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<document id="06B7AF6E1C654EF1987D667BF4A026B2" ID-CLB-Dataset="22365" ID-DOI="10.5852/ejt.2018.396" ID-GBIF-Dataset="2b95a851-e1bf-4bdd-9cce-441ddc4135d2" ID-Zenodo-Dep="3787049" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1588696404769" checkinUser="valdenar" docAuthor="Camfield, Rebecca &amp; Hughes, Mark" docDate="2018" docId="0397A96F2F2AFF9F03E9FAA1E4382AE9" docLanguage="en" docName="ejt-396_camfield_hughes.pdf.imf" docOrigin="European Journal of Taxonomy 396" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia palmata var. khasiana Golding &amp; Kareg." docType="treatment" docVersion="6" lastPageNumber="73" masterDocId="FFAED1172F6DFFD7025FFFB1E00A2E5C" masterDocTitle="A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India" masterLastPageNumber="116" masterPageNumber="1" pageNumber="72" updateTime="1698838542593" updateUser="ExternalLinkService">
<mods:mods id="FC299E15A592D6F1F67D18AFBBDB4BC5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4BFADEBF3E46EF542373277C1EC7197F">A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India</mods:title>
</mods:titleInfo>
<mods:name id="2764F4D672F764893C45DEF9EB6398EC" type="personal">
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</mods:role>
<mods:namePart id="C35EB84D25E543C8B43940C54A157A59">Camfield, Rebecca</mods:namePart>
</mods:name>
<mods:name id="039B3DA7BEF10F90B85433CED39D5CBC" type="personal">
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<mods:namePart id="1DECD590153CA4D0E1DEC6868FDF5866">Hughes, Mark</mods:namePart>
</mods:name>
<mods:typeOfResource id="DF4CE1623B7A507B0CB8D6A79C18B227">text</mods:typeOfResource>
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<mods:title id="AFC958D48A52882E80734CD49BE2336E">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="7E6413070017CEA6E42F1B62A8D9F73E">
<mods:date id="BA3E513CCD0EBCE6392871CEC152CE86">2018</mods:date>
<mods:detail id="6CB29BF1534086AD125DBFD314423F13" type="pubDate">
<mods:number id="75800AB3A6A4EC87A25E0B90802539DF">2018-01-19</mods:number>
</mods:detail>
<mods:detail id="E2A792EC6317FB7215A5F4FBB2269B5C" type="volume">
<mods:number id="36449B55961BE3B9AE63BE04FF728D5E">396</mods:number>
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<mods:start id="ED9F5C9F73511B4B31963818D1C006CD">1</mods:start>
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<mods:identifier id="8E043C0442B52458EE4FE0AB09B51C2A" type="CLB-Dataset">22365</mods:identifier>
<mods:identifier id="C74F1C7682450392ADD81FFEF2A688F3" type="DOI">10.5852/ejt.2018.396</mods:identifier>
<mods:identifier id="DD0A236BD0F4FA622AEBE31315FCC72C" type="GBIF-Dataset">2b95a851-e1bf-4bdd-9cce-441ddc4135d2</mods:identifier>
<mods:identifier id="0E5A02A3800F127D4A387BFA9B54639F" type="Zenodo-Dep">3787049</mods:identifier>
</mods:mods>
<treatment id="0397A96F2F2AFF9F03E9FAA1E4382AE9" ID-DOI="http://doi.org/10.5281/zenodo.3794257" ID-GBIF-Taxon="163963871" ID-Zenodo-Dep="3794257" LSID="urn:lsid:plazi:treatment:0397A96F2F2AFF9F03E9FAA1E4382AE9" httpUri="http://treatment.plazi.org/id/0397A96F2F2AFF9F03E9FAA1E4382AE9" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
<subSubSection id="C3244BF22F2AFF9003E9FAA1E5402BEF" pageId="71" pageNumber="72" type="nomenclature">
<paragraph id="8B8118792F2AFF9003E9FAA1E4772B76" blockId="71.[438,1149,1295,1323]" box="[438,1149,1295,1323]" pageId="71" pageNumber="72">
<heading id="D0C9AF152F2AFF9003E9FAA1E4772B76" box="[438,1149,1295,1323]" centered="true" fontSize="11" level="2" pageId="71" pageNumber="72" reason="2">
<taxonomicName id="4C3E63FA2F2AFF9003E9FAA1E4772B76" ID-CoL="7YMLV" authority="(Irmsch.) Golding &amp; Kareg." box="[438,1149,1295,1323]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="71" pageNumber="72" phylum="Tracheophyta" rank="variety" species="palmata" variety="khasiana">
<emphasis id="B94AC46B2F2AFF9003E9FAA1E28F2B76" bold="true" box="[438,645,1295,1322]" italics="true" pageId="71" pageNumber="72">Begonia palmata</emphasis>
var.
<emphasis id="B94AC46B2F2AFF9000E1FABEE3272B76" bold="true" box="[702,813,1295,1322]" italics="true" pageId="71" pageNumber="72">khasiana</emphasis>
(Irmsch.) Golding &amp; Kareg.
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8118792F2AFF9002E2FAE0E5402BEF" blockId="71.[189,1399,1361,1459]" pageId="71" pageNumber="72">
<emphasis id="B94AC46B2F2AFF9002E2FAE0E1342B37" box="[189,318,1361,1387]" italics="true" pageId="71" pageNumber="72">Phytologia</emphasis>
54: 495 (
<bibRefCitation id="EFAF65882F2AFF9003E2FAE0E3382B37" author="Golding J. &amp; Karegeannes C." box="[445,818,1361,1388]" pageId="71" pageNumber="72" pagination="493 - 499" refId="ref53956" refString="Golding J. &amp; Karegeannes C. 1984. Begonia nomenclature Notes, 7. Phytologia 54 (7): 493 - 499. Available from http: // www. biodiversitylibrary. org / page / 12978493 # page / 513 / mode / 1 up [accessed Jun. 2009]." type="journal article" year="1984">Golding &amp; Karegeannes 1984</bibRefCitation>
).
<taxonomicName id="4C3E63FA2F2AFF900139FAE3E1592BD3" authority="Irmsch., Mitteilungen" authorityName="Irmsch., Mitteilungen" authorityYear="1939" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="71" pageNumber="72" phylum="Tracheophyta" rank="subSpecies" species="laciniata" subSpecies="khasiana">
<emphasis id="B94AC46B2F2AFF900139FAE3E4362B37" box="[870,1084,1361,1387]" italics="true" pageId="71" pageNumber="72">Begonia laciniata</emphasis>
subsp.
<emphasis id="B94AC46B2F2AFF9006FFFAE0E5002B37" box="[1184,1290,1361,1387]" italics="true" pageId="71" pageNumber="72">khasiana</emphasis>
Irmsch.,
<emphasis id="B94AC46B2F2AFF9002E2FAC4E1592BD3" box="[189,339,1397,1423]" italics="true" pageId="71" pageNumber="72">Mitteilungen</emphasis>
</taxonomicName>
<emphasis id="B94AC46B2F2AFF900302FAC4E3DD2BD3" box="[349,983,1397,1423]" italics="true" pageId="71" pageNumber="72">aus dem Institut für allgemeine Botanik in Hamburg</emphasis>
10: 529 (
<bibRefCitation id="EFAF65882F2AFF90060FFAC5E50E2BD3" author="Irmscher E." box="[1104,1284,1396,1423]" pageId="71" pageNumber="72" pagination="525 - 534" refId="ref55230" refString="Irmscher E. 1939. Die Begoniaceen Chinas. Mitteilungen aus dem Institut fur allgemeine Botanik in Hamburg 10: 525 - 534." type="journal article" year="1939">Irmscher 1939</bibRefCitation>
).
<typeStatus id="5485A6DB2F2AFF90076BFAC4E5792BD3" box="[1332,1395,1397,1423]" pageId="71" pageNumber="72">Type</typeStatus>
:
<collectingCountry id="F32958E92F2AFF9002E2FA29E0F62BEE" box="[189,252,1432,1458]" name="India" pageId="71" pageNumber="72">India</collectingCountry>
,
<collectingRegion id="49FAD69B2F2AFF900357FA29E1872BEE" box="[264,397,1432,1458]" country="India" name="Meghalaya" pageId="71" pageNumber="72">Meghalaya</collectingRegion>
, Khasia mountains,
<emphasis id="B94AC46B2F2AFF900024FA29E3822BEE" box="[635,904,1432,1458]" italics="true" pageId="71" pageNumber="72">Hooker &amp; Thomson 10</emphasis>
(lecto-: E00265074, here designated).
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F2AFF9002E2FA68E22E28A9" pageId="71" pageNumber="72" type="materials_examined">
<paragraph id="8B8118792F2AFF9002E2FA68E1732BAF" blockId="71.[189,377,1497,1523]" box="[189,377,1497,1523]" pageId="71" pageNumber="72">
<heading id="D0C9AF152F2AFF9002E2FA68E1732BAF" bold="true" box="[189,377,1497,1523]" fontSize="11" level="3" pageId="71" pageNumber="72" reason="3">
<emphasis id="B94AC46B2F2AFF9002E2FA68E1732BAF" bold="true" box="[189,377,1497,1523]" pageId="71" pageNumber="72">Other material</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F2AFF9002E2F9B9E22E28A9" blockId="71.[189,1399,1544,1781]" pageId="71" pageNumber="72">
<materialsCitation id="3B5612242F2AFF9002E2F9B9E22E28A9" ID-GBIF-Occurrence="2610421315" pageId="71" pageNumber="72">
<collectingCountry id="F32958E92F2AFF9002E2F9B9E11F287E" box="[189,277,1544,1570]" name="India" pageId="71" pageNumber="72">INDIA</collectingCountry>
:
<emphasis id="B94AC46B2F2AFF90037DF9B9E212287E" bold="true" box="[290,536,1544,1570]" pageId="71" pageNumber="72">
<collectingRegion id="49FAD69B2F2AFF90037DF9B9E212287E" box="[290,536,1544,1570]" country="India" name="Arunachal-Pradesh" pageId="71" pageNumber="72">Arunachal-Pradesh</collectingRegion>
</emphasis>
:
<date id="FF803EB92F2AFF900076F9B9E2C3287E" box="[553,713,1544,1570]" pageId="71" pageNumber="72" value="2002-10-20">20 Oct. 2002</date>
,
<emphasis id="B94AC46B2F2AFF900089F9B9E444287E" box="[726,1102,1544,1570]" italics="true" pageId="71" pageNumber="72">Hutchinson &amp; Cox et al. 10081</emphasis>
(
<collectionCode id="ED2F80BC2F2AFF90063EF9B9E47D287E" box="[1121,1143,1544,1570]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="71" pageNumber="72" type="Herbarium">E</collectionCode>
).
<emphasis id="B94AC46B2F2AFF9006D1F9B9E4F4287E" bold="true" box="[1166,1278,1544,1570]" pageId="71" pageNumber="72">
<collectingRegion id="49FAD69B2F2AFF9006D1F9B9E4F4287E" box="[1166,1278,1544,1570]" country="India" name="Manipur" pageId="71" pageNumber="72">Manipur</collectingRegion>
</emphasis>
:
<emphasis id="B94AC46B2F2AFF90074FF9B9E0F22819" italics="true" pageId="71" pageNumber="72">Meebold 6395</emphasis>
(syn
<collectionCode id="ED2F80BC2F2AFF900365F99AE1442819" box="[314,334,1579,1605]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="71" pageNumber="72" type="Herbarium">B</collectionCode>
n.v.).
<emphasis id="B94AC46B2F2AFF9003C6F99AE22E2819" bold="true" box="[409,548,1579,1605]" pageId="71" pageNumber="72">
<collectingRegion id="49FAD69B2F2AFF9003C6F99AE22E2819" box="[409,548,1579,1605]" country="India" name="Meghalaya" pageId="71" pageNumber="72">Meghalaya</collectingRegion>
</emphasis>
: Khasi Hills, Cherrapunji,
<date id="FF803EB92F2AFF90013CF99AE3F5281A" box="[867,1023,1579,1606]" pageId="71" pageNumber="72" value="1952-04-15">15 Apr. 1952</date>
,
<emphasis id="B94AC46B2F2AFF900653F99AE4972819" box="[1036,1181,1579,1605]" italics="true" pageId="71" pageNumber="72">Chand 5356</emphasis>
(
<collectionCode id="ED2F80BC2F2AFF9006F2F99AE50A2819" box="[1197,1280,1579,1605]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="71" pageNumber="72" type="Herbarium">MICH</collectionCode>
); ibid.,
<date id="FF803EB92F2AFF900706F99AE1312835" pageId="71" pageNumber="72" value="1952-08-15">15 Aug. 1952</date>
,
<emphasis id="B94AC46B2F2AFF900317F9FFE1E12834" box="[328,491,1614,1640]" italics="true" pageId="71" pageNumber="72">Koelz 31101A</emphasis>
(
<collectionCode id="ED2F80BC2F2AFF9003A4F9FFE2442834" box="[507,590,1614,1640]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="71" pageNumber="72" type="Herbarium">MICH</collectionCode>
); Khasi Hills, Mawphlang,
<date id="FF803EB92F2AFF9001CEF9FFE4142835" box="[913,1054,1614,1641]" pageId="71" pageNumber="72" value="1954-04-08">8 Apr. 1954</date>
,
<emphasis id="B94AC46B2F2AFF900675F9FFE4B12834" box="[1066,1211,1614,1640]" italics="true" pageId="71" pageNumber="72">Chand 7503</emphasis>
(
<collectionCode id="ED2F80BC2F2AFF900694F9FFE5142834" box="[1227,1310,1614,1640]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="71" pageNumber="72" type="Herbarium">MICH</collectionCode>
); Khasi Hills, Mawryngkneng,
<date id="FF803EB92F2AFF90038FF9C0E26428D0" box="[464,622,1649,1676]" pageId="71" pageNumber="72" value="1951-10-18">18 Oct. 1951</date>
,
<emphasis id="B94AC46B2F2AFF900023F9C0E31B28D7" box="[636,785,1649,1675]" italics="true" pageId="71" pageNumber="72">Koelz 28876</emphasis>
(
<collectionCode id="ED2F80BC2F2AFF90017CF9C0E37C28D7" box="[803,886,1649,1675]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="71" pageNumber="72" type="Herbarium">MICH</collectionCode>
); ibid.,
<date id="FF803EB92F2AFF90018EF9C0E46828D0" box="[977,1122,1649,1676]" pageId="71" pageNumber="72" value="1951-09-04">4 Sep. 1951</date>
,
<emphasis id="B94AC46B2F2AFF90062FF9C0E50F28D0" box="[1136,1285,1649,1676]" italics="true" pageId="71" pageNumber="72">Koelz 28337</emphasis>
(
<collectionCode id="ED2F80BC2F2AFF900749F9C0E56328D7" box="[1302,1385,1649,1675]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="71" pageNumber="72" type="Herbarium">MICH</collectionCode>
); Khasi Hills, Pynursla,
<date id="FF803EB92F2AFF900397F924E26028F3" box="[456,618,1685,1711]" pageId="71" pageNumber="72" value="1949-08-17">17 Aug. 1949</date>
,
<emphasis id="B94AC46B2F2AFF900027F924E30128F2" box="[632,779,1684,1711]" italics="true" pageId="71" pageNumber="72">Koelz 23516</emphasis>
(
<collectionCode id="ED2F80BC2F2AFF900143F924E36528F3" box="[796,879,1685,1711]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="71" pageNumber="72" type="Herbarium">MICH</collectionCode>
); Khasia,
<emphasis id="B94AC46B2F2AFF9001BAF924E49228F3" box="[997,1176,1685,1711]" italics="true" pageId="71" pageNumber="72">Clarke 54285B</emphasis>
(syn
<collectionCode id="ED2F80BC2F2AFF900685F924E4E428F3" box="[1242,1262,1685,1711]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="71" pageNumber="72" type="Herbarium">B</collectionCode>
n.v.); ibid.,
<emphasis id="B94AC46B2F2AFF9002E2F909E15D288E" box="[189,343,1720,1746]" italics="true" pageId="71" pageNumber="72">Griffith 2560</emphasis>
(syn
<collectionCode id="ED2F80BC2F2AFF9003C5F909E1A4288E" box="[410,430,1720,1746]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="71" pageNumber="72" type="Herbarium">B</collectionCode>
n.v.); Khasia mountains,
<emphasis id="B94AC46B2F2AFF900080F909E3FA288E" box="[735,1008,1720,1746]" italics="true" pageId="71" pageNumber="72">Hooker &amp; Thomson 10</emphasis>
(
<collectionCode id="ED2F80BC2F2AFF90065EF909E439288E" box="[1025,1075,1720,1746]" country="United Kingdom" name="Bristol Museum" pageId="71" pageNumber="72">BM</collectionCode>
,
<collectionCode id="ED2F80BC2F2AFF90061FF909E453288E" box="[1088,1113,1720,1746]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="71" pageNumber="72" type="Herbarium">K</collectionCode>
); Shillong,
<date id="FF803EB92F2AFF9006BCF909E57B288F" box="[1251,1393,1720,1747]" pageId="71" pageNumber="72" value="1965-04-02">2 Apr. 1965</date>
,
<emphasis id="B94AC46B2F2AFF9002E2F96AE1C628A9" box="[189,460,1755,1781]" italics="true" pageId="71" pageNumber="72">Cox &amp; Hutchinson 306</emphasis>
(
<collectionCode id="ED2F80BC2F2AFF900382F96AE1F828A9" box="[477,498,1755,1781]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="71" pageNumber="72" type="Herbarium">E</collectionCode>
,
<collectionCode id="ED2F80BC2F2AFF9003A1F96AE21D28A9" box="[510,535,1755,1781]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="71" pageNumber="72" type="Herbarium">K</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F2AFF9002E2F8ADE1092993" pageId="71" pageNumber="72" type="description">
<paragraph id="8B8118792F2AFF9002E2F8ADE144296A" blockId="71.[189,334,1820,1846]" box="[189,334,1820,1846]" pageId="71" pageNumber="72">
<heading id="D0C9AF152F2AFF9002E2F8ADE144296A" bold="true" box="[189,334,1820,1846]" fontSize="11" level="3" pageId="71" pageNumber="72" reason="3">
<emphasis id="B94AC46B2F2AFF9002E2F8ADE144296A" bold="true" box="[189,334,1820,1846]" pageId="71" pageNumber="72">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F2AFF9002E2F8FAE1092993" blockId="71.[189,1398,1867,1999]" pageId="71" pageNumber="72">
Stem: red tomentose. Stipules:
<quantity id="4CC6B59C2F2AFF900074F8FAE2BA2939" box="[555,688,1867,1893]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.7" metricValueMin="1.0" pageId="71" pageNumber="72" unit="mm" value="18.5" valueMax="27.0" valueMin="10.0">1027 mm</quantity>
long, red tomentose. Leaves: petiole densely red tomentose; lamina ovate, upper surface with short red pubescence all over, denser on veins, underside red, red pubescence all over, denser on veins; several lobes, long, triangular, not curved; apex acuminate or acute.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F25FF9F02E2FEBDE4382AE9" pageId="72" pageNumber="73" type="discussion">
<paragraph id="8B8118792F25FF9F02E2FEBDE1252F7A" blockId="72.[189,303,268,294]" box="[189,303,268,294]" pageId="72" pageNumber="73">
<heading id="D0C9AF152F25FF9F02E2FEBDE1252F7A" bold="true" box="[189,303,268,294]" fontSize="11" level="3" pageId="72" pageNumber="73" reason="3">
<emphasis id="B94AC46B2F25FF9F02E2FEBDE1252F7A" bold="true" box="[189,303,268,294]" pageId="72" pageNumber="73">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F25FF9F02E2FE8BE1632CCD" blockId="72.[189,1399,314,657]" pageId="72" pageNumber="73">
A very variable species which has a wide distribution throughout Eastern Asia and Indochina. The lobes on the leaf margin vary from elongate acuminate to shortly triangular. Young leaves have very dense hairs which become less dense as the leaf matures. The leaves can also bear markings (
<emphasis id="B94AC46B2F25FF9F0688FE31E5662FC6" box="[1239,1388,384,410]" italics="true" pageId="72" pageNumber="73">Koelz 33555</emphasis>
) ranging from a few simple spots to larger bands of colour similar to
<taxonomicName id="4C3E63FA2F25FF9F01BFFE15E4632FE1" authorityName="K. Koch (Koch" authorityYear="1857" box="[992,1129,419,445]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="annulata">
<emphasis id="B94AC46B2F25FF9F01BFFE15E4632FE1" box="[992,1129,419,445]" italics="true" pageId="72" pageNumber="73">B. annulata</emphasis>
</taxonomicName>
or
<taxonomicName id="4C3E63FA2F25FF9F06CFFE15E4DF2FE1" authorityName="Putz. (Putzeys" authorityYear="1857" box="[1168,1237,420,445]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="rex">
<emphasis id="B94AC46B2F25FF9F06CFFE15E4DF2FE1" box="[1168,1237,420,445]" italics="true" pageId="72" pageNumber="73">B. rex</emphasis>
</taxonomicName>
, although the deeply lobed margin separates
<taxonomicName id="4C3E63FA2F25FF9F0079FE79E2AC2FBD" authorityName="D. Don (Don" authorityYear="1825" box="[550,678,455,481]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F25FF9F0079FE79E2AC2FBD" box="[550,678,455,481]" italics="true" pageId="72" pageNumber="73">B. palmata</emphasis>
</taxonomicName>
from these two species. This species is also vegetatively very similar to
<taxonomicName id="4C3E63FA2F25FF9F036BFE5AE1B52C58" authorityName="H. Hara (Hara" authorityYear="1970" box="[308,447,490,516]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="flaviflora">
<emphasis id="B94AC46B2F25FF9F036BFE5AE1B52C58" box="[308,447,490,516]" italics="true" pageId="72" pageNumber="73">B. flaviflora</emphasis>
</taxonomicName>
, which is distinguished by its smaller yellow flowers.
<taxonomicName id="4C3E63FA2F25FF9F0661FE5AE51B2C58" authorityName="H. Hara (Hara" authorityYear="1970" box="[1086,1297,490,516]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="flaviflora">
<emphasis id="B94AC46B2F25FF9F0661FE5AE51B2C58" box="[1086,1297,490,516]" italics="true" pageId="72" pageNumber="73">Begonia flaviflora</emphasis>
</taxonomicName>
also has shorter stipules and the upper surface of the leaves are almost glabrous with a few sparse hairs on veins, unlike those of
<taxonomicName id="4C3E63FA2F25FF9F032AFD80E1FD2C16" authorityName="D. Don (Don" authorityYear="1825" box="[373,503,560,586]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F25FF9F032AFD80E1FD2C16" box="[373,503,560,586]" italics="true" pageId="72" pageNumber="73">B. palmata</emphasis>
</taxonomicName>
which usually has bristles scattered all over leaf from sparse to very dense. The longest wing on the fruit is only
<quantity id="4CC6B59C2F25FF9F0026FDE5E2BE2C32" box="[633,692,596,622]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="cm" value="1.0">1 cm</quantity>
long in
<taxonomicName id="4C3E63FA2F25FF9F0147FDE5E3AE2C31" authorityName="H. Hara (Hara" authorityYear="1970" box="[792,932,595,621]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="flaviflora">
<emphasis id="B94AC46B2F25FF9F0147FDE5E3AE2C31" box="[792,932,595,621]" italics="true" pageId="72" pageNumber="73">B. flaviflora</emphasis>
</taxonomicName>
and has a crenate tip which is not seen on
<taxonomicName id="4C3E63FA2F25FF9F02BDFDC9E1682CCD" authorityName="D. Don (Don" authorityYear="1825" box="[226,354,631,657]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F25FF9F02BDFDC9E1682CCD" box="[226,354,631,657]" italics="true" pageId="72" pageNumber="73">B. palmata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8118792F25FF9F02E2FD08E4252D60" blockId="72.[189,1399,696,828]" pageId="72" pageNumber="73">
<taxonomicName id="4C3E63FA2F25FF9F02E2FD08E18D2C8E" authorityName="D. Don (Don" authorityYear="1825" box="[189,391,696,722]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F25FF9F02E2FD08E18D2C8E" box="[189,391,696,722]" italics="true" pageId="72" pageNumber="73">Begonia palmata</emphasis>
</taxonomicName>
forms a natural hybrid with
<taxonomicName id="4C3E63FA2F25FF9F0086FD08E3662C8E" authorityName=": Blume" authorityYear="1827" box="[729,876,696,722]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="longifolia">
<emphasis id="B94AC46B2F25FF9F0086FD08E3662C8E" box="[729,876,696,722]" italics="true" pageId="72" pageNumber="73">B. longifolia</emphasis>
</taxonomicName>
where the two are growing together, known as
<taxonomicName id="4C3E63FA2F25FF9F0283FD6DE1342CA9" box="[220,318,732,757]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AC46B2F25FF9F0283FD6DE1342CA9" box="[220,318,732,757]" italics="true" pageId="72" pageNumber="73">Begonia</emphasis>
</taxonomicName>
×
<emphasis id="B94AC46B2F25FF9F0303FD6AE1BE2CA9" box="[348,436,731,757]" italics="true" pageId="72" pageNumber="73">chungii</emphasis>
(
<bibRefCitation id="EFAF65882F25FF9F039CFD6AE2872CA9" author="Peng C. &amp; Ku S. - M." box="[451,653,731,758]" pageId="72" pageNumber="73" pagination="241 - 250" refId="ref56960" refString="Peng C. - I &amp; Ku S. - M. 2009. Begonia × chungii (Begoniaceae), a new natural hybrid in Taiwan. Botanical Studies 50 (2): 241 - 250." type="journal article" year="2009">Peng &amp; Ku 2009</bibRefCitation>
;
<bibRefCitation id="EFAF65882F25FF9F00C6FD6AE3312CA9" author="Morris R." box="[665,827,731,757]" pageId="72" pageNumber="73" refId="ref56663" refString="Morris R. 2011 a. Begonia × chungii natural hybrid in Arunachal-Pradesh. The Begonian 78: 18 - 1." type="journal volume" year="2011">Morris 2011a</bibRefCitation>
). This hybrid has baccate fruit with wings, and is intermediate in form between the two parents. The leaves are deltoid and lobed like
<taxonomicName id="4C3E63FA2F25FF9F06D0FD4EE5052D44" authorityName="D. Don (Don" authorityYear="1825" box="[1167,1295,766,792]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F25FF9F06D0FD4EE5052D44" box="[1167,1295,766,792]" italics="true" pageId="72" pageNumber="73">B. palmata</emphasis>
</taxonomicName>
with red margin and centre, sometimes with light green spots between the red veins.
</paragraph>
<paragraph id="8B8118792F25FF9F02E2FCD2E30F2A56" blockId="72.[189,1399,867,1034]" pageId="72" pageNumber="73">
There are several infra-specific taxa under
<emphasis id="B94AC46B2F25FF9F00E9FCD5E3352D21" box="[694,831,867,893]" italics="true" pageId="72" pageNumber="73">
<taxonomicName id="4C3E63FA2F25FF9F00E9FCD5E3312D21" authorityName="D. Don (Don" authorityYear="1825" box="[694,827,867,893]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="palmata">B. palmata</taxonomicName>
.
</emphasis>
Two are present in the study area, with the
<typeStatus id="5485A6DB2F25FF9F071BFCD5E57D2D22" box="[1348,1399,868,894]" pageId="72" pageNumber="73">type</typeStatus>
<taxonomicName id="4C3E63FA2F25FF9F02E2FC37E15F2DFC" box="[189,341,902,928]" pageId="72" pageNumber="73" rank="variety" variety="being">variety being</taxonomicName>
far more common.
<taxonomicName id="4C3E63FA2F25FF9F0067FC36E3A92DFC" box="[568,931,902,929]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="variety" species="palmata" variety="khasiana">
<emphasis id="B94AC46B2F25FF9F0067FC36E30A2DFC" box="[568,768,902,928]" italics="true" pageId="72" pageNumber="73">Begonia palmata</emphasis>
var.
<emphasis id="B94AC46B2F25FF9F0166FC37E3A92DFC" box="[825,931,902,928]" italics="true" pageId="72" pageNumber="73">khasiana</emphasis>
</taxonomicName>
is distinct for having densely red villose stems, petioles and peduncles. The upper surface of the leaves is quite rough, and feels like sandpaper to the touch when dried, due to the short bristles about
<quantity id="4CC6B59C2F25FF9F0176FC7CE38E2DBB" box="[809,900,973,999]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="72" pageNumber="73" unit="mm" value="200.0">200 µm</quantity>
long, in
<taxonomicName id="4C3E63FA2F25FF9F01B1FC7FE5002DBB" authority="var. palmata" box="[1006,1290,973,999]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="variety" species="palmata" variety="palmata">
<emphasis id="B94AC46B2F25FF9F01B1FC7FE4652DBB" box="[1006,1135,973,999]" italics="true" pageId="72" pageNumber="73">B. palmata</emphasis>
var.
<emphasis id="B94AC46B2F25FF9F06F5FC7CE5002DBB" box="[1194,1290,973,999]" italics="true" pageId="72" pageNumber="73">palmata</emphasis>
</taxonomicName>
the hairs are softer and
<quantity id="4CC6B59C2F25FF9F033BFC41E1F72A56" box="[356,509,1008,1034]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="72" pageNumber="73" unit="mm" value="550.0" valueMax="700.0" valueMin="400.0">400700 µm</quantity>
long (
<bibRefCitation id="EFAF65882F25FF9F0018FC41E2F22A56" author="Irmscher E." box="[583,760,1008,1034]" pageId="72" pageNumber="73" pagination="525 - 534" refId="ref55230" refString="Irmscher E. 1939. Die Begoniaceen Chinas. Mitteilungen aus dem Institut fur allgemeine Botanik in Hamburg 10: 525 - 534." type="journal article" year="1939">Irmscher 1939</bibRefCitation>
).
</paragraph>
<paragraph id="8B8118792F25FF9F02E2FB80E4382AE9" blockId="72.[189,1399,1073,1206]" pageId="72" pageNumber="73">
There is potential for other varieties to be present in the study area, such as
<taxonomicName id="4C3E63FA2F25FF9F0677FB83E2922A33" authority="(Champ. ex Benth.) Golding &amp; Kareg." authorityName="Golding &amp; Kareg." baseAuthorityName="Benth." class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="variety" species="palmata" variety="bowringiana">
<emphasis id="B94AC46B2F25FF9F0677FB83E4A22A17" box="[1064,1192,1073,1099]" italics="true" pageId="72" pageNumber="73">B. palmata</emphasis>
var.
<emphasis id="B94AC46B2F25FF9F06BFFB80E57C2A17" box="[1248,1398,1073,1099]" italics="true" pageId="72" pageNumber="73">bowringiana</emphasis>
(Champ. ex Benth.) Golding &amp; Kareg.
</taxonomicName>
(
<bibRefCitation id="EFAF65882F25FF9F00F3FBE5E4142A33" author="Golding J. &amp; Karegeannes C." box="[684,1054,1108,1135]" pageId="72" pageNumber="73" pagination="493 - 499" refId="ref53956" refString="Golding J. &amp; Karegeannes C. 1984. Begonia nomenclature Notes, 7. Phytologia 54 (7): 493 - 499. Available from http: // www. biodiversitylibrary. org / page / 12978493 # page / 513 / mode / 1 up [accessed Jun. 2009]." type="journal article" year="1984">Golding &amp; Karegeannes 1984</bibRefCitation>
; basionym:
<bibRefCitation id="EFAF65882F25FF9F06EAFBE5E5602A33" author="Bentham G." box="[1205,1386,1108,1135]" pageId="72" pageNumber="73" pagination="116 - 123" refId="ref52537" refString="Bentham G. 1852. Florula Hongkongensis: an enumeration of the plants collected in the island of Hong- Kong, by Major J. G. Champion, 95 th regiment; the determination revised and the new species described by George Bentham, Esq. Hooker's journal of botany and Kew Garden miscellany 4: 116 - 123." type="journal article" year="1852">Bentham 1852</bibRefCitation>
), which are recorded from nearby
<collectingRegion id="49FAD69B2F25FF9F001DFBC9E2AA2ACE" box="[578,672,1144,1170]" country="China" name="Yunnan" pageId="72" pageNumber="73">Yunnan</collectingRegion>
. However, given the variation we see in
<taxonomicName id="4C3E63FA2F25FF9F06DAFBC8E50D2ACE" authorityName="D. Don (Don" authorityYear="1825" box="[1157,1287,1144,1170]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F25FF9F06DAFBC8E50D2ACE" box="[1157,1287,1144,1170]" italics="true" pageId="72" pageNumber="73">B. palmata</emphasis>
</taxonomicName>
applying these names is extremely difficult, and may not be biologically meaningful.
</paragraph>
</subSubSection>
</treatment>
</document>