plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/97/A1/0397A138743CFFA04DBEFDD9FD2AF833.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

1050 lines
104 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="21E6DEC47FB4114ACDE00BD6C5C27175" ID-DOI="10.5281/zenodo.282649" ID-GBIF-Dataset="951e5fe3-bdf1-4b9d-9e7f-4a2c1e0ee2ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="282649" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460545564130" checkinUser="plazi" docAuthor="Jagersbacher-Baumann, Julia &amp; Ebermann, Ernst" docDate="2012" docId="0397A138743CFFA04DBEFDD9FD2AF833" docLanguage="en" docName="zt03520p070.pdf" docOrigin="Zootaxa 3520" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Heterodispus cordidiscus Jagersbacher-Baumann &amp; Ebermann, 2012, n. sp." docType="treatment" docVersion="8" lastPageNumber="64" masterDocId="FFAED940743DFFA84D29FFBBFFEFFFE7" masterDocTitle="Heterodispus cordidiscus n. sp., a new soil mite from Egypt, and remarks on the nominal species H. elongatus Trägårdh, 1905 (Acari: Heterostigmatina: Scutacaridae)" masterLastPageNumber="70" masterPageNumber="56" pageNumber="57" updateTime="1698317682431" updateUser="plazi">
<mods:mods id="61D7325E0834C3FEB699573B603B22FD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="74AFF53DA64DA7CB846848D252F64B4D">
<mods:title id="5D0A97ED162CDCA059AA6F99E67F6FAC">Heterodispus cordidiscus n. sp., a new soil mite from Egypt, and remarks on the nominal species H. elongatus Trägårdh, 1905 (Acari: Heterostigmatina: Scutacaridae)</mods:title>
</mods:titleInfo>
<mods:name id="ADE653774A2662B0E50A265D42F5EEF0" type="personal">
<mods:role id="F3D2DDFB0707B72995965A70F1A5651E">
<mods:roleTerm id="CA647E6675DAC0210171A8836585B796">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8BF83C5E57CB4812F4E10BD717D76E66">Jagersbacher-Baumann, Julia</mods:namePart>
</mods:name>
<mods:name id="7328B9ADB40D3388DCADF03065824D74" type="personal">
<mods:role id="D685DACA2AFA8DEAE4105340A0EFF683">
<mods:roleTerm id="E079C44894CC84722BB0C680BB66BB60">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C666DB41359468C7F4548EA08043ADAE">Ebermann, Ernst</mods:namePart>
</mods:name>
<mods:typeOfResource id="0B5A670B9B6FDECD7562D20C9B8A6811">text</mods:typeOfResource>
<mods:relatedItem id="CBF42ACF2F076B965C44C4268C9902CE" type="host">
<mods:titleInfo id="6A59CDD399E4B5909F4BFF4035192403">
<mods:title id="32453183691FE85663B2CB6CEB592214">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B6AAE48FBE75D8C333CEE4AED6BBBD01">
<mods:date id="BB56BAB474DCB82EA7926F31E3ED09B0">2012</mods:date>
<mods:detail id="769D30FBD8FD5FEAEEF72C71CDF2C859" type="volume">
<mods:number id="D806AE82F94B807062EEB09B905B85B8">3520</mods:number>
</mods:detail>
<mods:extent id="4A63E2279C9549D85EE14B84162494A5" unit="page">
<mods:start id="18EA759E2145757B29FB94F3200B2429">56</mods:start>
<mods:end id="38280EC7E2B1F2E6A4AE102C2C677449">70</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B4EF205B452CDC81C43626EFA1C9720E">journal article</mods:classification>
<mods:identifier id="74865B3C34B8C286A310D05E9CF5D187" type="DOI">10.5281/zenodo.282649</mods:identifier>
<mods:identifier id="42D4A9FBB597CD33129B947889F412CD" type="GBIF-Dataset">951e5fe3-bdf1-4b9d-9e7f-4a2c1e0ee2ca</mods:identifier>
<mods:identifier id="644180692107B149276B41261F58BD43" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0278F143CBF3F4512EF091F969E8829B" type="Zenodo-Dep">282649</mods:identifier>
</mods:mods>
<treatment id="0397A138743CFFA04DBEFDD9FD2AF833" ID-DOI="http://doi.org/10.5281/zenodo.6173572" ID-GBIF-Taxon="119528970" ID-Zenodo-Dep="6173572" LSID="urn:lsid:plazi:treatment:0397A138743CFFA04DBEFDD9FD2AF833" httpUri="http://treatment.plazi.org/id/0397A138743CFFA04DBEFDD9FD2AF833" lastPageId="8" lastPageNumber="64" pageId="1" pageNumber="57">
<subSubSection id="C32443A5743CFFA94DBEFDD9FECFFD7A" pageId="1" pageNumber="57" type="nomenclature">
<paragraph id="8B81102E743CFFA94DBEFDD9FDFFFD9B" blockId="1.[151,528,610,669]" box="[151,528,610,636]" pageId="1" pageNumber="57">
<heading id="D0C9A742743CFFA94DBEFDD9FDFFFD9B" bold="true" box="[151,528,610,636]" fontSize="11" level="1" pageId="1" pageNumber="57" reason="1">
<emphasis id="B94ACC3C743CFFA94DBEFDD9FDFFFD9B" bold="true" box="[151,528,610,636]" pageId="1" pageNumber="57">
<taxonomicName id="4C3E6BAD743CFFA94DBEFDD9FE29FD9B" box="[151,454,610,636]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="cordidiscus" status="sp. nov.">
<emphasis id="B94ACC3C743CFFA94DBEFDD9FE29FD9B" bold="true" box="[151,454,610,636]" italics="true" pageId="1" pageNumber="57">Heterodispus cordidiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2797147743CFFA94CE7FDD9FDFFFD9B" box="[462,528,610,636]" pageId="1" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B81102E743CFFA94DBEFD3EFECFFD7A" blockId="1.[151,528,610,669]" box="[151,288,645,669]" pageId="1" pageNumber="57">
(
<figureCitation id="13050CAB743CFFA94DB6FD3EFEF8FD7A" box="[159,279,645,669]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="2.[151,250,1792,1815]" captionStartId-1="3.[151,250,1755,1778]" captionStartId-10="11.[151,250,1810,1833]" captionStartId-11="12.[151,250,1002,1025]" captionStartId-2="4.[151,250,1188,1211]" captionStartId-3="5.[151,250,1956,1979]" captionStartId-4="6.[151,250,590,613]" captionStartId-5="6.[151,250,2009,2032]" captionStartId-6="7.[151,250,1697,1720]" captionStartId-7="8.[151,250,1579,1602]" captionStartId-8="9.[151,250,1861,1884]" captionStartId-9="10.[151,250,1835,1858]" captionTargetBox-0="[158,1400,186,1791]" captionTargetBox-1="[184,1334,184,1754]" captionTargetBox-10="[227,1393,211,1751]" captionTargetBox-11="[200,1418,200,962]" captionTargetBox-2="[510,1088,186,1160]" captionTargetBox-3="[190,1426,329,1913]" captionTargetBox-4="[161,1421,159,569]" captionTargetBox-5="[185,1384,622,1986]" captionTargetBox-6="[275,1305,196,1663]" captionTargetBox-7="[455,1134,932,1555]" captionTargetBox-8="[191,1397,209,1798]" captionTargetBox-9="[179,1357,196,1793]" captionTargetId-0="figure@2.[151,1436,151,1791]" captionTargetId-1="figure@3.[151,1436,137,1760]" captionTargetId-10="figure@11.[167,1419,193,1789]" captionTargetId-11="figure@12.[151,1436,193,979]" captionTargetId-2="figure@4.[485,1109,161,1169]" captionTargetId-3="figure@5.[160,1445,303,1926]" captionTargetId-4="figure@6.[153,1435,153,575]" captionTargetId-5="figure@6.[185,1396,622,2000]" captionTargetId-6="figure@7.[236,1342,163,1681]" captionTargetId-7="figure@8.[450,1142,926,1562]" captionTargetId-8="figure@9.[151,1436,200,1823]" captionTargetId-9="figure@10.[151,1436,185,1808]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-10="11" captionTargetPageId-11="12" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="6" captionTargetPageId-6="7" captionTargetPageId-7="8" captionTargetPageId-8="9" captionTargetPageId-9="10" captionText-0="FIGURE 1. Heterodispus cordidiscus n. sp. female holotype, dorsal view." captionText-1="FIGURE 2. Heterodispus cordidiscus n. sp. female holotype, ventral view." captionText-10="FIGURE 11. Heterodispus cordidiscus n. sp. male larva, ventral view. The posterior setae ps are covered with particles. gn = gnathosoma." captionText-11="FIGURE 12. Heterodispus cordidiscus n. sp. male larva. a) legI; b) leg II; c) leg III." captionText-2="FIGURE 3. Heterodispus cordidiscus n. sp. female. Examples of the variability in the positions of setae 4 a and 4 b in relation to each other; insertions of the respective specimens are connected by lines." captionText-3="FIGURE 4. Heterodispus cordidiscus n. sp. female holotype. a) leg I; b) leg II; c) leg III; d) leg IV; e) lateral part of prodorsum." captionText-4="FIGURE 5. Heterodispus cordidiscus n. sp. a) gnathosoma of male larva, dorsal view; b) gnathosoma of female, ventral view; c)" captionText-5="FIGURE 6. Heterodispus cordidiscus n. sp. male, dorsal view. ae = aedeagus, dej = ejaculatory duct, sv = seminar vesicle." captionText-6="FIGURE 7. Heterodispus cordidiscus n. sp. male, ventral view. gn = gnathosoma." captionText-7="FIGURE 8. Heterodispus cordidiscus n. sp. male, lateral view of the genital apparatus. ae = aedeagus, dej = ejaculatory duct, lo = internal loop of the aedeagus, sv = seminar vesicle." captionText-8="FIGURE 9. Heterodispus cordidiscus n. sp. male. a) leg I; b) leg II; c) leg III; d) leg IV." captionText-9="FIGURE 10. Heterodispus cordidiscus n. sp. male larva, dorsal view." httpUri-0="https://zenodo.org/record/282650/files/figure.png" httpUri-1="https://zenodo.org/record/282651/files/figure.png" httpUri-10="https://zenodo.org/record/282660/files/figure.png" httpUri-11="https://zenodo.org/record/282661/files/figure.png" httpUri-2="https://zenodo.org/record/282652/files/figure.png" httpUri-3="https://zenodo.org/record/282653/files/figure.png" httpUri-4="https://zenodo.org/record/282654/files/figure.png" httpUri-5="https://zenodo.org/record/282655/files/figure.png" httpUri-6="https://zenodo.org/record/282656/files/figure.png" httpUri-7="https://zenodo.org/record/282657/files/figure.png" httpUri-8="https://zenodo.org/record/282658/files/figure.png" httpUri-9="https://zenodo.org/record/282659/files/figure.png" pageId="1" pageNumber="57">Figs. 1–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32443A5743CFFAD4DBEFD76FD07FF10" lastPageId="5" lastPageNumber="61" pageId="1" pageNumber="57" type="description">
<paragraph id="8B81102E743CFFA94DBEFD76FBBDFCEE" blockId="1.[151,1437,717,2037]" pageId="1" pageNumber="57">
<emphasis id="B94ACC3C743CFFA94DBEFD76FF1AFD02" bold="true" box="[151,245,717,741]" pageId="1" pageNumber="57">Female.</emphasis>
Measurements of
<typeStatus id="5485AE8C743CFFA94CEDFD76FDC5FD02" box="[452,554,717,741]" pageId="1" pageNumber="57" type="holotype">holotype</typeStatus>
: body length 232, body width (measured on the broadest part of tergite C) 168, anterior width of posterior sternal plate (measured between insertions of setae
<emphasis id="B94ACC3C743CFFA9492DFD4AFBF0FCEE" box="[1028,1055,753,777]" italics="true" pageId="1" pageNumber="57">3c</emphasis>
) 71.
</paragraph>
<paragraph id="8B81102E743CFFA94DEEFCADFE99FBE2" blockId="1.[151,1437,717,2037]" pageId="1" pageNumber="57">
<emphasis id="B94ACC3C743CFFA94DEEFCADFECEFCCA" box="[199,289,790,813]" italics="true" pageId="1" pageNumber="57">Dorsum</emphasis>
(
<figureCitation id="13050CAB743CFFA94C1AFCAEFE99FCCA" box="[307,374,789,813]" captionStart="FIGURE 1" captionStartId="2.[151,250,1792,1815]" captionTargetBox="[158,1400,186,1791]" captionTargetId="figure@2.[151,1436,151,1791]" captionTargetPageId="2" captionText="FIGURE 1. Heterodispus cordidiscus n. sp. female holotype, dorsal view." httpUri="https://zenodo.org/record/282650/files/figure.png" pageId="1" pageNumber="57">Fig. 1</figureCitation>
) All dorsal setae barbed,
<emphasis id="B94ACC3C743CFFA94F8FFCAEFD2EFCCA" box="[678,705,789,813]" italics="true" pageId="1" pageNumber="57">c1</emphasis>
,
<emphasis id="B94ACC3C743CFFA94FFBFCAEFD03FCCA" box="[722,748,789,813]" italics="true" pageId="1" pageNumber="57">c2</emphasis>
well developed, with alveolar canal,
<emphasis id="B94ACC3C743CFFA949B6FCAEFB48FCCA" box="[1183,1191,789,813]" italics="true" pageId="1" pageNumber="57">f</emphasis>
,
<emphasis id="B94ACC3C743CFFA94991FCADFB2BFCCA" box="[1208,1220,790,813]" italics="true" pageId="1" pageNumber="57">e</emphasis>
,
<emphasis id="B94ACC3C743CFFA949FCFCAEFB1EFCCA" box="[1237,1265,789,813]" italics="true" pageId="1" pageNumber="57">h1</emphasis>
,
<emphasis id="B94ACC3C743CFFA9482BFCAEFAF0FCCA" box="[1282,1311,789,813]" italics="true" pageId="1" pageNumber="57">h2</emphasis>
thickened. Setae
<emphasis id="B94ACC3C743CFFA94DF2FC82FF1AFCB6" box="[219,245,825,849]" italics="true" pageId="1" pageNumber="57">c1</emphasis>
,
<emphasis id="B94ACC3C743CFFA94C2AFC82FEF1FCB6" box="[259,286,825,849]" italics="true" pageId="1" pageNumber="57">c2</emphasis>
sharply pointed,
<emphasis id="B94ACC3C743CFFA94CC8FC82FE00FCB6" box="[481,495,825,849]" italics="true" pageId="1" pageNumber="57">d</emphasis>
pointed,
<emphasis id="B94ACC3C743CFFA94F71FC81FD8BFCB6" box="[600,612,826,849]" italics="true" pageId="1" pageNumber="57">e</emphasis>
,
<emphasis id="B94ACC3C743CFFA94F5AFC82FD94FCB6" box="[627,635,825,849]" italics="true" pageId="1" pageNumber="57">f</emphasis>
,
<emphasis id="B94ACC3C743CFFA94FA0FC82FD4BFCB6" box="[649,676,825,849]" italics="true" pageId="1" pageNumber="57">h1</emphasis>
and
<emphasis id="B94ACC3C743CFFA94FF5FC82FD18FCB6" box="[732,759,825,849]" italics="true" pageId="1" pageNumber="57">h2</emphasis>
blunt-ended. Average length of dorsal setae:
<emphasis id="B94ACC3C743CFFA949DCFC82FAFFFCB6" box="[1269,1296,825,849]" italics="true" pageId="1" pageNumber="57">c1</emphasis>
35 (n = 19),
<emphasis id="B94ACC3C743CFFA94DBEFCE6FF5EFC92" box="[151,177,861,885]" italics="true" pageId="1" pageNumber="57">c2</emphasis>
38 (n = 20),
<emphasis id="B94ACC3C743CFFA94C6AFCE6FEBEFC92" box="[323,337,861,885]" italics="true" pageId="1" pageNumber="57">d</emphasis>
51 (n = 19),
<emphasis id="B94ACC3C743CFFA94CCDFCE5FE1FFC92" box="[484,496,862,885]" italics="true" pageId="1" pageNumber="57">e</emphasis>
43 (n = 20),
<emphasis id="B94ACC3C743CFFA94FABFCE6FD65FC92" box="[642,650,861,885]" italics="true" pageId="1" pageNumber="57">f</emphasis>
50 (n = 20),
<emphasis id="B94ACC3C743CFFA94E35FCE6FCD7FC92" box="[796,824,861,885]" italics="true" pageId="1" pageNumber="57">h1</emphasis>
35 (n = 20),
<emphasis id="B94ACC3C743CFFA94EE3FCE6FC0AFC92" box="[970,997,861,885]" italics="true" pageId="1" pageNumber="57">h2</emphasis>
37 (n = 20). Average distance between dorsal setae:
<emphasis id="B94ACC3C743CFFA94C00FC3AFE84FC7E" box="[297,363,897,921]" italics="true" pageId="1" pageNumber="57">c1–c1</emphasis>
78 (n = 20),
<emphasis id="B94ACC3C743CFFA94F29FC3AFDC5FC7E" box="[512,554,897,921]" italics="true" pageId="1" pageNumber="57">d–d</emphasis>
51 (n = 20),
<emphasis id="B94ACC3C743CFFA94F94FC3AFD34FC7E" box="[701,731,897,921]" italics="true" pageId="1" pageNumber="57">f–f</emphasis>
58 (n = 20),
<emphasis id="B94ACC3C743CFFA94E47FC3AFC5BFC7E" box="[878,948,897,921]" italics="true" pageId="1" pageNumber="57">h1–h1</emphasis>
26 (n = 20). Measurements of
<typeStatus id="5485AE8C743CFFA94839FC3AFA99FC7E" box="[1296,1398,897,921]" pageId="1" pageNumber="57" type="holotype">holotype</typeStatus>
:
<emphasis id="B94ACC3C743CFFA948A8FC3AFA74FC7E" box="[1409,1435,897,921]" italics="true" pageId="1" pageNumber="57">c1</emphasis>
32,
<emphasis id="B94ACC3C743CFFA94DE8FC1EFF34FC5A" box="[193,219,933,957]" italics="true" pageId="1" pageNumber="57">c2</emphasis>
35
<emphasis id="B94ACC3C743CFFA94C2DFC1EFEFDFC5A" box="[260,274,933,957]" italics="true" pageId="1" pageNumber="57">d</emphasis>
44,
<emphasis id="B94ACC3C743CFFA94C6AFC1DFEA0FC5A" box="[323,335,934,957]" italics="true" pageId="1" pageNumber="57">e</emphasis>
38,
<emphasis id="B94ACC3C743CFFA94CA9FC1EFE67FC5A" box="[384,392,933,957]" italics="true" pageId="1" pageNumber="57">f</emphasis>
47,
<emphasis id="B94ACC3C743CFFA94C91FC1EFE3BFC5A" box="[440,468,933,957]" italics="true" pageId="1" pageNumber="57">h1</emphasis>
32,
<emphasis id="B94ACC3C743CFFA94F2CFC1EFDCEFC5A" box="[517,545,933,957]" italics="true" pageId="1" pageNumber="57">h2</emphasis>
34;
<emphasis id="B94ACC3C743CFFA94F7BFC1EFD7AFC5A" box="[594,661,933,957]" italics="true" pageId="1" pageNumber="57">c1–c1</emphasis>
67,
<emphasis id="B94ACC3C743CFFA94FECFC1EFD1FFC5A" box="[709,752,933,957]" italics="true" pageId="1" pageNumber="57">d–d</emphasis>
71,
<emphasis id="B94ACC3C743CFFA94E09FC1EFCD1FC5A" box="[800,830,933,957]" italics="true" pageId="1" pageNumber="57">f–f</emphasis>
50,
<emphasis id="B94ACC3C743CFFA94E47FC1EFC5BFC5A" box="[878,948,933,957]" italics="true" pageId="1" pageNumber="57">h1–h1</emphasis>
20. Posterior margin of tergits C, D and EF smooth or scalloped. Cupulae
<emphasis id="B94ACC3C743CFFA94CD2FC71FDFEFC06" box="[507,529,970,993]" italics="true" pageId="1" pageNumber="57">ia</emphasis>
and
<emphasis id="B94ACC3C743CFFA94F79FC72FD89FC06" box="[592,614,969,993]" italics="true" pageId="1" pageNumber="57">ih</emphasis>
roundish. Body surface finely stippled, margin of tergite C with fine, longitudinal stripes.
</paragraph>
<paragraph id="8B81102E743CFFA94DEEFBA9FAB8FAC2" blockId="1.[151,1437,717,2037]" pageId="1" pageNumber="57">
<emphasis id="B94ACC3C743CFFA94DEEFBA9FEE3FBCE" box="[199,268,1042,1065]" italics="true" pageId="1" pageNumber="57">Venter</emphasis>
(
<figureCitation id="13050CAB743CFFA94C34FBAAFE8EFBCE" box="[285,353,1041,1065]" captionStart="FIGURE 2" captionStartId="3.[151,250,1755,1778]" captionTargetBox="[184,1334,184,1754]" captionTargetId="figure@3.[151,1436,137,1760]" captionTargetPageId="3" captionText="FIGURE 2. Heterodispus cordidiscus n. sp. female holotype, ventral view." httpUri="https://zenodo.org/record/282651/files/figure.png" pageId="1" pageNumber="57">Fig. 2</figureCitation>
) Ap1, 2 well developed, ap3 very broad, ap4, 5 weakly developed. Ventral plates finely stippled. Setae
<emphasis id="B94ACC3C743CFFA94DF4FB8EFF15FBAA" box="[221,250,1077,1101]" italics="true" pageId="1" pageNumber="57">1a</emphasis>
,
<emphasis id="B94ACC3C743CFFA94C23FB8EFEC8FBAA" box="[266,295,1077,1101]" italics="true" pageId="1" pageNumber="57">1b</emphasis>
,
<emphasis id="B94ACC3C743CFFA94C1EFB8EFEBCFBAA" box="[311,339,1077,1101]" italics="true" pageId="1" pageNumber="57">2a</emphasis>
,
<emphasis id="B94ACC3C743CFFA94C4DFB8EFE6FFBAA" box="[356,384,1077,1101]" italics="true" pageId="1" pageNumber="57">2b</emphasis>
barbed.
<emphasis id="B94ACC3C743CFFA94CCEFB8EFDECFBAA" box="[487,515,1077,1101]" italics="true" pageId="1" pageNumber="57">3a</emphasis>
,
<emphasis id="B94ACC3C743CFFA94F3DFB8EFDDFFBAA" box="[532,560,1077,1101]" italics="true" pageId="1" pageNumber="57">3b</emphasis>
long, smooth, barbed,
<emphasis id="B94ACC3C743CFFA94E14FB8EFCB7FBAA" box="[829,856,1077,1101]" italics="true" pageId="1" pageNumber="57">3c</emphasis>
thicker, barbed.
<emphasis id="B94ACC3C743CFFA94935FB8EFBD6FBAA" box="[1052,1081,1077,1101]" italics="true" pageId="1" pageNumber="57">4b</emphasis>
inserted anteriorly of
<emphasis id="B94ACC3C743CFFA94814FB8EFAB6FBAA" box="[1341,1369,1077,1101]" italics="true" pageId="1" pageNumber="57">4a</emphasis>
, both barbed. Positions of insertions of
<emphasis id="B94ACC3C743CFFA94F25FBE2FDC6FB96" box="[524,553,1113,1137]" italics="true" pageId="1" pageNumber="57">4a</emphasis>
and
<emphasis id="B94ACC3C743CFFA94F74FBE2FD95FB96" box="[605,634,1113,1137]" italics="true" pageId="1" pageNumber="57">4b</emphasis>
may vary (
<figureCitation id="13050CAB743CFFA94FDEFBE2FCD4FB96" box="[759,827,1113,1137]" captionStart="FIGURE 3" captionStartId="4.[151,250,1188,1211]" captionTargetBox="[510,1088,186,1160]" captionTargetId="figure@4.[485,1109,161,1169]" captionTargetPageId="4" captionText="FIGURE 3. Heterodispus cordidiscus n. sp. female. Examples of the variability in the positions of setae 4 a and 4 b in relation to each other; insertions of the respective specimens are connected by lines." httpUri="https://zenodo.org/record/282652/files/figure.png" pageId="1" pageNumber="57">Fig. 3</figureCitation>
).
<emphasis id="B94ACC3C743CFFA94E66FBE2FC85FB96" box="[847,874,1113,1137]" italics="true" pageId="1" pageNumber="57">4c</emphasis>
thick, barbed, straight.
<emphasis id="B94ACC3C743CFFA94958FBE2FB63FB96" box="[1137,1164,1113,1137]" italics="true" pageId="1" pageNumber="57">2b</emphasis>
,
<emphasis id="B94ACC3C743CFFA949B3FBE2FB5BFB96" box="[1178,1204,1113,1137]" italics="true" pageId="1" pageNumber="57">3c</emphasis>
and
<emphasis id="B94ACC3C743CFFA949C3FBE2FAEBFB96" box="[1258,1284,1113,1137]" italics="true" pageId="1" pageNumber="57">4c</emphasis>
blunt–ended, all other ventral setae sharply pointed. Pseudanal setae
<emphasis id="B94ACC3C743CFFA94E2FFBC6FCC2FB72" box="[774,813,1149,1173]" italics="true" pageId="1" pageNumber="57">ps1</emphasis>
,
<emphasis id="B94ACC3C743CFFA94E15FBC6FC8CFB72" box="[828,867,1149,1173]" italics="true" pageId="1" pageNumber="57">ps2</emphasis>
&gt;
<emphasis id="B94ACC3C743CFFA94EABFBC6FC47FB72" box="[898,936,1149,1173]" italics="true" pageId="1" pageNumber="57">ps3</emphasis>
.
<emphasis id="B94ACC3C743CFFA94E9EFBC6FC31FB72" box="[951,990,1149,1173]" italics="true" pageId="1" pageNumber="57">ps1</emphasis>
,
<emphasis id="B94ACC3C743CFFA94EC5FBC6FBFCFB72" box="[1004,1043,1149,1173]" italics="true" pageId="1" pageNumber="57">ps2</emphasis>
thickened, barbed. Average length of ventral setae (n = 20):
<emphasis id="B94ACC3C743CFFA94C95FB1AFE38FB5E" box="[444,471,1185,1209]" italics="true" pageId="1" pageNumber="57">1a</emphasis>
35,
<emphasis id="B94ACC3C743CFFA94F25FB1AFDF6FB5E" box="[524,537,1185,1209]" italics="true" pageId="1" pageNumber="57">1</emphasis>
b 33,
<emphasis id="B94ACC3C743CFFA94F75FB1AFD85FB5E" box="[604,618,1185,1209]" italics="true" pageId="1" pageNumber="57">2</emphasis>
a 33,
<emphasis id="B94ACC3C743CFFA94F85FB1AFD55FB5E" box="[684,698,1185,1209]" italics="true" pageId="1" pageNumber="57">2</emphasis>
b 48,
<emphasis id="B94ACC3C743CFFA94FD5FB1AFCF7FB5E" box="[764,792,1185,1209]" italics="true" pageId="1" pageNumber="57">3a</emphasis>
65,
<emphasis id="B94ACC3C743CFFA94E64FB1AFCB4FB5E" box="[845,859,1185,1209]" italics="true" pageId="1" pageNumber="57">3</emphasis>
b 61,
<emphasis id="B94ACC3C743CFFA94EB4FB1AFC44FB5E" box="[925,939,1185,1209]" italics="true" pageId="1" pageNumber="57">3</emphasis>
c 50,
<emphasis id="B94ACC3C743CFFA94EC5FB1AFBE7FB5E" box="[1004,1032,1185,1209]" italics="true" pageId="1" pageNumber="57">4a</emphasis>
71,
<emphasis id="B94ACC3C743CFFA94915FB1AFBA4FB5E" box="[1084,1099,1185,1209]" italics="true" pageId="1" pageNumber="57">4</emphasis>
b 77,
<emphasis id="B94ACC3C743CFFA949A5FB1AFB74FB5E" box="[1164,1179,1185,1209]" italics="true" pageId="1" pageNumber="57">4</emphasis>
c 58. Average distance between ventral setae (n = 20):
<emphasis id="B94ACC3C743CFFA94CDCFB7EFDD4FB3A" box="[501,571,1221,1245]" italics="true" pageId="1" pageNumber="57">1a–1a</emphasis>
20,
<emphasis id="B94ACC3C743CFFA94F42FB7EFD5DFB3A" box="[619,690,1221,1245]" italics="true" pageId="1" pageNumber="57">1b–1b</emphasis>
53,
<emphasis id="B94ACC3C743CFFA94FC8FB7EFD00FB3A" box="[737,751,1221,1245]" italics="true" pageId="1" pageNumber="57">2</emphasis>
a–2a 39,
<emphasis id="B94ACC3C743CFFA94E7EFB7EFC8AFB3A" box="[855,869,1221,1245]" italics="true" pageId="1" pageNumber="57">2</emphasis>
b–2b 38,
<emphasis id="B94ACC3C743CFFA94EE4FB7EFC34FB3A" box="[973,987,1221,1245]" italics="true" pageId="1" pageNumber="57">3</emphasis>
a–3a 13,
<emphasis id="B94ACC3C743CFFA9496AFB7EFBBEFB3A" box="[1091,1105,1221,1245]" italics="true" pageId="1" pageNumber="57">3</emphasis>
b–3b 37,
<emphasis id="B94ACC3C743CFFA94990FB7EFB28FB3A" box="[1209,1223,1221,1245]" italics="true" pageId="1" pageNumber="57">3</emphasis>
c–3c 82,
<emphasis id="B94ACC3C743CFFA94805FB7EFAD5FB3A" box="[1324,1338,1221,1245]" italics="true" pageId="1" pageNumber="57">4</emphasis>
a–4a 27,
<emphasis id="B94ACC3C743CFFA94DBEFB52FF49FAE6" box="[151,166,1257,1281]" italics="true" pageId="1" pageNumber="57">4</emphasis>
b–4b 63,
<emphasis id="B94ACC3C743CFFA94C26FB52FEF2FAE6" box="[271,285,1257,1281]" italics="true" pageId="1" pageNumber="57">4</emphasis>
c–4c 77. Measurements of
<typeStatus id="5485AE8C743CFFA94F66FB52FD5AFAE6" box="[591,693,1257,1281]" pageId="1" pageNumber="57" type="holotype">holotype</typeStatus>
:
<emphasis id="B94ACC3C743CFFA94FE9FB52FD32FAE6" box="[704,733,1257,1281]" italics="true" pageId="1" pageNumber="57">1a</emphasis>
37,
<emphasis id="B94ACC3C743CFFA94E26FB52FCF3FAE6" box="[783,796,1257,1281]" italics="true" pageId="1" pageNumber="57">1</emphasis>
b 29,
<emphasis id="B94ACC3C743CFFA94E74FB52FC84FAE6" box="[861,875,1257,1281]" italics="true" pageId="1" pageNumber="57">2</emphasis>
a 26,
<emphasis id="B94ACC3C743CFFA94E85FB52FC55FAE6" box="[940,954,1257,1281]" italics="true" pageId="1" pageNumber="57">2</emphasis>
b 43,
<emphasis id="B94ACC3C743CFFA94ED3FB52FBF9FAE6" box="[1018,1046,1257,1281]" italics="true" pageId="1" pageNumber="57">3a</emphasis>
59,
<emphasis id="B94ACC3C743CFFA94960FB52FBB8FAE6" box="[1097,1111,1257,1281]" italics="true" pageId="1" pageNumber="57">3</emphasis>
b 53,
<emphasis id="B94ACC3C743CFFA949BEFB52FB4AFAE6" box="[1175,1189,1257,1281]" italics="true" pageId="1" pageNumber="57">3</emphasis>
c 45,
<emphasis id="B94ACC3C743CFFA949CDFB52FAEEFAE6" box="[1252,1281,1257,1281]" italics="true" pageId="1" pageNumber="57">4a</emphasis>
61,
<emphasis id="B94ACC3C743CFFA9481AFB52FAADFAE6" box="[1331,1346,1257,1281]" italics="true" pageId="1" pageNumber="57">4</emphasis>
b 64,
<emphasis id="B94ACC3C743CFFA948A8FB52FA60FAE6" box="[1409,1423,1257,1281]" italics="true" pageId="1" pageNumber="57">4</emphasis>
c 56;
<emphasis id="B94ACC3C743CFFA94DEBFAB6FEE7FAC2" box="[194,264,1293,1317]" italics="true" pageId="1" pageNumber="57">1a–1a</emphasis>
18,
<emphasis id="B94ACC3C743CFFA94C10FAB6FE90FAC2" box="[313,383,1293,1317]" italics="true" pageId="1" pageNumber="57">1b–1b</emphasis>
48,
<emphasis id="B94ACC3C743CFFA94C86FAB6FE53FAC2" box="[431,444,1293,1317]" italics="true" pageId="1" pageNumber="57">2</emphasis>
a–2a 37,
<emphasis id="B94ACC3C743CFFA94F0FFAB6FDDBFAC2" box="[550,564,1293,1317]" italics="true" pageId="1" pageNumber="57">2</emphasis>
b–2b 35,
<emphasis id="B94ACC3C743CFFA94FB4FAB6FD44FAC2" box="[669,683,1293,1317]" italics="true" pageId="1" pageNumber="57">3</emphasis>
a–3a 11,
<emphasis id="B94ACC3C743CFFA94E3BFAB6FCCFFAC2" box="[786,800,1293,1317]" italics="true" pageId="1" pageNumber="57">3</emphasis>
b–3b 31,
<emphasis id="B94ACC3C743CFFA94EA0FAB6FC78FAC2" box="[905,919,1293,1317]" italics="true" pageId="1" pageNumber="57">3</emphasis>
c–3c 70,
<emphasis id="B94ACC3C743CFFA94ED4FAB6FBE4FAC2" box="[1021,1035,1293,1317]" italics="true" pageId="1" pageNumber="57">4</emphasis>
a–4a 22,
<emphasis id="B94ACC3C743CFFA9495DFAB6FB6DFAC2" box="[1140,1154,1293,1317]" italics="true" pageId="1" pageNumber="57">4</emphasis>
b–4b 54,
<emphasis id="B94ACC3C743CFFA949C2FAB6FB15FAC2" box="[1259,1274,1293,1317]" italics="true" pageId="1" pageNumber="57">4</emphasis>
c–4c 63.
</paragraph>
<paragraph id="8B81102E743CFFA94DEEFA8AFC36FAAE" blockId="1.[151,1437,717,2037]" box="[199,985,1329,1353]" pageId="1" pageNumber="57">
Trichobothrium thin-stemmed, club-shaped, with fine barbs. (
<figureCitation id="13050CAB743CFFA94E53FA8AFC51FAAE" box="[890,958,1329,1353]" captionStart="FIGURE 4" captionStartId="5.[151,250,1956,1979]" captionTargetBox="[190,1426,329,1913]" captionTargetId="figure@5.[160,1445,303,1926]" captionTargetPageId="5" captionText="FIGURE 4. Heterodispus cordidiscus n. sp. female holotype. a) leg I; b) leg II; c) leg III; d) leg IV; e) lateral part of prodorsum." httpUri="https://zenodo.org/record/282653/files/figure.png" pageId="1" pageNumber="57">Fig. 4</figureCitation>
e).
</paragraph>
<paragraph id="8B81102E743CFFA94DEEFAEEFE91FA52" blockId="1.[151,1437,717,2037]" pageId="1" pageNumber="57">
<emphasis id="B94ACC3C743CFFA94DEEFAEEFEB9FA8A" box="[199,342,1365,1389]" italics="true" pageId="1" pageNumber="57">Gnathosoma</emphasis>
(
<figureCitation id="13050CAB743CFFA94C4FFAEEFE46FA8A" box="[358,425,1365,1389]" captionStart="FIGURE 5" captionStartId="6.[151,250,590,613]" captionTargetBox="[161,1421,159,569]" captionTargetId="figure@6.[153,1435,153,575]" captionTargetPageId="6" captionText="FIGURE 5. Heterodispus cordidiscus n. sp. a) gnathosoma of male larva, dorsal view; b) gnathosoma of female, ventral view; c)" httpUri="https://zenodo.org/record/282654/files/figure.png" pageId="1" pageNumber="57">Fig. 5</figureCitation>
b) roundish, average width 24 (n = 20), average length 22 (n = 13); dorsal setae
<emphasis id="B94ACC3C743CFFA94815FAEEFA8BFA8A" box="[1340,1380,1365,1389]" italics="true" pageId="1" pageNumber="57">ch1</emphasis>
,
<emphasis id="B94ACC3C743CFFA9485DFAEEFA73FA8A" box="[1396,1436,1365,1389]" italics="true" pageId="1" pageNumber="57">ch2</emphasis>
and
<emphasis id="B94ACC3C743CFFA94DEEFAC1FF0CFA76" box="[199,227,1402,1425]" italics="true" pageId="1" pageNumber="57">pp</emphasis>
thin, smooth; ventral setae
<emphasis id="B94ACC3C743CFFA94F34FAC1FDD9FA76" box="[541,566,1402,1425]" italics="true" pageId="1" pageNumber="57">su</emphasis>
thin, smooth. Each palp with setae
<emphasis id="B94ACC3C743CFFA94EE5FAC2FC35FA76" box="[972,986,1401,1425]" italics="true" pageId="1" pageNumber="57">d</emphasis>
Ge and
<emphasis id="B94ACC3C743CFFA9491AFAC2FBAEFA76" box="[1075,1089,1401,1425]" italics="true" pageId="1" pageNumber="57">d</emphasis>
Fe, 1 solenidion and accessory setigenous structure.
</paragraph>
<paragraph id="8B81102E743CFFA94DEEFA79FC54F96A" blockId="1.[151,1437,717,2037]" pageId="1" pageNumber="57">
<emphasis id="B94ACC3C743CFFA94DEEFA79FF15FA3E" box="[199,250,1474,1497]" italics="true" pageId="1" pageNumber="57">Legs</emphasis>
. Leg I (
<figureCitation id="13050CAB743CFFA94C7AFA7AFE7BFA3E" box="[339,404,1473,1497]" captionStart="FIGURE 4" captionStartId="5.[151,250,1956,1979]" captionTargetBox="[190,1426,329,1913]" captionTargetId="figure@5.[160,1445,303,1926]" captionTargetPageId="5" captionText="FIGURE 4. Heterodispus cordidiscus n. sp. female holotype. a) leg I; b) leg II; c) leg III; d) leg IV; e) lateral part of prodorsum." httpUri="https://zenodo.org/record/282653/files/figure.png" pageId="1" pageNumber="57">Fig. 4</figureCitation>
a) Formula of setae: Tr1-Fe3-Ge4-TiTa16(4). Solenidia ω
<emphasis id="B94ACC3C743CFFA94922FA7AFBF6FA3E" box="[1035,1049,1473,1497]" italics="true" pageId="1" pageNumber="57">1</emphasis>
and φ
<emphasis id="B94ACC3C743CFFA9494DFA7AFB9DFA3E" box="[1124,1138,1473,1497]" italics="true" pageId="1" pageNumber="57">1</emphasis>
finger-shaped, ω
<emphasis id="B94ACC3C743CFFA9481FFA7AFAABFA3E" box="[1334,1348,1473,1497]" italics="true" pageId="1" pageNumber="57">2</emphasis>
and φ
<emphasis id="B94ACC3C743CFFA948A7FA7AFA73FA3E" box="[1422,1436,1473,1497]" italics="true" pageId="1" pageNumber="57">2</emphasis>
small and thin. ω
<emphasis id="B94ACC3C743CFFA94C72FA5EFE86FA1A" box="[347,361,1509,1533]" italics="true" pageId="1" pageNumber="57">1</emphasis>
&gt; φ
<emphasis id="B94ACC3C743CFFA94CB0FA5EFE48FA1A" box="[409,423,1509,1533]" italics="true" pageId="1" pageNumber="57">1</emphasis>
&gt; φ
<emphasis id="B94ACC3C743CFFA94CFEFA5EFE0AFA1A" box="[471,485,1509,1533]" italics="true" pageId="1" pageNumber="57">2</emphasis>
≥ ω
<emphasis id="B94ACC3C743CFFA94F31FA5EFDC9FA1A" box="[536,550,1509,1533]" italics="true" pageId="1" pageNumber="57">2</emphasis>
. TiTa with well developed claw. Leg II (
<figureCitation id="13050CAB743CFFA94ECAFA5EFBCBFA1A" box="[995,1060,1509,1533]" captionStart="FIGURE 4" captionStartId="5.[151,250,1956,1979]" captionTargetBox="[190,1426,329,1913]" captionTargetId="figure@5.[160,1445,303,1926]" captionTargetPageId="5" captionText="FIGURE 4. Heterodispus cordidiscus n. sp. female holotype. a) leg I; b) leg II; c) leg III; d) leg IV; e) lateral part of prodorsum." httpUri="https://zenodo.org/record/282653/files/figure.png" pageId="1" pageNumber="57">Fig. 4</figureCitation>
b) Formula of setae: Tr1-Fe3-Ge3- Ti4(1)-Ta6(1). Solenidion ω&gt; φ, both finger-shaped, tarsal setae
<emphasis id="B94ACC3C743CFFA94E76F9B1FC94F9C6" box="[863,891,1546,1569]" italics="true" pageId="1" pageNumber="57">tc’</emphasis>
and
<emphasis id="B94ACC3C743CFFA94E9AF9B2FC39F9C6" box="[947,982,1545,1569]" italics="true" pageId="1" pageNumber="57">pl’’</emphasis>
and tibial seta
<emphasis id="B94ACC3C743CFFA94957F9B1FB76F9C6" box="[1150,1177,1546,1569]" italics="true" pageId="1" pageNumber="57">v’’</emphasis>
spine-like. Ta with two small claws and empodium. Leg III (
<figureCitation id="13050CAB743CFFA94F1BF996FD9AF9A2" box="[562,629,1581,1605]" captionStart="FIGURE 4" captionStartId="5.[151,250,1956,1979]" captionTargetBox="[190,1426,329,1913]" captionTargetId="figure@5.[160,1445,303,1926]" captionTargetPageId="5" captionText="FIGURE 4. Heterodispus cordidiscus n. sp. female holotype. a) leg I; b) leg II; c) leg III; d) leg IV; e) lateral part of prodorsum." httpUri="https://zenodo.org/record/282653/files/figure.png" pageId="1" pageNumber="57">Fig. 4</figureCitation>
c) Formula of setae: Tr1-Fe2-Ge2-Ti4(1)-Ta6. Solenidion φ small, fingershaped, tarsal setae
<emphasis id="B94ACC3C743CFFA94C40F9E9FE6AF98E" box="[361,389,1618,1641]" italics="true" pageId="1" pageNumber="57">tc’</emphasis>
,
<emphasis id="B94ACC3C743CFFA94CB8F9EAFE5BF98E" box="[401,436,1617,1641]" italics="true" pageId="1" pageNumber="57">pl’’</emphasis>
,
<emphasis id="B94ACC3C743CFFA94CE8F9E9FE0CF98E" box="[449,483,1618,1641]" italics="true" pageId="1" pageNumber="57">pv’</emphasis>
and
<emphasis id="B94ACC3C743CFFA94F3FF9E9FDD1F98E" box="[534,574,1618,1641]" italics="true" pageId="1" pageNumber="57">pv’’</emphasis>
spine-like. Ta with two small claws and empodium. Leg IV (
<figureCitation id="13050CAB743CFFA949E0F9EAFAE7F98E" box="[1225,1288,1617,1641]" captionStart="FIGURE 4" captionStartId="5.[151,250,1956,1979]" captionTargetBox="[190,1426,329,1913]" captionTargetId="figure@5.[160,1445,303,1926]" captionTargetPageId="5" captionText="FIGURE 4. Heterodispus cordidiscus n. sp. female holotype. a) leg I; b) leg II; c) leg III; d) leg IV; e) lateral part of prodorsum." httpUri="https://zenodo.org/record/282653/files/figure.png" pageId="1" pageNumber="57">Fig. 4</figureCitation>
d) Formula of setae: Tr1-Fe2-Ge1-Ti3-Ta4. Tibial seta
<emphasis id="B94ACC3C743CFFA94F6CF9CEFDBAF96A" box="[581,597,1653,1677]" italics="true" pageId="1" pageNumber="57">l’</emphasis>
spine-like. PrTa with empodium.
</paragraph>
<paragraph id="8B81102E743CFFA94DEEF921FD38F956" blockId="1.[151,1437,717,2037]" box="[199,727,1689,1714]" pageId="1" pageNumber="57">
<emphasis id="B94ACC3C743CFFA94DEEF921FEEBF955" bold="true" box="[199,260,1690,1714]" pageId="1" pageNumber="57">Male</emphasis>
. Idiosoma length: 158 (134–201), n = 11.
</paragraph>
<paragraph id="8B81102E743CFFA94DEEF905FCCCF86E" blockId="1.[151,1437,717,2037]" pageId="1" pageNumber="57">
<emphasis id="B94ACC3C743CFFA94DEEF905FECEF932" box="[199,289,1726,1749]" italics="true" pageId="1" pageNumber="57">Dorsum</emphasis>
(
<figureCitation id="13050CAB743CFFA94C19F906FE9AF932" box="[304,373,1725,1749]" captionStart="FIGURE 6" captionStartId="6.[151,250,2009,2032]" captionTargetBox="[185,1384,622,1986]" captionTargetId="figure@6.[185,1396,622,2000]" captionTargetPageId="6" captionText="FIGURE 6. Heterodispus cordidiscus n. sp. male, dorsal view. ae = aedeagus, dej = ejaculatory duct, sv = seminar vesicle." httpUri="https://zenodo.org/record/282655/files/figure.png" pageId="1" pageNumber="57">Fig. 6</figureCitation>
). Idiosoma with four finely stippled sclerotized plates. Most posterior plate forms dorsal cover for genital capsule. Setae
<date id="FF8036EE743CFFA94C90F95AFE3CF91E" box="[441,467,1761,1785]" pageId="1" pageNumber="57">
<emphasis id="B94ACC3C743CFFA94C90F95AFE3CF91E" box="[441,467,1761,1785]" italics="true" pageId="1" pageNumber="57">v2</emphasis>
</date>
,
<emphasis id="B94ACC3C743CFFA94CC8F95AFDE9F91E" box="[481,518,1761,1785]" italics="true" pageId="1" pageNumber="57">sc1</emphasis>
,
<emphasis id="B94ACC3C743CFFA94F3CF95AFDD4F91E" box="[533,571,1761,1785]" italics="true" pageId="1" pageNumber="57">sc2</emphasis>
,
<emphasis id="B94ACC3C743CFFA94F60F95AFD8DF91E" box="[585,610,1761,1785]" italics="true" pageId="1" pageNumber="57">c1</emphasis>
,
<emphasis id="B94ACC3C743CFFA94F5BF95AFD63F91E" box="[626,652,1761,1785]" italics="true" pageId="1" pageNumber="57">c2</emphasis>
,
<emphasis id="B94ACC3C743CFFA94FB3F95AFD47F91E" box="[666,680,1761,1785]" italics="true" pageId="1" pageNumber="57">d</emphasis>
,
<emphasis id="B94ACC3C743CFFA94F9EF959FD2CF91E" box="[695,707,1762,1785]" italics="true" pageId="1" pageNumber="57">e</emphasis>
,
<emphasis id="B94ACC3C743CFFA94FFBF95AFD35F91E" box="[722,730,1761,1785]" italics="true" pageId="1" pageNumber="57">f</emphasis>
barbed,
<emphasis id="B94ACC3C743CFFA94E12F95AFCB8F91E" box="[827,855,1761,1785]" italics="true" pageId="1" pageNumber="57">h1</emphasis>
smooth,
<emphasis id="B94ACC3C743CFFA94E96F95AFC33F91E" box="[959,988,1761,1785]" italics="true" pageId="1" pageNumber="57">h2</emphasis>
minute. All setae except
<emphasis id="B94ACC3C743CFFA949D3F95AFAE7F91E" box="[1274,1288,1761,1785]" italics="true" pageId="1" pageNumber="57">d</emphasis>
blunt-ended,
<emphasis id="B94ACC3C743CFFA94DBEF8BEFF4AF8FA" box="[151,165,1797,1821]" italics="true" pageId="1" pageNumber="57">d</emphasis>
pointed. Average length of dorsal setae:
<date id="FF8036EE743CFFA94F45F8BEFD46F8FA" box="[620,681,1797,1821]" pageId="1" pageNumber="57" value="2010-05-02">
<emphasis id="B94ACC3C743CFFA94F45F8BEFD68F8FA" box="[620,647,1797,1821]" italics="true" pageId="1" pageNumber="57">v2</emphasis>
10
</date>
(n = 11),
<emphasis id="B94ACC3C743CFFA94E3FF8BEFCD3F8FA" box="[790,828,1797,1821]" italics="true" pageId="1" pageNumber="57">sc1</emphasis>
19 (n = 11),
<emphasis id="B94ACC3C743CFFA94EE5F8BEFC1EF8FA" box="[972,1009,1797,1821]" italics="true" pageId="1" pageNumber="57">sc2</emphasis>
38 (n = 11),
<emphasis id="B94ACC3C743CFFA949A9F8BEFB74F8FA" box="[1152,1179,1797,1821]" italics="true" pageId="1" pageNumber="57">c1</emphasis>
20 (n = 11),
<emphasis id="B94ACC3C743CFFA94803F8BEFAABF8FA" box="[1322,1348,1797,1821]" italics="true" pageId="1" pageNumber="57">c2</emphasis>
35 (n = 12),
<emphasis id="B94ACC3C743CFFA94DE3F892FF37F8A6" box="[202,216,1833,1857]" italics="true" pageId="1" pageNumber="57">d</emphasis>
34 (n = 11),
<emphasis id="B94ACC3C743CFFA94C43F891FE99F8A6" box="[362,374,1834,1857]" italics="true" pageId="1" pageNumber="57">e</emphasis>
18 (n = 12),
<emphasis id="B94ACC3C743CFFA94F20F892FDFEF8A6" box="[521,529,1833,1857]" italics="true" pageId="1" pageNumber="57">f</emphasis>
26 (n = 12),
<emphasis id="B94ACC3C743CFFA94F8AF892FD50F8A6" box="[675,703,1833,1857]" italics="true" pageId="1" pageNumber="57">h1</emphasis>
9 (n = 12). Average distance between dorsal setae:
<date id="FF8036EE743CFFA94828F892FA88F8A6" box="[1281,1383,1833,1857]" pageId="1" pageNumber="57" value="1917-05-02" valueMax="1917-05-02" valueMin="1917-05-02">
<emphasis id="B94ACC3C743CFFA94828F892FAABF8A6" box="[1281,1348,1833,1857]" italics="true" pageId="1" pageNumber="57">v2–v2</emphasis>
17
</date>
(n = 11),
<emphasis id="B94ACC3C743CFFA94DE3F8F6FECCF882" box="[202,291,1869,1893]" italics="true" pageId="1" pageNumber="57">sc1–sc1</emphasis>
13 (n = 11),
<emphasis id="B94ACC3C743CFFA94C91F8F6FDFDF882" box="[440,530,1869,1893]" italics="true" pageId="1" pageNumber="57">sc2–sc2</emphasis>
27 (n = 11),
<emphasis id="B94ACC3C743CFFA94F8EF8F6FD05F882" box="[679,746,1869,1893]" italics="true" pageId="1" pageNumber="57">c1–c1</emphasis>
50 (n = 11),
<emphasis id="B94ACC3C743CFFA94EA9F8F6FC2CF882" box="[896,963,1869,1893]" italics="true" pageId="1" pageNumber="57">c2–c2</emphasis>
86 (n = 12),
<emphasis id="B94ACC3C743CFFA94970F8F6FB6CF882" box="[1113,1155,1869,1893]" italics="true" pageId="1" pageNumber="57">d–d</emphasis>
69 (n = 12),
<emphasis id="B94ACC3C743CFFA94833F8F5FAAFF882" box="[1306,1344,1870,1893]" italics="true" pageId="1" pageNumber="57">e–e</emphasis>
50 (n = 12),
<emphasis id="B94ACC3C743CFFA94DE3F8CAFF07F86E" box="[202,232,1905,1929]" italics="true" pageId="1" pageNumber="57">f–f</emphasis>
31 (n = 12),
<emphasis id="B94ACC3C743CFFA94C53F8CAFE2FF86E" box="[378,448,1905,1929]" italics="true" pageId="1" pageNumber="57">h1–h1</emphasis>
32 (n = 12),
<emphasis id="B94ACC3C743CFFA94F7AF8CAFD76F86E" box="[595,665,1905,1929]" italics="true" pageId="1" pageNumber="57">h2–h2</emphasis>
26 (n = 12).
</paragraph>
<paragraph id="8B81102E743CFFA94DEEF82DFEA7F812" blockId="1.[151,1437,717,2037]" pageId="1" pageNumber="57">
<emphasis id="B94ACC3C743CFFA94DEEF82DFEE3F84A" box="[199,268,1942,1965]" italics="true" pageId="1" pageNumber="57">Venter</emphasis>
(
<figureCitation id="13050CAB743CFFA94C34F82EFE8EF84A" box="[285,353,1941,1965]" captionStart="FIGURE 7" captionStartId="7.[151,250,1697,1720]" captionTargetBox="[275,1305,196,1663]" captionTargetId="figure@7.[236,1342,163,1681]" captionTargetPageId="7" captionText="FIGURE 7. Heterodispus cordidiscus n. sp. male, ventral view. gn = gnathosoma." httpUri="https://zenodo.org/record/282656/files/figure.png" pageId="1" pageNumber="57">Fig. 7</figureCitation>
). Sternal plates finely stippled. ApI, apII clearly sclerotized. Presternal ap continuous from apI to sejugal apodeme. ApIII incomplete, apIV complete. Poststernal ap incomplete, present distally from point where apIV branch of.
</paragraph>
<caption id="DF4140A6743FFFAA4DBEF8BBFC68F8F0" box="[151,903,1792,1815]" httpUri="https://zenodo.org/record/282650/files/figure.png" pageId="2" pageNumber="58" targetBox="[158,1400,186,1791]" targetPageId="2">
<paragraph id="8B81102E743FFFAA4DBEF8BBFC68F8F0" blockId="2.[151,903,1792,1815]" box="[151,903,1792,1815]" pageId="2" pageNumber="58">
<emphasis id="B94ACC3C743FFFAA4DBEF8BBFEFCF8F1" bold="true" box="[151,275,1792,1815]" pageId="2" pageNumber="58">FIGURE 1.</emphasis>
<taxonomicName id="4C3E6BAD743FFFAA4C30F8BAFDF6F8F0" box="[281,537,1793,1815]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="cordidiscus" status="sp. nov.">
<emphasis id="B94ACC3C743FFFAA4C30F8BAFDF6F8F0" box="[281,537,1793,1815]" italics="true" pageId="2" pageNumber="58">Heterodispus cordidiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2797147743FFFAA4F36F8B9FDB9F8F0" box="[543,598,1794,1815]" pageId="2" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
female holotype, dorsal view.
</paragraph>
</caption>
<paragraph id="8B81102E743FFFAA4DEEF8F8FDD3F80C" blockId="2.[151,1436,1859,2028]" pageId="2" pageNumber="58">
Setae
<emphasis id="B94ACC3C743FFFAA4C23F8FFFECAF8BB" box="[266,293,1860,1884]" italics="true" pageId="2" pageNumber="58">1a</emphasis>
,
<emphasis id="B94ACC3C743FFFAA4C1DF8FFFEA0F8BB" box="[308,335,1860,1884]" italics="true" pageId="2" pageNumber="58">1b</emphasis>
,
<emphasis id="B94ACC3C743FFFAA4C74F8FFFE96F8BB" box="[349,377,1860,1884]" italics="true" pageId="2" pageNumber="58">2a</emphasis>
,
<emphasis id="B94ACC3C743FFFAA4CAEF8FFFE4CF8BB" box="[391,419,1860,1884]" italics="true" pageId="2" pageNumber="58">2b</emphasis>
sparsely barbed, setae
<emphasis id="B94ACC3C743FFFAA4F8DF8FFFD2FF8BB" box="[676,704,1860,1884]" italics="true" pageId="2" pageNumber="58">3a</emphasis>
,
<emphasis id="B94ACC3C743FFFAA4FE7F8FFFD05F8BB" box="[718,746,1860,1884]" italics="true" pageId="2" pageNumber="58">3b</emphasis>
,
<emphasis id="B94ACC3C743FFFAA4FD1F8FFFCFDF8BB" box="[760,786,1860,1884]" italics="true" pageId="2" pageNumber="58">3c</emphasis>
,
<emphasis id="B94ACC3C743FFFAA4E09F8FFFCD4F8BB" box="[800,827,1860,1884]" italics="true" pageId="2" pageNumber="58">4a</emphasis>
,
<emphasis id="B94ACC3C743FFFAA4E63F8FFFC8AF8BB" box="[842,869,1860,1884]" italics="true" pageId="2" pageNumber="58">4b</emphasis>
,
<emphasis id="B94ACC3C743FFFAA4E5AF8FFFC61F8BB" box="[883,910,1860,1884]" italics="true" pageId="2" pageNumber="58">4c</emphasis>
smooth. Average length of ventral setae:
<emphasis id="B94ACC3C743FFFAA4874F8FFFA95F8BB" box="[1373,1402,1860,1884]" italics="true" pageId="2" pageNumber="58">1a</emphasis>
12 (n = 10),
<emphasis id="B94ACC3C743FFFAA4DD6F8DCFEF3F898" box="[255,284,1895,1919]" italics="true" pageId="2" pageNumber="58">1b</emphasis>
15 (n = 10),
<emphasis id="B94ACC3C743FFFAA4C84F8DCFE26F898" box="[429,457,1895,1919]" italics="true" pageId="2" pageNumber="58">2a</emphasis>
15 (n = 9),
<emphasis id="B94ACC3C743FFFAA4F64F8DCFD86F898" box="[589,617,1895,1919]" italics="true" pageId="2" pageNumber="58">2b</emphasis>
15 (n = 12),
<emphasis id="B94ACC3C743FFFAA4FD5F8DCFCF8F898" box="[764,791,1895,1919]" italics="true" pageId="2" pageNumber="58">3a</emphasis>
12 (n = 9),
<emphasis id="B94ACC3C743FFFAA4EB5F8DCFC57F898" box="[924,952,1895,1919]" italics="true" pageId="2" pageNumber="58">3b</emphasis>
14 (n = 11),
<emphasis id="B94ACC3C743FFFAA4960F8DCFB8CF898" box="[1097,1123,1895,1919]" italics="true" pageId="2" pageNumber="58">3c</emphasis>
14 (n = 11),
<emphasis id="B94ACC3C743FFFAA49DDF8DCFAFEF898" box="[1268,1297,1895,1919]" italics="true" pageId="2" pageNumber="58">4a</emphasis>
12 (n = 10),
<emphasis id="B94ACC3C743FFFAA4DBEF837FF5CF843" box="[151,179,1932,1956]" italics="true" pageId="2" pageNumber="58">4b</emphasis>
15 (n = 12),
<emphasis id="B94ACC3C743FFFAA4C78F837FE83F843" box="[337,364,1932,1956]" italics="true" pageId="2" pageNumber="58">4c</emphasis>
12 (n = 12). Average distance between ventral setae:
<emphasis id="B94ACC3C743FFFAA4EF5F837FBCCF843" box="[988,1059,1932,1956]" italics="true" pageId="2" pageNumber="58">1a–1a</emphasis>
15 (n = 10),
<emphasis id="B94ACC3C743FFFAA49E8F837FAE8F843" box="[1217,1287,1932,1956]" italics="true" pageId="2" pageNumber="58">1b–1b</emphasis>
47 (n = 10),
<emphasis id="B94ACC3C743FFFAA4DBEF814FF33F820" box="[151,220,1967,1991]" italics="true" pageId="2" pageNumber="58">2a–2a</emphasis>
24 (n = 11),
<emphasis id="B94ACC3C743FFFAA4C47F814FE5BF820" box="[366,436,1967,1991]" italics="true" pageId="2" pageNumber="58">2b–2b</emphasis>
42 (n = 12),
<emphasis id="B94ACC3C743FFFAA4F6FF814FD63F820" box="[582,652,1967,1991]" italics="true" pageId="2" pageNumber="58">3a–3a</emphasis>
22 (n = 11),
<emphasis id="B94ACC3C743FFFAA4E37F814FC8CF820" box="[798,867,1967,1991]" italics="true" pageId="2" pageNumber="58">3b–3b</emphasis>
41 (n = 11),
<emphasis id="B94ACC3C743FFFAA4EDCF814FBD7F820" box="[1013,1080,1967,1991]" italics="true" pageId="2" pageNumber="58">3c–3c</emphasis>
68 (n = 12),
<emphasis id="B94ACC3C743FFFAA49E2F814FAFEF820" box="[1227,1297,1967,1991]" italics="true" pageId="2" pageNumber="58">4a–4a</emphasis>
17 (n = 12),
<emphasis id="B94ACC3C743FFFAA4DBEF86FFF32F80B" box="[151,221,2004,2028]" italics="true" pageId="2" pageNumber="58">4b–4b</emphasis>
11 (n = 12),
<emphasis id="B94ACC3C743FFFAA4C47F86FFE5EF80B" box="[366,433,2004,2028]" italics="true" pageId="2" pageNumber="58">4c–4c</emphasis>
55 (n = 12).
</paragraph>
<caption id="DF4140A6743EFFAB4DBEF960FC60F916" box="[151,911,1755,1778]" httpUri="https://zenodo.org/record/282651/files/figure.png" pageId="3" pageNumber="59" targetBox="[184,1334,184,1754]" targetPageId="3">
<paragraph id="8B81102E743EFFAB4DBEF960FC60F916" blockId="3.[151,911,1755,1778]" box="[151,911,1755,1778]" pageId="3" pageNumber="59">
<emphasis id="B94ACC3C743EFFAB4DBEF960FEFCF916" bold="true" box="[151,275,1755,1778]" pageId="3" pageNumber="59">FIGURE 2.</emphasis>
<taxonomicName id="4C3E6BAD743EFFAB4C30F967FDF6F915" box="[281,537,1756,1778]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="cordidiscus" status="sp. nov.">
<emphasis id="B94ACC3C743EFFAB4C30F967FDF6F915" box="[281,537,1756,1778]" italics="true" pageId="3" pageNumber="59">Heterodispus cordidiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2797147743EFFAB4F36F966FDB9F915" box="[543,598,1757,1778]" pageId="3" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
female holotype, ventral view.
</paragraph>
</caption>
<paragraph id="8B81102E743EFFAB4DEEF8A5FD07F80D" blockId="3.[151,1436,1822,2026]" pageId="3" pageNumber="59">
<emphasis id="B94ACC3C743EFFAB4DEEF8A5FEB9F8D1" box="[199,342,1822,1846]" italics="true" pageId="3" pageNumber="59">Gnathosoma</emphasis>
(
<figureCitation id="13050CAB743EFFAB4C4FF8A5FE46F8D1" box="[358,425,1822,1846]" captionStart="FIGURE 5" captionStartId="6.[151,250,590,613]" captionTargetBox="[161,1421,159,569]" captionTargetId="figure@6.[153,1435,153,575]" captionTargetPageId="6" captionText="FIGURE 5. Heterodispus cordidiscus n. sp. a) gnathosoma of male larva, dorsal view; b) gnathosoma of female, ventral view; c)" httpUri="https://zenodo.org/record/282654/files/figure.png" pageId="3" pageNumber="59">Fig. 5</figureCitation>
c) extended, tube-shaped, average length 8 (n= 10), no functional mouthparts. One pair of finger-shaped solenidia, two pairs of dorsal setae
<emphasis id="B94ACC3C743EFFAB4FE8F8F8FD05F8BC" box="[705,746,1859,1883]" italics="true" pageId="3" pageNumber="59">ch1</emphasis>
and
<emphasis id="B94ACC3C743EFFAB4E09F8F8FCA7F8BC" box="[800,840,1859,1883]" italics="true" pageId="3" pageNumber="59">ch2</emphasis>
thin, smooth, ventral pair of setae
<emphasis id="B94ACC3C743EFFAB49E7F8FFFB08F8BC" box="[1230,1255,1860,1883]" italics="true" pageId="3" pageNumber="59">su</emphasis>
longer, barbed. Genital apparatus (
<figureCitation id="13050CAB743EFFAB4C46F8DDFE2FF899" box="[367,448,1894,1918]" captionStart="FIGURE 6" captionStartId="6.[151,250,2009,2032]" captionTargetBox="[185,1384,622,1986]" captionTargetId="figure@6.[185,1396,622,2000]" captionTargetPageId="6" captionText="FIGURE 6. Heterodispus cordidiscus n. sp. male, dorsal view. ae = aedeagus, dej = ejaculatory duct, sv = seminar vesicle." httpUri="https://zenodo.org/record/282655/files/figure.png" pageId="3" pageNumber="59">Figs. 6</figureCitation>
,
<figureCitation id="13050CAB743EFFAB4CE7F8DDFE33F899" box="[462,476,1894,1918]" captionStart="FIGURE 8" captionStartId="8.[151,250,1579,1602]" captionTargetBox="[455,1134,932,1555]" captionTargetId="figure@8.[450,1142,926,1562]" captionTargetPageId="8" captionText="FIGURE 8. Heterodispus cordidiscus n. sp. male, lateral view of the genital apparatus. ae = aedeagus, dej = ejaculatory duct, lo = internal loop of the aedeagus, sv = seminar vesicle." httpUri="https://zenodo.org/record/282657/files/figure.png" pageId="3" pageNumber="59">8</figureCitation>
) Length of genital capsule (tergite H and Ps) 43 (42–51), n = 10. Posterior edge of genital capsule narrowed. Surface with round, sucker-like structures with a diameter of about 2 and four ridges (two on each side) forming a heart-shaped structure. Aedeagus with diagonal stripes, internally forming a loop and continuing as ejaculatory duct into a seminal vesicle.
</paragraph>
<caption id="DF4140A67439FFAC4DBEFB1FFC93FB3E" httpUri="https://zenodo.org/record/282652/files/figure.png" pageId="4" pageNumber="60" targetBox="[510,1088,186,1160]" targetPageId="4">
<paragraph id="8B81102E7439FFAC4DBEFB1FFC93FB3E" blockId="4.[151,1436,1188,1242]" pageId="4" pageNumber="60">
<emphasis id="B94ACC3C7439FFAC4DBEFB1FFEFDFB5D" bold="true" box="[151,274,1188,1211]" pageId="4" pageNumber="60">FIGURE 3.</emphasis>
<taxonomicName id="4C3E6BAD7439FFAC4C31FB1EFDF7FB5C" box="[280,536,1189,1211]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="cordidiscus" status="sp. nov.">
<emphasis id="B94ACC3C7439FFAC4C31FB1EFDF7FB5C" box="[280,536,1189,1211]" italics="true" pageId="4" pageNumber="60">Heterodispus cordidiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27971477439FFAC4F34FB1DFDBCFB5C" box="[541,595,1190,1211]" pageId="4" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
female. Examples of the variability in the positions of setae
<emphasis id="B94ACC3C7439FFAC499AFB1EFB23FB5C" box="[1203,1228,1189,1211]" italics="true" pageId="4" pageNumber="60">4a</emphasis>
and
<emphasis id="B94ACC3C7439FFAC49D5FB1EFAF9FB5C" box="[1276,1302,1189,1211]" italics="true" pageId="4" pageNumber="60">4b</emphasis>
in relation to each other; insertions of the respective specimens are connected by lines.
</paragraph>
</caption>
<paragraph id="8B81102E7439FFAC4DEEFA9AFC06F9E8" blockId="4.[151,1436,1311,2020]" pageId="4" pageNumber="60">
<emphasis id="B94ACC3C7439FFAC4DEEFA9AFF15FADF" box="[199,250,1313,1336]" italics="true" pageId="4" pageNumber="60">Legs</emphasis>
. Leg I (
<figureCitation id="13050CAB7439FFAC4C7FFAA4FE76FAD0" box="[342,409,1311,1335]" captionStart="FIGURE 9" captionStartId="9.[151,250,1861,1884]" captionTargetBox="[191,1397,209,1798]" captionTargetId="figure@9.[151,1436,200,1823]" captionTargetPageId="9" captionText="FIGURE 9. Heterodispus cordidiscus n. sp. male. a) leg I; b) leg II; c) leg III; d) leg IV." httpUri="https://zenodo.org/record/282658/files/figure.png" pageId="4" pageNumber="60">Fig. 9</figureCitation>
a). Setal formula: Tr1-Fe3-Ge4-Ti6(2)-Ta13(2). Ta with one small claw. Solenidion ω
<emphasis id="B94ACC3C7439FFAC4860FA9BFAB8FADF" box="[1353,1367,1312,1336]" italics="true" pageId="4" pageNumber="60">1</emphasis>
thick, long, ω
<emphasis id="B94ACC3C7439FFAC4DC0FAFFFF18FABB" box="[233,247,1348,1372]" italics="true" pageId="4" pageNumber="60">2</emphasis>
thinner, finger-shaped, long, φ
<emphasis id="B94ACC3C7439FFAC4F7CFAFFFD8CFABB" box="[597,611,1348,1372]" italics="true" pageId="4" pageNumber="60">1</emphasis>
short, thin, distally thickened, φ
<emphasis id="B94ACC3C7439FFAC4EFAFAFFFC0EFABB" box="[979,993,1348,1372]" italics="true" pageId="4" pageNumber="60">2</emphasis>
short, thin, ω
<emphasis id="B94ACC3C7439FFAC4956FAFFFB62FABB" box="[1151,1165,1348,1372]" italics="true" pageId="4" pageNumber="60">1</emphasis>
&lt;ω
<emphasis id="B94ACC3C7439FFAC49E8FAFFFB20FABB" box="[1217,1231,1348,1372]" italics="true" pageId="4" pageNumber="60">2</emphasis>
&gt; φ
<emphasis id="B94ACC3C7439FFAC49D7FAFFFAE3FABB" box="[1278,1292,1348,1372]" italics="true" pageId="4" pageNumber="60">1</emphasis>
&gt; φ
<emphasis id="B94ACC3C7439FFAC4815FAFFFAA5FABB" box="[1340,1354,1348,1372]" italics="true" pageId="4" pageNumber="60">2</emphasis>
. Leg II (
<figureCitation id="13050CAB7439FFAC4DB6FADCFF0DFA98" box="[159,226,1383,1407]" captionStart="FIGURE 9" captionStartId="9.[151,250,1861,1884]" captionTargetBox="[191,1397,209,1798]" captionTargetId="figure@9.[151,1436,200,1823]" captionTargetPageId="9" captionText="FIGURE 9. Heterodispus cordidiscus n. sp. male. a) leg I; b) leg II; c) leg III; d) leg IV." httpUri="https://zenodo.org/record/282658/files/figure.png" pageId="4" pageNumber="60">Fig. 9</figureCitation>
b). Setal formula: Tr1-Fe3-Ge3-Ti4(1)-Ta7(1). Ta with two small claws and empodium. Solenidion ω thick, long, φ very short. Leg III (
<figureCitation id="13050CAB7439FFAC4CEBFA37FDEBFA43" box="[450,516,1420,1444]" captionStart="FIGURE 9" captionStartId="9.[151,250,1861,1884]" captionTargetBox="[191,1397,209,1798]" captionTargetId="figure@9.[151,1436,200,1823]" captionTargetPageId="9" captionText="FIGURE 9. Heterodispus cordidiscus n. sp. male. a) leg I; b) leg II; c) leg III; d) leg IV." httpUri="https://zenodo.org/record/282658/files/figure.png" pageId="4" pageNumber="60">Fig. 9</figureCitation>
c). Setal formula: Tr1-Fe2-Ge2-Ti4(1)-Ta7. Ta with two small claws and empodium. Solenidion φ small, distally thickened. Leg IV (
<figureCitation id="13050CAB7439FFAC4F9AFA14FD17FA20" box="[691,760,1455,1479]" captionStart="FIGURE 9" captionStartId="9.[151,250,1861,1884]" captionTargetBox="[191,1397,209,1798]" captionTargetId="figure@9.[151,1436,200,1823]" captionTargetPageId="9" captionText="FIGURE 9. Heterodispus cordidiscus n. sp. male. a) leg I; b) leg II; c) leg III; d) leg IV." httpUri="https://zenodo.org/record/282658/files/figure.png" pageId="4" pageNumber="60">Fig. 9</figureCitation>
d). Setal formula: Tr1-Fe2-Ge1-Ti4(1)-Ta6. Distal Ta with 2 adhesive pads of differing size. Solenidion φ thin, finger-shaped, long. Tarsal setae
<emphasis id="B94ACC3C7439FFAC490DFA6EFBAFFA0B" box="[1060,1088,1493,1516]" italics="true" pageId="4" pageNumber="60">tc’</emphasis>
,
<emphasis id="B94ACC3C7439FFAC4967FA6EFB9FFA0B" box="[1102,1136,1493,1516]" italics="true" pageId="4" pageNumber="60">tc”</emphasis>
the longest, average length (n = 12)
<emphasis id="B94ACC3C7439FFAC4DDDFA43FEFFF9E8" box="[244,272,1528,1551]" italics="true" pageId="4" pageNumber="60">tc’</emphasis>
79,
<emphasis id="B94ACC3C7439FFAC4C69FA43FE8DF9E8" box="[320,354,1528,1551]" italics="true" pageId="4" pageNumber="60">tc”</emphasis>
49. Tarsal seta
<emphasis id="B94ACC3C7439FFAC4F23FA43FDC2F9F7" box="[522,557,1528,1552]" italics="true" pageId="4" pageNumber="60">pl’’</emphasis>
and tibial seta
<emphasis id="B94ACC3C7439FFAC4FE6FA43FD04F9E8" box="[719,747,1528,1551]" italics="true" pageId="4" pageNumber="60">v’’</emphasis>
thick, short, spine-like.
</paragraph>
<paragraph id="8B81102E7439FFAC4DEEF9A7FDFCF9D3" blockId="4.[151,1436,1311,2020]" box="[199,531,1564,1588]" pageId="4" pageNumber="60">
<emphasis id="B94ACC3C7439FFAC4DEEF9A7FEFFF9D3" bold="true" box="[199,272,1564,1588]" pageId="4" pageNumber="60">Larva</emphasis>
. Idiosoma length: 171.
</paragraph>
<paragraph id="8B81102E7439FFAC4DEEF9FAFE23F923" blockId="4.[151,1436,1311,2020]" pageId="4" pageNumber="60">
<emphasis id="B94ACC3C7439FFAC4DEEF9FAFECEF9BF" box="[199,289,1601,1624]" italics="true" pageId="4" pageNumber="60">Dorsum</emphasis>
(
<figureCitation id="13050CAB7439FFAC4C19F984FE6BF9B0" box="[304,388,1599,1623]" captionStart="FIGURE 10" captionStartId="10.[151,250,1835,1858]" captionTargetBox="[179,1357,196,1793]" captionTargetId="figure@10.[151,1436,185,1808]" captionTargetPageId="10" captionText="FIGURE 10. Heterodispus cordidiscus n. sp. male larva, dorsal view." httpUri="https://zenodo.org/record/282659/files/figure.png" pageId="4" pageNumber="60">Fig. 10</figureCitation>
). All dorsal setae barbed, on smooth, finely stippled dorsal shields. All setae except
<emphasis id="B94ACC3C7439FFAC481FF9FBFABDF9BF" box="[1334,1362,1600,1624]" italics="true" pageId="4" pageNumber="60">h2</emphasis>
bluntended, setae
<emphasis id="B94ACC3C7439FFAC4C0EF9DFFEABF99B" box="[295,324,1636,1660]" italics="true" pageId="4" pageNumber="60">h2</emphasis>
pointed, situated on outgrowths. Length of dorsal setae
<date id="FF8036EE7439FFAC4E95F9DFFC13F99B" box="[956,1020,1636,1660]" pageId="4" pageNumber="60" value="2010-05-02">
<emphasis id="B94ACC3C7439FFAC4E95F9DFFC38F99B" box="[956,983,1636,1660]" italics="true" pageId="4" pageNumber="60">v2</emphasis>
10
</date>
,
<emphasis id="B94ACC3C7439FFAC4920F9DEFBCFF99B" box="[1033,1056,1637,1660]" italics="true" pageId="4" pageNumber="60">sc</emphasis>
1 25,
<emphasis id="B94ACC3C7439FFAC4949F9DEFB98F99B" box="[1120,1143,1637,1660]" italics="true" pageId="4" pageNumber="60">sc</emphasis>
2 24,
<emphasis id="B94ACC3C7439FFAC499EF9DFFB3DF99B" box="[1207,1234,1636,1660]" italics="true" pageId="4" pageNumber="60">c1</emphasis>
18,
<emphasis id="B94ACC3C7439FFAC482AF9DFFAF1F99B" box="[1283,1310,1636,1660]" italics="true" pageId="4" pageNumber="60">c2</emphasis>
24,
<emphasis id="B94ACC3C7439FFAC4879F9DFFAB1F99B" box="[1360,1374,1636,1660]" italics="true" pageId="4" pageNumber="60">d</emphasis>
29,
<emphasis id="B94ACC3C7439FFAC48A6F9DEFA74F99B" box="[1423,1435,1637,1660]" italics="true" pageId="4" pageNumber="60">e</emphasis>
26,
<emphasis id="B94ACC3C7439FFAC4DE8F933FF26F947" box="[193,201,1672,1696]" italics="true" pageId="4" pageNumber="60">f</emphasis>
24,
<emphasis id="B94ACC3C7439FFAC4DD3F933FEF9F947" box="[250,278,1672,1696]" italics="true" pageId="4" pageNumber="60">h1</emphasis>
30,
<emphasis id="B94ACC3C7439FFAC4C6EF933FE8CF947" box="[327,355,1672,1696]" italics="true" pageId="4" pageNumber="60">h2</emphasis>
40. Distance between dorsal setae:
<date id="FF8036EE7439FFAC4FDDF933FCB3F978" box="[756,860,1671,1696]" pageId="4" pageNumber="60" value="1920-05-02" valueMax="1920-05-02" valueMin="1920-05-02">
<emphasis id="B94ACC3C7439FFAC4FDDF933FCD8F947" box="[756,823,1672,1696]" italics="true" pageId="4" pageNumber="60">v2–v2</emphasis>
20
</date>
,
<emphasis id="B94ACC3C7439FFAC4E41F933FC2FF947" box="[872,960,1672,1696]" italics="true" pageId="4" pageNumber="60">sc1–sc1</emphasis>
20,
<emphasis id="B94ACC3C7439FFAC4EDBF933FBA4F947" box="[1010,1099,1672,1696]" italics="true" pageId="4" pageNumber="60">sc2–sc2</emphasis>
40,
<emphasis id="B94ACC3C7439FFAC4955F933FB50F947" box="[1148,1215,1672,1696]" italics="true" pageId="4" pageNumber="60">c1–c1</emphasis>
28,
<emphasis id="B94ACC3C7439FFAC49D9F933FAF5F947" box="[1264,1306,1672,1696]" italics="true" pageId="4" pageNumber="60">d–d</emphasis>
44,
<emphasis id="B94ACC3C7439FFAC4862F932FA9EF947" box="[1355,1393,1672,1696]" italics="true" pageId="4" pageNumber="60">e–e</emphasis>
41,
<emphasis id="B94ACC3C7439FFAC4DBEF917FF5BF923" box="[151,180,1708,1732]" italics="true" pageId="4" pageNumber="60">f–f</emphasis>
24,
<emphasis id="B94ACC3C7439FFAC4DCCF917FEC4F923" box="[229,299,1708,1732]" italics="true" pageId="4" pageNumber="60">h1–h1</emphasis>
15,
<emphasis id="B94ACC3C7439FFAC4C75F917FE4EF923" box="[348,417,1708,1732]" italics="true" pageId="4" pageNumber="60">h2–h2</emphasis>
13.
</paragraph>
<paragraph id="8B81102E7439FFAC4DEEF96AFABAF8C8" blockId="4.[151,1436,1311,2020]" pageId="4" pageNumber="60">
<emphasis id="B94ACC3C7439FFAC4DEEF96AFEE3F90F" box="[199,268,1745,1768]" italics="true" pageId="4" pageNumber="60">Venter</emphasis>
(
<figureCitation id="13050CAB7439FFAC4C34F974FE82F900" box="[285,365,1743,1767]" captionStart="FIGURE 11" captionStartId="11.[151,250,1810,1833]" captionTargetBox="[227,1393,211,1751]" captionTargetId="figure@11.[167,1419,193,1789]" captionTargetPageId="11" captionText="FIGURE 11. Heterodispus cordidiscus n. sp. male larva, ventral view. The posterior setae ps are covered with particles. gn = gnathosoma." httpUri="https://zenodo.org/record/282660/files/figure.png" pageId="4" pageNumber="60">Fig. 11</figureCitation>
). All ventral setae smooth, thin, pointed. Epimeres I with
<emphasis id="B94ACC3C7439FFAC4ED8F96BFBE2F90F" box="[1009,1037,1744,1768]" italics="true" pageId="4" pageNumber="60">1a</emphasis>
and
<emphasis id="B94ACC3C7439FFAC496BF96BFBB2F90F" box="[1090,1117,1744,1768]" italics="true" pageId="4" pageNumber="60">1b</emphasis>
, epimeres II with
<emphasis id="B94ACC3C7439FFAC4801F96BFAACF90F" box="[1320,1347,1744,1768]" italics="true" pageId="4" pageNumber="60">2a</emphasis>
and
<emphasis id="B94ACC3C7439FFAC4850F96BFA7AF90F" box="[1401,1429,1744,1768]" italics="true" pageId="4" pageNumber="60">2b</emphasis>
, epimeres I and II joined; epimeres III separated, with
<emphasis id="B94ACC3C7439FFAC4FDDF94FFCFFF8EB" box="[756,784,1780,1804]" italics="true" pageId="4" pageNumber="60">3a</emphasis>
and
<emphasis id="B94ACC3C7439FFAC4E6EF94FFC8CF8EB" box="[839,867,1780,1804]" italics="true" pageId="4" pageNumber="60">3b</emphasis>
. Length of ventral setae
<emphasis id="B94ACC3C7439FFAC4952F94FFB77F8EB" box="[1147,1176,1780,1804]" italics="true" pageId="4" pageNumber="60">1a</emphasis>
8,
<emphasis id="B94ACC3C7439FFAC4992F94FFB28F8EB" box="[1211,1223,1780,1804]" italics="true" pageId="4" pageNumber="60">1</emphasis>
b 10,
<emphasis id="B94ACC3C7439FFAC4820F94FFACBF8EB" box="[1289,1316,1780,1804]" italics="true" pageId="4" pageNumber="60">2a</emphasis>
11,
<emphasis id="B94ACC3C7439FFAC487CF94FFA8CF8EB" box="[1365,1379,1780,1804]" italics="true" pageId="4" pageNumber="60">2</emphasis>
b 12, 3
<emphasis id="B94ACC3C7439FFAC4D8CF8A2FF5CF8D7" box="[165,179,1817,1840]" italics="true" pageId="4" pageNumber="60">a</emphasis>
10, 3
<emphasis id="B94ACC3C7439FFAC4DDBF8A3FEEFF8D7" box="[242,256,1816,1840]" italics="true" pageId="4" pageNumber="60">b</emphasis>
10. Distance between ventral setae
<emphasis id="B94ACC3C7439FFAC4FBBF8A3FD37F8D7" box="[658,728,1816,1840]" italics="true" pageId="4" pageNumber="60">1a–1a</emphasis>
18,
<emphasis id="B94ACC3C7439FFAC4E20F8A3FCA0F8D7" box="[777,847,1816,1840]" italics="true" pageId="4" pageNumber="60">1b–1b</emphasis>
31,
<emphasis id="B94ACC3C7439FFAC4EA9F8A3FC62F8D7" box="[896,909,1816,1840]" italics="true" pageId="4" pageNumber="60">2</emphasis>
a–2a 21,
<emphasis id="B94ACC3C7439FFAC4EDEF8A3FBEBF8D7" box="[1015,1028,1816,1840]" italics="true" pageId="4" pageNumber="60">2</emphasis>
b–2b 48,
<emphasis id="B94ACC3C7439FFAC4944F8A3FB95F8D7" box="[1133,1146,1816,1840]" italics="true" pageId="4" pageNumber="60">3</emphasis>
a–3a 38,
<emphasis id="B94ACC3C7439FFAC49CDF8A3FB1DF8D7" box="[1252,1266,1816,1840]" italics="true" pageId="4" pageNumber="60">3</emphasis>
b–3b 37.
</paragraph>
<paragraph id="8B81102E7439FFAC4DEEF887FC64F87B" blockId="4.[151,1436,1311,2020]" pageId="4" pageNumber="60">
<emphasis id="B94ACC3C7439FFAC4DEEF887FEB9F8B3" box="[199,342,1852,1876]" italics="true" pageId="4" pageNumber="60">Gnathosoma</emphasis>
(
<figureCitation id="13050CAB7439FFAC4C4CF887FE48F8B3" box="[357,423,1852,1876]" captionStart="FIGURE 5" captionStartId="6.[151,250,590,613]" captionTargetBox="[161,1421,159,569]" captionTargetId="figure@6.[153,1435,153,575]" captionTargetPageId="6" captionText="FIGURE 5. Heterodispus cordidiscus n. sp. a) gnathosoma of male larva, dorsal view; b) gnathosoma of female, ventral view; c)" httpUri="https://zenodo.org/record/282654/files/figure.png" pageId="4" pageNumber="60">Fig. 5</figureCitation>
a) roundish, width 20, length 15; two pairs of dorsal setae
<emphasis id="B94ACC3C7439FFAC491BF887FBB4F8B3" box="[1074,1115,1852,1876]" italics="true" pageId="4" pageNumber="60">ch1</emphasis>
and
<emphasis id="B94ACC3C7439FFAC49B8F887FB55F8B3" box="[1169,1210,1852,1876]" italics="true" pageId="4" pageNumber="60">ch2</emphasis>
,
<emphasis id="B94ACC3C7439FFAC49EEF887FB00F8B3" box="[1223,1263,1852,1876]" italics="true" pageId="4" pageNumber="60">ch1</emphasis>
short and oval,
<emphasis id="B94ACC3C7439FFAC4DBEF8DBFF2FF89F" box="[151,192,1888,1912]" italics="true" pageId="4" pageNumber="60">ch2</emphasis>
thin, smooth; one pair of ventral setae
<emphasis id="B94ACC3C7439FFAC4FA0F8DAFD4DF89F" box="[649,674,1889,1912]" italics="true" pageId="4" pageNumber="60">su</emphasis>
, thin, smooth; two pairs of palpal setae, each palpus with 1 small ventral palpal solenidion and small accessory setigenous structures.
</paragraph>
<paragraph id="8B81102E7439FFAC4DEEF812FE64F803" blockId="4.[151,1436,1311,2020]" pageId="4" pageNumber="60">
<emphasis id="B94ACC3C7439FFAC4DEEF812FF13F827" box="[199,252,1961,1984]" italics="true" pageId="4" pageNumber="60">Legs</emphasis>
. Leg I (
<figureCitation id="13050CAB7439FFAC4C7AF81CFE4AF858" box="[339,421,1959,1983]" captionStart="FIGURE 12" captionStartId="12.[151,250,1002,1025]" captionTargetBox="[200,1418,200,962]" captionTargetId="figure@12.[151,1436,193,979]" captionTargetPageId="12" captionText="FIGURE 12. Heterodispus cordidiscus n. sp. male larva. a) legI; b) leg II; c) leg III." httpUri="https://zenodo.org/record/282661/files/figure.png" pageId="4" pageNumber="60">Fig. 12</figureCitation>
a). Setal formula: Tr0-Fe3-Ge4-Ti6(1)-Ta11(1). Solenidia ω and φ finger-shaped, ω&gt; φ. Ta with two small claws.
</paragraph>
<paragraph id="8B81102E7438FFAD4DEEFF23FD07FF10" blockId="5.[151,1436,151,247]" pageId="5" pageNumber="61">
Leg II (
<figureCitation id="13050CAB7438FFAD4C36FF2CFE9CFF48" box="[287,371,151,175]" captionStart="FIGURE 12" captionStartId="12.[151,250,1002,1025]" captionTargetBox="[200,1418,200,962]" captionTargetId="figure@12.[151,1436,193,979]" captionTargetPageId="12" captionText="FIGURE 12. Heterodispus cordidiscus n. sp. male larva. a) legI; b) leg II; c) leg III." httpUri="https://zenodo.org/record/282661/files/figure.png" pageId="5" pageNumber="61">Fig. 12</figureCitation>
b). Setal formula: Tr0-Fe3-Ge3-Ti4(1)-Ta7(1). Solenidia ω and φ short, thick, ω ≥ φ. Ta with two small claws and empodium. Leg III (
<figureCitation id="13050CAB7438FFAD4F5EFF07FD23FF33" box="[631,716,188,212]" captionStart="FIGURE 12" captionStartId="12.[151,250,1002,1025]" captionTargetBox="[200,1418,200,962]" captionTargetId="figure@12.[151,1436,193,979]" captionTargetPageId="12" captionText="FIGURE 12. Heterodispus cordidiscus n. sp. male larva. a) legI; b) leg II; c) leg III." httpUri="https://zenodo.org/record/282661/files/figure.png" pageId="5" pageNumber="61">Fig. 12</figureCitation>
c) Setal formula: Tr0-Fe2-Ge2-Ti4(1)-Ta7. Solenidion φ small, finger-shaped. Ta with 2 small claws and empodium.
</paragraph>
</subSubSection>
<caption id="DF4140A67438FFAD4DBEF81FFA72F85D" box="[151,1437,1956,1979]" httpUri="https://zenodo.org/record/282653/files/figure.png" pageId="5" pageNumber="61" targetBox="[190,1426,329,1913]" targetPageId="5">
<paragraph id="8B81102E7438FFAD4DBEF81FFA72F85D" blockId="5.[151,1437,1956,1979]" box="[151,1437,1956,1979]" pageId="5" pageNumber="61">
<emphasis id="B94ACC3C7438FFAD4DBEF81FFEFDF85D" bold="true" box="[151,274,1956,1979]" pageId="5" pageNumber="61">FIGURE 4.</emphasis>
<taxonomicName id="4C3E6BAD7438FFAD4C31F81EFDF7F85C" box="[280,536,1957,1979]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="61" phylum="Arthropoda" rank="species" species="cordidiscus" status="sp. nov.">
<emphasis id="B94ACC3C7438FFAD4C31F81EFDF7F85C" box="[280,536,1957,1979]" italics="true" pageId="5" pageNumber="61">Heterodispus cordidiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27971477438FFAD4F34F81EFDBCF85D" box="[541,595,1957,1978]" pageId="5" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
female holotype. a) leg I; b) leg II; c) leg III; d) leg IV; e) lateral part of prodorsum.
</paragraph>
</caption>
<caption id="DF4140A6743BFFAE4DBEFDF5FF44FD65" httpUri="https://zenodo.org/record/282654/files/figure.png" pageId="6" pageNumber="62" targetBox="[161,1421,159,569]" targetPageId="6">
<paragraph id="8B81102E743BFFAE4DBEFDF5FF44FD65" blockId="6.[151,1436,590,642]" pageId="6" pageNumber="62">
<emphasis id="B94ACC3C743BFFAE4DBEFDF5FEFCFD83" bold="true" box="[151,275,590,613]" pageId="6" pageNumber="62">FIGURE 5.</emphasis>
<taxonomicName id="4C3E6BAD743BFFAE4C30FDF5FDF6FD83" box="[281,537,590,612]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="cordidiscus" status="sp. nov.">
<emphasis id="B94ACC3C743BFFAE4C30FDF5FDF6FD83" box="[281,537,590,612]" italics="true" pageId="6" pageNumber="62">Heterodispus cordidiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2797147743BFFAE4F09FDF4FDB8FD83" box="[544,599,591,612]" pageId="6" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
a) gnathosoma of male larva, dorsal view; b) gnathosoma of female, ventral view; c)
</paragraph>
</caption>
<caption id="DF4140A6743BFFAE4DBEF862FA9CF817" box="[151,1395,2009,2032]" httpUri="https://zenodo.org/record/282655/files/figure.png" pageId="6" pageNumber="62" targetBox="[185,1384,622,1986]" targetPageId="6">
<paragraph id="8B81102E743BFFAE4DBEF862FA9CF817" blockId="6.[151,1395,2009,2032]" box="[151,1395,2009,2032]" pageId="6" pageNumber="62">
<emphasis id="B94ACC3C743BFFAE4DBEF862FEFDF808" bold="true" box="[151,274,2009,2032]" pageId="6" pageNumber="62">FIGURE 6.</emphasis>
<taxonomicName id="4C3E6BAD743BFFAE4C30F861FDF6F817" box="[281,537,2010,2032]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="cordidiscus" status="sp. nov.">
<emphasis id="B94ACC3C743BFFAE4C30F861FDF6F817" box="[281,537,2010,2032]" italics="true" pageId="6" pageNumber="62">Heterodispus cordidiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2797147743BFFAE4F36F860FDB9F817" box="[543,598,2011,2032]" pageId="6" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
male, dorsal view. ae = aedeagus, dej = ejaculatory duct, sv = seminar vesicle.
</paragraph>
</caption>
<caption id="DF4140A6743AFFAF4DBEF91AFC3DF950" box="[151,978,1697,1720]" httpUri="https://zenodo.org/record/282656/files/figure.png" pageId="7" pageNumber="63" targetBox="[275,1305,196,1663]" targetPageId="7">
<paragraph id="8B81102E743AFFAF4DBEF91AFC3DF950" blockId="7.[151,978,1697,1720]" box="[151,978,1697,1720]" pageId="7" pageNumber="63">
<emphasis id="B94ACC3C743AFFAF4DBEF91AFEFCF950" bold="true" box="[151,275,1697,1720]" pageId="7" pageNumber="63">FIGURE 7.</emphasis>
<taxonomicName id="4C3E6BAD743AFFAF4C30F91AFDF6F950" box="[281,537,1697,1719]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="63" phylum="Arthropoda" rank="species" species="cordidiscus" status="sp. nov.">
<emphasis id="B94ACC3C743AFFAF4C30F91AFDF6F950" box="[281,537,1697,1719]" italics="true" pageId="7" pageNumber="63">Heterodispus cordidiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2797147743AFFAF4F36F919FDB9F950" box="[543,598,1698,1719]" pageId="7" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
male, ventral view. gn =gnathosoma.
</paragraph>
</caption>
<subSubSection id="C32443A5743AFFAF4DEEF964FE25F8FE" pageId="7" pageNumber="63" type="etymology">
<paragraph id="8B81102E743AFFAF4DEEF964FE25F8FE" blockId="7.[151,1436,1759,2026]" pageId="7" pageNumber="63">
<emphasis id="B94ACC3C743AFFAF4DEEF964FEA9F910" bold="true" box="[199,326,1759,1783]" pageId="7" pageNumber="63">Etymology</emphasis>
. The species name
<taxonomicName id="4C3E6BAD743AFFAF4F0AF964FD4CF910" box="[547,675,1759,1783]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="63" phylum="Arthropoda" rank="species" species="cordidiscus">
<emphasis id="B94ACC3C743AFFAF4F0AF964FD4CF910" box="[547,675,1759,1783]" italics="true" pageId="7" pageNumber="63">cordidiscus</emphasis>
</taxonomicName>
(lat. cor = heart; discus = plate) refers to the distinctly heart-shaped genital capsule of the male.
</paragraph>
</subSubSection>
<subSubSection id="C32443A5743AFFA04DEEF89EFD2AF833" lastPageId="8" lastPageNumber="64" pageId="7" pageNumber="63" type="materials_examined">
<paragraph id="8B81102E743AFFAF4DEEF89EFE8EF866" blockId="7.[151,1436,1759,2026]" pageId="7" pageNumber="63">
<emphasis id="B94ACC3C743AFFAF4DEEF89EFE82F8DA" bold="true" box="[199,365,1828,1853]" pageId="7" pageNumber="63">
<typeStatus id="5485AE8C743AFFAF4DEEF89EFEEDF8DA" box="[199,258,1829,1853]" pageId="7" pageNumber="63">Type</typeStatus>
material
</emphasis>
. Male
<typeStatus id="5485AE8C743AFFAF4C93F89FFDF0F8DB" box="[442,543,1828,1852]" pageId="7" pageNumber="63" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D38DBA7743AFFAF4F00F89FFD6CF8DB" box="[553,643,1828,1852]" pageId="7" pageNumber="63" type="male">11 male</specimenCount>
,
<specimenCount id="9D38DBA7743AFFAF4FA7F89FFCEFF8DB" box="[654,768,1828,1852]" pageId="7" pageNumber="63" type="female">57 female</specimenCount>
, and 1 larva
<typeStatus id="5485AE8C743AFFAF4EA5F89EFC17F8DA" box="[908,1016,1829,1853]" pageId="7" pageNumber="63" type="paratype">paratypes</typeStatus>
(laboratory culture),
<collectingCountry id="F32950BE743AFFAF49CDF89FFAACF8DA" box="[1252,1347,1828,1853]" name="Egypt" pageId="7" pageNumber="63">
<emphasis id="B94ACC3C743AFFAF49CDF89FFAACF8DA" bold="true" box="[1252,1347,1828,1853]" pageId="7" pageNumber="63">EGYPT</emphasis>
</collectingCountry>
: Luxor, Nile riverside (2541’38.02’’N, 3237’41.18’’E), soil sample taken from sandy ground with roots;
<date id="FF8036EE743AFFAF49CFF8FCFA77F8B8" box="[1254,1432,1863,1888]" pageId="7" pageNumber="63" value="1985-01-01">1 January 1985</date>
, leg. E. Ebermann.
</paragraph>
<paragraph id="8B81102E743AFFAF4DEEF836FBC8F843" blockId="7.[151,1436,1759,2026]" box="[199,1063,1932,1957]" pageId="7" pageNumber="63">
<emphasis id="B94ACC3C743AFFAF4DEEF836FE6CF842" bold="true" box="[199,387,1932,1957]" pageId="7" pageNumber="63">
<typeStatus id="5485AE8C743AFFAF4DEEF836FEEDF842" box="[199,258,1933,1957]" pageId="7" pageNumber="63">Type</typeStatus>
deposition
</emphasis>
. The
<typeStatus id="5485AE8C743AFFAF4CEDF836FE1BF842" box="[452,500,1933,1957]" pageId="7" pageNumber="63">type</typeStatus>
material is deposited at the following institutions:
</paragraph>
<paragraph id="8B81102E743AFFAF4DEEF814FB89F80E" blockId="7.[151,1436,1759,2026]" pageId="7" pageNumber="63">
<typeStatus id="5485AE8C743AFFAF4DEEF814FEDDF820" box="[199,306,1967,1991]" pageId="7" pageNumber="63" type="holotype">Holotype</typeStatus>
, 10
<typeStatus id="5485AE8C743AFFAF4C4DF80BFE20F82F" box="[356,463,1968,1992]" pageId="7" pageNumber="63" type="paratype">paratypes</typeStatus>
(females), 2
<typeStatus id="5485AE8C743AFFAF4F43F80BFD3AF82F" box="[618,725,1968,1992]" pageId="7" pageNumber="63" type="paratype">paratypes</typeStatus>
(males): Zoologisches Institut und Zoologisches Museum der Universität Hamburg
<collectionCode id="ED2F88EB743AFFAF4CA3F869FE30F80E" box="[394,479,2002,2025]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="7" pageNumber="63">ZMHU</collectionCode>
,
<collectingCountry id="F32950BE743AFFAF4CC2F86AFDBCF80E" box="[491,595,2001,2025]" name="Germany" pageId="7" pageNumber="63">Germany</collectingCountry>
(
<typeStatus id="5485AE8C743AFFAF4F48F86AFD24F80E" box="[609,715,2001,2025]" pageId="7" pageNumber="63" type="holotype">Holotype</typeStatus>
Inv.No. A35/12,
<typeStatus id="5485AE8C743AFFAF4EA6F869FC15F80D" box="[911,1018,2002,2026]" pageId="7" pageNumber="63" type="paratype">paratypes</typeStatus>
A36/12);
</paragraph>
<paragraph id="8B81102E7435FFA04DEEFF2CFE28FF35" blockId="8.[151,1436,151,838]" pageId="8" pageNumber="64">
10
<typeStatus id="5485AE8C7435FFA04DDAFF23FEB0FF57" box="[243,351,152,176]" pageId="8" pageNumber="64" type="paratype">paratypes</typeStatus>
(females), 2
<typeStatus id="5485AE8C7435FFA04F27FF23FD96FF57" box="[526,633,152,176]" pageId="8" pageNumber="64" type="paratype">paratypes</typeStatus>
(males): Senckenberg Museum für Naturkunde Görlitz,
<collectingCountry id="F32950BE7435FFA0481DFF2CFA73FF48" box="[1332,1436,151,175]" name="Germany" pageId="8" pageNumber="64">Germany</collectingCountry>
(Collection No. 85/47331);
</paragraph>
<paragraph id="8B81102E7435FFA04DEEFF66FE4FFEF0" blockId="8.[151,1436,151,838]" pageId="8" pageNumber="64">
10
<typeStatus id="5485AE8C7435FFA04DDBFF65FEB2FF11" box="[242,349,222,246]" pageId="8" pageNumber="64" type="paratype">paratypes</typeStatus>
(females), 2
<typeStatus id="5485AE8C7435FFA04F20FF65FD9BFF11" box="[521,628,222,246]" pageId="8" pageNumber="64" type="paratype">paratypes</typeStatus>
(males): Naturhistorisches Museum Wien,
<collectingCountry id="F32950BE7435FFA049ADFF66FB39FF12" box="[1156,1238,221,245]" name="Austria" pageId="8" pageNumber="64">Austria</collectingCountry>
(Collection No.
<collectionCode id="ED2F88EB7435FFA04DBEFEBBFF1CFEF0" box="[151,243,256,279]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="64">NHMW</collectionCode>
21855-21866);
</paragraph>
<paragraph id="8B81102E7435FFA04DEEFE99FB09FEDD" blockId="8.[151,1436,151,838]" box="[199,1254,290,315]" pageId="8" pageNumber="64">
10
<typeStatus id="5485AE8C7435FFA04DC3FE98FEBAFEDC" box="[234,341,291,315]" pageId="8" pageNumber="64" type="paratype">paratypes</typeStatus>
(females), 2
<typeStatus id="5485AE8C7435FFA04CC0FE98FDBAFEDC" box="[489,597,291,315]" pageId="8" pageNumber="64" type="paratype">paratypes</typeStatus>
(males): Shmalgausen Institute of Zoology Kiev,
<collectingCountry id="F32950BE7435FFA049ADFE99FB0DFEDD" box="[1156,1250,290,314]" name="Ukraine" pageId="8" pageNumber="64">Ukraine</collectingCountry>
;
</paragraph>
<paragraph id="8B81102E7435FFA04DEEFEFEFEDAFE98" blockId="8.[151,1436,151,838]" pageId="8" pageNumber="64">
16
<typeStatus id="5485AE8C7435FFA04DC6FEFDFEB5FEB9" box="[239,346,326,350]" pageId="8" pageNumber="64" type="paratype">paratypes</typeStatus>
(females), 4
<typeStatus id="5485AE8C7435FFA04CD5FEFDFD88FEB9" box="[508,615,326,350]" pageId="8" pageNumber="64" type="paratype">paratypes</typeStatus>
(males),
<specimenCount id="9D38DBA77435FFA04FF3FEFEFCC5FEBA" box="[730,810,325,349]" pageId="8" pageNumber="64" type="male">1 male</specimenCount>
larva: Institute of Zoology, Karl-Franzens-University Graz,
<collectingCountry id="F32950BE7435FFA04DF2FEDCFEDEFE98" box="[219,305,359,383]" name="Austria" pageId="8" pageNumber="64">Austria</collectingCountry>
.
</paragraph>
<paragraph id="8B81102E7435FFA04DEEFE31FB9CFD5F" blockId="8.[151,1436,151,838]" pageId="8" pageNumber="64">
<emphasis id="B94ACC3C7435FFA04DEEFE31FE20FE44" bold="true" box="[199,463,394,419]" pageId="8" pageNumber="64">Differential diagnosis.</emphasis>
<taxonomicName id="4C3E6BAD7435FFA04CFEFE30FD1CFE44" box="[471,755,395,419]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="cordidiscus">
<emphasis id="B94ACC3C7435FFA04CFEFE30FD1CFE44" box="[471,755,395,419]" italics="true" pageId="8" pageNumber="64">Heterodispus cordidiscus</emphasis>
</taxonomicName>
is separated from all other
<taxonomicName id="4C3E6BAD7435FFA04904FE30FB2EFE44" box="[1069,1217,395,419]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B94ACC3C7435FFA04904FE30FB2EFE44" box="[1069,1217,395,419]" italics="true" pageId="8" pageNumber="64">Heterodispus</emphasis>
</taxonomicName>
-species because of length differences of particular setae on the body and, most clearly, by length differences of tibial and tarsal setae on leg IV. In
<taxonomicName id="4C3E6BAD7435FFA04C06FE6AFE3CFE0F" box="[303,467,464,488]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="cordidiscus">
<emphasis id="B94ACC3C7435FFA04C06FE6AFE3CFE0F" box="[303,467,464,488]" italics="true" pageId="8" pageNumber="64">H. cordidiscus</emphasis>
</taxonomicName>
, the ratio of the three most peculiar tarsal setae is
<emphasis id="B94ACC3C7435FFA0493EFE6BFBDBFE00" box="[1047,1076,464,487]" italics="true" pageId="8" pageNumber="64">tc’</emphasis>
~
<emphasis id="B94ACC3C7435FFA0497DFE6BFB97FE00" box="[1108,1144,464,487]" italics="true" pageId="8" pageNumber="64">tc’’</emphasis>
&gt;
<emphasis id="B94ACC3C7435FFA049B0FE6BFB2CFE00" box="[1177,1219,464,487]" italics="true" pageId="8" pageNumber="64">pv’’</emphasis>
, while in the other species it is
<emphasis id="B94ACC3C7435FFA04C06FE48FEA3FDED" box="[303,332,499,522]" italics="true" pageId="8" pageNumber="64">tc’</emphasis>
&gt;
<emphasis id="B94ACC3C7435FFA04C5DFE48FE70FDED" box="[372,415,499,522]" italics="true" pageId="8" pageNumber="64">pv’’</emphasis>
&gt;
<emphasis id="B94ACC3C7435FFA04CEEFE48FE05FDED" box="[455,490,499,522]" italics="true" pageId="8" pageNumber="64">tc’’</emphasis>
. Moreover, the tibial seta
<emphasis id="B94ACC3C7435FFA04E02FE48FCD6FDEC" box="[811,825,499,523]" italics="true" pageId="8" pageNumber="64">d</emphasis>
, which is comparatively short in
<taxonomicName id="4C3E6BAD7435FFA049E7FE4FFA99FDEC" box="[1230,1398,499,523]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="cordidiscus">
<emphasis id="B94ACC3C7435FFA049E7FE4FFA99FDEC" box="[1230,1398,499,523]" italics="true" pageId="8" pageNumber="64">H. cordidiscus</emphasis>
</taxonomicName>
, is pronounced in all other Egyptian species and most other
<taxonomicName id="4C3E6BAD7435FFA04E02FDAEFC50FDCA" box="[811,959,533,557]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B94ACC3C7435FFA04E02FDAEFC50FDCA" box="[811,959,533,557]" italics="true" pageId="8" pageNumber="64">Heterodispus</emphasis>
</taxonomicName>
-species. The tarsal setae on leg IV of
<taxonomicName id="4C3E6BAD7435FFA048A8FDADFE79FDA8" authority="Mahunka, 1977" authorityName="Mahunka" authorityYear="1977" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="paolii">
<emphasis id="B94ACC3C7435FFA048A8FDADFF36FDB7" italics="true" pageId="8" pageNumber="64">H. paolii</emphasis>
<bibRefCitation id="EFAF6DDF7435FFA04DC8FD8CFE79FDA8" author="Mahunka" box="[225,406,567,591]" pageId="8" pageNumber="64" refString="Mahunka, S. (1977) Beitrage zur Kenntnis der Systematik, Taxonomie, Ontogenie, Okologie und Verbreitung der Tarsonemiden. III. Folia Entomologica Hungarica, 30, 1, 85 - 97." type="journal article" year="1977">Mahunka, 1977</bibRefCitation>
</taxonomicName>
, are similar to those of
<taxonomicName id="4C3E6BAD7435FFA04F80FD82FCA2FDB7" box="[681,845,568,592]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="cordidiscus">
<emphasis id="B94ACC3C7435FFA04F80FD82FCA2FDB7" box="[681,845,568,592]" italics="true" pageId="8" pageNumber="64">H. cordidiscus</emphasis>
</taxonomicName>
, but the tibial seta
<emphasis id="B94ACC3C7435FFA04907FD83FBD3FDB7" box="[1070,1084,568,592]" italics="true" pageId="8" pageNumber="64">d</emphasis>
of this species is long and the dorsal setae
<emphasis id="B94ACC3C7435FFA04C0BFDE0FED4FD94" box="[290,315,603,627]" italics="true" pageId="8" pageNumber="64">c1</emphasis>
,
<emphasis id="B94ACC3C7435FFA04C62FDE0FE8AFD94" box="[331,357,603,627]" italics="true" pageId="8" pageNumber="64">c2</emphasis>
and
<emphasis id="B94ACC3C7435FFA04CB5FDE0FE45FD94" box="[412,426,603,627]" italics="true" pageId="8" pageNumber="64">d</emphasis>
are smooth. In the first description of
<taxonomicName id="4C3E6BAD7435FFA04E72FDE7FC05FD94" box="[859,1002,603,627]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B94ACC3C7435FFA04E72FDE7FC05FD94" box="[859,1002,603,627]" italics="true" pageId="8" pageNumber="64">H. elongatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFAF6DDF7435FFA0493DFDE1FB21FD95" author="Tragardh" box="[1044,1230,602,626]" pageId="8" pageNumber="64" refString="Tragardh, I. (1905) Acariden aus Agypten und dem Sudan. Zugleich ein Beitrag zur Kenntnis der Entwicklungsgeschichte der Gattungen Phytoptipalpus, Pimeliaphilus, Pterygosoma und Podapolipus. Results of the Swedish Zoological Expedition to Egypt and the White Nile 1901 under the direction of L. A. Jagerskiold, Part II, 20, 1 - 124." type="journal article" year="1905">Trägårdh (1905)</bibRefCitation>
, the illustration of the setae on the legs is not clear. However,
<taxonomicName id="4C3E6BAD7435FFA04FA9FDC5FCCCFD72" box="[640,803,637,661]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="cordidiscus">
<emphasis id="B94ACC3C7435FFA04FA9FDC5FCCCFD72" box="[640,803,637,661]" italics="true" pageId="8" pageNumber="64">H. cordidiscus</emphasis>
</taxonomicName>
is separated from this species by its longer setae
<emphasis id="B94ACC3C7435FFA04878FDC6FA83FD72" box="[1361,1388,637,661]" italics="true" pageId="8" pageNumber="64">c2</emphasis>
and by the non-existent length difference
<emphasis id="B94ACC3C7435FFA04F10FD1BFDBAFD5F" box="[569,597,672,696]" italics="true" pageId="8" pageNumber="64">3b</emphasis>
&gt;
<emphasis id="B94ACC3C7435FFA04F5AFD1BFD60FD5F" box="[627,655,672,696]" italics="true" pageId="8" pageNumber="64">3a</emphasis>
which is clearly displayed in
<taxonomicName id="4C3E6BAD7435FFA04EF4FD1AFB82FD5F" box="[989,1133,672,696]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B94ACC3C7435FFA04EF4FD1AFB82FD5F" box="[989,1133,672,696]" italics="true" pageId="8" pageNumber="64">H. elongatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B81102E7435FFA04DEEFD79FD7EFCA1" blockId="8.[151,1436,151,838]" pageId="8" pageNumber="64">
Males of
<taxonomicName id="4C3E6BAD7435FFA04C11FD7FFE32FD3C" box="[312,477,707,731]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="cordidiscus">
<emphasis id="B94ACC3C7435FFA04C11FD7FFE32FD3C" box="[312,477,707,731]" italics="true" pageId="8" pageNumber="64">H. cordidiscus</emphasis>
</taxonomicName>
do not possess piston-like appendages on the gnathosoma that can be seen in the genera
<taxonomicName id="4C3E6BAD7435FFA04DC4FD5CFE85FD19" box="[237,362,743,766]" class="Arachnida" family="Scutacaridae" genus="Scutacarus" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B94ACC3C7435FFA04DC4FD5CFE85FD19" box="[237,362,743,766]" italics="true" pageId="8" pageNumber="64">Scutacarus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFAF6DDF7435FFA04C55FD5DFDD9FD19" author="Ebermann" box="[380,566,742,766]" pageId="8" pageNumber="64" refString="Ebermann, E. (1983) Zur Morphologie der Milbenart Scutacarus peractus Karafiat, 1959 (Acari, Trombidiformes). Mittteilungen des naturwissenschaftlichen Vereines fur Steiermark, 113, 91 - 98." type="journal article" year="1983">Ebermann 1983</bibRefCitation>
,
<bibRefCitation id="EFAF6DDF7435FFA04F6EFD5CFCD1FD19" author="Norton" box="[583,830,742,766]" pageId="8" pageNumber="64" refString="Norton, R. A. &amp; Ide, G. S. (1974) Scutacarus baculitarsus agaricus n. subsp. (Acarina: Scutacaridae) from commercial mushroom houses, with notes on phoretic behaviour. Journal of the Kansas Entomological Society, 47, 4, 527 - 534." type="journal article" year="1974">Norton and Ide 1974</bibRefCitation>
) and
<taxonomicName id="4C3E6BAD7435FFA04EACFD5DFB17FD19" authority="(Ebermann 1982a)" baseAuthorityName="Ebermann" baseAuthorityYear="1982" box="[901,1272,742,766]" class="Arachnida" family="Scutacaridae" genus="Lophodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B94ACC3C7435FFA04EACFD5DFBFBFD19" box="[901,1044,742,766]" italics="true" pageId="8" pageNumber="64">Lophodispus</emphasis>
(
<bibRefCitation id="EFAF6DDF7435FFA0490EFD5DFB1FFD19" author="Ebermann" box="[1063,1264,742,766]" pageId="8" pageNumber="64" refString="Ebermann, E. (1982 a) Zuchtversuche und morphologische Untersuchungen an heimischen Milben (Acari, Scutacaridae). Mittteilungen des naturwissenschaftlichen Vereines fur Steiermark, 112, 155 - 165." type="journal article" year="1982" yearSuffix="a">Ebermann 1982a</bibRefCitation>
)
</taxonomicName>
. Males of the genera
<taxonomicName id="4C3E6BAD7435FFA04DC4FCB7FEB0FCC4" box="[237,351,780,803]" class="Arachnida" family="Scutacaridae" genus="Imparipes" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B94ACC3C7435FFA04DC4FCB7FEB0FCC4" box="[237,351,780,803]" italics="true" pageId="8" pageNumber="64">Imparipes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFAF6DDF7435FFA04C5BFCB1FDD4FCC5" author="Ebermann" box="[370,571,778,802]" pageId="8" pageNumber="64" refString="Ebermann, E. (1982 b) Fortpflanzungsbiologische Studien an Scutacariden (Acari, Trombidiformes). Zoologische Jahrbucher (Systematik), 109, 98 - 116." type="journal article" year="1982" yearSuffix="b">Ebermann 1982b</bibRefCitation>
,
<bibRefCitation id="EFAF6DDF7435FFA04F62FCB1FC9DFCC5" author="Delfinado" box="[587,882,778,802]" pageId="8" pageNumber="64" refString="Delfinado, M. D. &amp; Baker, E. W. (1976) New species of Scutacaridae (Acarina) associated with insects. Acarologia, 18, 2, 264 - 301." type="journal article" year="1976">
Delfinado &amp;
<collectingCountry id="F32950BE7435FFA04FC1FCB1FCC4FCC5" box="[744,811,778,802]" name="United States Pacific Island Wildlife Refuges" pageId="8" pageNumber="64">Baker</collectingCountry>
1976
</bibRefCitation>
,
<bibRefCitation id="EFAF6DDF7435FFA04EA8FCB1FB3FFCC5" author="Khaustov" box="[897,1232,778,802]" pageId="8" pageNumber="64" refString="Khaustov, A. A. &amp; Chydydrov, P. R. (2004) New species of mites of the family Scutacaridae (Acari: Heterostigmata) associated with ants (Hymenoptera, Formicidae) from Turkmenistan. Acarina, 12, 2, 87 - 103." type="journal article" year="2004">Khaustov &amp; Chydyrov 2004</bibRefCitation>
) and
<taxonomicName id="4C3E6BAD7435FFA0483EFCB0FEB4FCA1" authority="Ebermann 1994" authorityName="Ebermann" authorityYear="1994" class="Arachnida" family="Scutacaridae" genus="Archidispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B94ACC3C7435FFA0483EFCB0FA73FCC4" box="[1303,1436,779,803]" italics="true" pageId="8" pageNumber="64">Archidispus</emphasis>
(
<bibRefCitation id="EFAF6DDF7435FFA04DB7FC95FEBCFCA1" author="Ebermann" box="[158,339,814,838]" pageId="8" pageNumber="64" refString="Ebermann, E. (1994) First demonstration of males in species of the genus Archidispus Karafiat (Acari, Tarsonemina, Scutacaridae). International Journal of Acarology, 20, 3, 169 - 182." type="journal article" year="1994">Ebermann 1994</bibRefCitation>
)
</taxonomicName>
also lack such appendages.
</paragraph>
<caption id="DF4140A67435FFA04DBEF990FD5DF987" httpUri="https://zenodo.org/record/282657/files/figure.png" pageId="8" pageNumber="64" targetBox="[455,1134,932,1555]" targetPageId="8">
<paragraph id="8B81102E7435FFA04DBEF990FD5DF987" blockId="8.[151,1436,1579,1632]" pageId="8" pageNumber="64">
<emphasis id="B94ACC3C7435FFA04DBEF990FEFDF9A6" bold="true" box="[151,274,1579,1602]" pageId="8" pageNumber="64">FIGURE 8.</emphasis>
<taxonomicName id="4C3E6BAD7435FFA04C31F997FDF7F9A5" box="[280,536,1580,1602]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="cordidiscus" status="sp. nov.">
<emphasis id="B94ACC3C7435FFA04C31F997FDF7F9A5" box="[280,536,1580,1602]" italics="true" pageId="8" pageNumber="64">Heterodispus cordidiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27971477435FFA04F34F996FDBCF9A5" box="[541,595,1581,1602]" pageId="8" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
male, lateral view of the genital apparatus. ae = aedeagus, dej = ejaculatory duct, lo = internal loop of the aedeagus, sv = seminar vesicle.
</paragraph>
</caption>
<paragraph id="8B81102E7435FFA04DEEF915FCF1F88A" blockId="8.[151,1436,1710,2004]" pageId="8" pageNumber="64">
The dorsal setae
<date id="FF8036EE7435FFA04CACF915FE4FF921" box="[389,416,1710,1734]" pageId="8" pageNumber="64">
<emphasis id="B94ACC3C7435FFA04CACF915FE4FF921" box="[389,416,1710,1734]" italics="true" pageId="8" pageNumber="64">v2</emphasis>
</date>
and
<emphasis id="B94ACC3C7435FFA04CFEF915FE12F921" box="[471,509,1710,1734]" italics="true" pageId="8" pageNumber="64">sc1</emphasis>
of
<taxonomicName id="4C3E6BAD7435FFA04F0AF90BFD29F921" box="[547,710,1710,1735]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="cordidiscus">
<emphasis id="B94ACC3C7435FFA04F0AF90BFD29F921" box="[547,710,1710,1735]" italics="true" pageId="8" pageNumber="64">H. cordidiscus</emphasis>
</taxonomicName>
are located close to each other and are distinctly anterior to
<emphasis id="B94ACC3C7435FFA04846F915FA7AF921" box="[1391,1429,1710,1734]" italics="true" pageId="8" pageNumber="64">sc2</emphasis>
. A similar arrangement can only be seen in
<taxonomicName id="4C3E6BAD7435FFA04F85F96BFBCCF900" authority="(Ebermann 1982a)" baseAuthorityName="Ebermann" baseAuthorityYear="1982" box="[684,1059,1743,1768]" class="Arachnida" family="Scutacaridae" genus="Lophodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B94ACC3C7435FFA04F85F96BFCD4F90F" box="[684,827,1744,1768]" italics="true" pageId="8" pageNumber="64">Lophodispus</emphasis>
(
<bibRefCitation id="EFAF6DDF7435FFA04E79F974FBF4F900" author="Ebermann" box="[848,1051,1743,1767]" pageId="8" pageNumber="64" refString="Ebermann, E. (1982 a) Zuchtversuche und morphologische Untersuchungen an heimischen Milben (Acari, Scutacaridae). Mittteilungen des naturwissenschaftlichen Vereines fur Steiermark, 112, 155 - 165." type="journal article" year="1982" yearSuffix="a">Ebermann 1982a</bibRefCitation>
)
</taxonomicName>
. In the genera
<taxonomicName id="4C3E6BAD7435FFA049C8F96BFA8AF90F" box="[1249,1381,1744,1768]" class="Arachnida" family="Scutacaridae" genus="Archidispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B94ACC3C7435FFA049C8F96BFA8AF90F" box="[1249,1381,1744,1768]" italics="true" pageId="8" pageNumber="64">Archidispus</emphasis>
</taxonomicName>
and
<emphasis id="B94ACC3C7435FFA04DBEF949FEC6F8EE" box="[151,297,1777,1801]" italics="true" pageId="8" pageNumber="64">
<taxonomicName id="4C3E6BAD7435FFA04DBEF949FEE6F8EE" box="[151,265,1778,1801]" class="Arachnida" family="Scutacaridae" genus="Imparipes" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">Imparipes</taxonomicName>
<date id="FF8036EE7435FFA04C39F94AFEC6F8EE" box="[272,297,1777,1801]" pageId="8" pageNumber="64">v2</date>
</emphasis>
is located anterior of both
<emphasis id="B94ACC3C7435FFA04F74F94AFD6CF8EE" box="[605,643,1777,1801]" italics="true" pageId="8" pageNumber="64">sc1</emphasis>
and
<emphasis id="B94ACC3C7435FFA04F92F94AFD0EF8EE" box="[699,737,1777,1801]" italics="true" pageId="8" pageNumber="64">sc2</emphasis>
(
<bibRefCitation id="EFAF6DDF7435FFA04FC6F94AFC5AF8EE" author="Ebermann" box="[751,949,1777,1801]" pageId="8" pageNumber="64" refString="Ebermann, E. (1982 b) Fortpflanzungsbiologische Studien an Scutacariden (Acari, Trombidiformes). Zoologische Jahrbucher (Systematik), 109, 98 - 116." type="journal article" year="1982" yearSuffix="b">Ebermann 1982b</bibRefCitation>
,
<bibRefCitation id="EFAF6DDF7435FFA04EE8F94AFC13F8EE" author="Ebermann" box="[961,1020,1777,1801]" pageId="8" pageNumber="64" refString="Ebermann, E. (1994) First demonstration of males in species of the genus Archidispus Karafiat (Acari, Tarsonemina, Scutacaridae). International Journal of Acarology, 20, 3, 169 - 182." type="journal article" year="1994">1994</bibRefCitation>
;
<bibRefCitation id="EFAF6DDF7435FFA04921F94AFACBF8EE" author="Delfinado" box="[1032,1316,1777,1801]" pageId="8" pageNumber="64" refString="Delfinado, M. D. &amp; Baker, E. W. (1976) New species of Scutacaridae (Acarina) associated with insects. Acarologia, 18, 2, 264 - 301." type="journal article" year="1976">
Delfinado &amp;
<collectingCountry id="F32950BE7435FFA049B7F94AFB0EF8EE" box="[1182,1249,1777,1801]" name="United States Pacific Island Wildlife Refuges" pageId="8" pageNumber="64">Baker</collectingCountry>
1976
</bibRefCitation>
;
<bibRefCitation id="EFAF6DDF7435FFA04819F94AFE8BF8CD" author="Khaustov" pageId="8" pageNumber="64" refString="Khaustov, A. A. &amp; Chydydrov, P. R. (2004) New species of mites of the family Scutacaridae (Acari: Heterostigmata) associated with ants (Hymenoptera, Formicidae) from Turkmenistan. Acarina, 12, 2, 87 - 103." type="journal article" year="2004">Khaustov &amp; Chydyrov 2004</bibRefCitation>
) while in
<taxonomicName id="4C3E6BAD7435FFA04CFDF8A8FDBFF8CD" box="[468,592,1811,1834]" class="Arachnida" family="Scutacaridae" genus="Scutacarus" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B94ACC3C7435FFA04CFDF8A8FDBFF8CD" box="[468,592,1811,1834]" italics="true" pageId="8" pageNumber="64">Scutacarus</emphasis>
</taxonomicName>
the arrangement of these setae seems to be more variable (
<bibRefCitation id="EFAF6DDF7435FFA049C9F8A9FA78F8CD" author="Ebermann" box="[1248,1431,1810,1834]" pageId="8" pageNumber="64" refString="Ebermann, E. (1983) Zur Morphologie der Milbenart Scutacarus peractus Karafiat, 1959 (Acari, Trombidiformes). Mittteilungen des naturwissenschaftlichen Vereines fur Steiermark, 113, 91 - 98." type="journal article" year="1983">Ebermann 1983</bibRefCitation>
;
<bibRefCitation id="EFAF6DDF7435FFA04DBEF88EFE9DF8AB" author="Norton" box="[151,370,1844,1868]" pageId="8" pageNumber="64" refString="Norton, R. A. &amp; Ide, G. S. (1974) Scutacarus baculitarsus agaricus n. subsp. (Acarina: Scutacaridae) from commercial mushroom houses, with notes on phoretic behaviour. Journal of the Kansas Entomological Society, 47, 4, 527 - 534." type="journal article" year="1974">Norton &amp; Ide 1974</bibRefCitation>
). Since the numbers of known males are low in each genus, it cannot be stated yet whether the position of the dorsal setae indeed may be genus-specific.
</paragraph>
<paragraph id="8B81102E7435FFA04DEEF8CDFD2AF833" blockId="8.[151,1436,1710,2004]" pageId="8" pageNumber="64">
The setation for legs I, II and III is the same in all known males from different genera. Differences are, however, present in leg IV. The formula Tr1-Fe2-Ge1-Ti4(1)-Ta6 that is displayed in
<taxonomicName id="4C3E6BAD7435FFA0497EF821FB16F856" box="[1111,1273,1945,1969]" class="Arachnida" family="Scutacaridae" genus="Heterodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="cordidiscus">
<emphasis id="B94ACC3C7435FFA0497EF821FB16F856" box="[1111,1273,1945,1969]" italics="true" pageId="8" pageNumber="64">H. cordidiscus</emphasis>
</taxonomicName>
can otherwise only be found in
<taxonomicName id="4C3E6BAD7435FFA04C7FF807FD2EF833" authority="(Ebermann 1982a)" baseAuthorityName="Ebermann" baseAuthorityYear="1982" box="[342,705,1980,2004]" class="Arachnida" family="Scutacaridae" genus="Lophodispus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B94ACC3C7435FFA04C7FF807FE0AF833" box="[342,485,1980,2004]" italics="true" pageId="8" pageNumber="64">Lophodispus</emphasis>
(
<bibRefCitation id="EFAF6DDF7435FFA04CDDF807FD56F833" author="Ebermann" box="[500,697,1980,2004]" pageId="8" pageNumber="64" refString="Ebermann, E. (1982 a) Zuchtversuche und morphologische Untersuchungen an heimischen Milben (Acari, Scutacaridae). Mittteilungen des naturwissenschaftlichen Vereines fur Steiermark, 112, 155 - 165." type="journal article" year="1982" yearSuffix="a">Ebermann 1982a</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink