely setose, cuticle alveolate (
<figureCitation id="13232A04FFCE9D2E4A820090DB512589" box="[432,530,1450,1476]" captionStart="FIGURES 117–120" captionStartId="39.[151,264,1199,1223]" captionTargetBox="[151,1436,181,1176]" captionTargetId="figure-17@39.[151,1436,181,1176]" captionTargetPageId="39" captionText="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805069" httpUri="https://zenodo.org/record/4805069/files/figure.png" pageId="39" pageNumber="120">Fig. 118</figureCitation>
). Collum (
<figureCitation id="13232A04FFCE9D2E49BA0090DBA42589" box="[648,743,1450,1476]" captionStart="FIGURES 117–120" captionStartId="39.[151,264,1199,1223]" captionTargetBox="[151,1436,181,1176]" captionTargetId="figure-17@39.[151,1436,181,1176]" captionTargetPageId="39" captionText="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805069" httpUri="https://zenodo.org/record/4805069/files/figure.png" pageId="39" pageNumber="120">Fig. 118</figureCitation>
) with anterior marginal row of 18 setae. Anterior metazonites with three rows of setae, transitioning to four rows (
<figureCitation id="13232A04FFCE9D2E49E400F4DA7025A5" box="[726,819,1486,1512]" captionStart="FIGURES 117–120" captionStartId="39.[151,264,1199,1223]" captionTargetBox="[151,1436,181,1176]" captionTargetId="figure-17@39.[151,1436,181,1176]" captionTargetPageId="39" captionText="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805069" httpUri="https://zenodo.org/record/4805069/files/figure.png" pageId="39" pageNumber="120">Fig. 119</figureCitation>
) about ring 7, setal tubercles becoming almost obsolete on midbody to posterior rings. Alveolate cuticle absent from metazonite posterior to collum. Epiproct (
<figureCitation id="13232A04FFCE9D2E4E0100C8DCD72641" box="[1331,1428,1522,1548]" captionStart="FIGURES 117–120" captionStartId="39.[151,264,1199,1223]" captionTargetBox="[151,1436,181,1176]" captionTargetId="figure-17@39.[151,1436,181,1176]" captionTargetPageId="39" captionText="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805069" httpUri="https://zenodo.org/record/4805069/files/figure.png" pageId="39" pageNumber="120">Fig. 120</figureCitation>
) greatly swollen, short, strongly decurved at nearly right angle, without alveolate cuticle. Anterior legs crassate, tarsi with sphaerotrichomes. Gonopod with flattened prefemorite. Acropodite short, robust. Pulvillus (
<emphasis id="B96CEA93FFCE9D2E4FE70300DDA82619" bold="true" box="[1237,1259,1594,1620]" pageId="39" pageNumber="120">p,</emphasis>
<figureCitation id="13232A04FFCE9D2E4FC00300DC132618" box="[1266,1360,1594,1621]" captionStart="FIGURES 133–137" captionStartId="42.[151,264,1252,1276]" captionTargetBox="[160,1429,181,1218]" captionTargetId="figure-24@42.[151,1436,181,1228]" captionTargetPageId="42" captionText="FIGURES 133–137. Drawings of gonopods of polydesmid species. 133. Right gonopod of Benedictesmus aureus, n. sp., mesal view. 134. Right gonopod of Benedictesmus ellenae,n. sp., lateral view. 135, Right gonopod of Benedictesmus yaquina, n. sp., lateral view. Fig. 136, 137, Benedictesmus timber.n. sp. 136, Right gonopod, lateral view. 137, Tip of right gonopod, mesal view. Abbreviation: p, pulvillus." figureDoi="http://doi.org/10.5281/zenodo.4805079" httpUri="https://zenodo.org/record/4805079/files/figure.png" pageId="39" pageNumber="120">Fig. 136</figureCitation>
) distal just below terminal zone, pulvillar process absent. Terminal zone bifurcate, both processes strongly curved, hooklike (
<figureCitation id="13232A04FFCE9D2E4BE203B8D82C26D1" box="[208,367,1666,1692]" captionStart="FIGURES 133–137" captionStartId="42.[151,264,1252,1276]" captionTargetBox="[160,1429,181,1218]" captionTargetId="figure-24@42.[151,1436,181,1228]" captionTargetPageId="42" captionText="FIGURES 133–137. Drawings of gonopods of polydesmid species. 133. Right gonopod of Benedictesmus aureus, n. sp., mesal view. 134. Right gonopod of Benedictesmus ellenae,n. sp., lateral view. 135, Right gonopod of Benedictesmus yaquina, n. sp., lateral view. Fig. 136, 137, Benedictesmus timber.n. sp. 136, Right gonopod, lateral view. 137, Tip of right gonopod, mesal view. Abbreviation: p, pulvillus." figureDoi="http://doi.org/10.5281/zenodo.4805079" httpUri="https://zenodo.org/record/4805079/files/figure.png" pageId="39" pageNumber="120">Figs 136, 137</figureCitation>
). Females unknown.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFCE9D2E4BF5039FDBB2268D" box="[199,753,1701,1728]" pageId="39" pageNumber="120" type="distribution">
<paragraph id="8BA73681FFCE9D2E4BF5039FDBB2268D" blockId="39.[151,1437,1378,1728]" box="[199,753,1701,1728]" pageId="39" pageNumber="120">
<emphasis id="B96CEA93FFCE9D2E4BF5039FD82226F2" bold="true" box="[199,353,1701,1727]" pageId="39" pageNumber="120">Distribution.</emphasis>
Known only from the
<typeStatus id="54A38823FFCE9D2E4952039CDBD3268D" box="[608,656,1702,1728]" pageId="39" pageNumber="120">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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