287 lines
36 KiB
XML
287 lines
36 KiB
XML
<document id="1F379AE865145679B09D93055238D7A7" ID-DOI="10.1649/0010-065X-72.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5102840" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626277672172" checkinUser="carolina" docAuthor="Mullen, Logan J., Campbell, J. M. & Sikes, Derek S." docDate="2018" docId="03971078FFEFBC64FD62F9BEFC87CB81" docLanguage="en" docName="TheColeopBull.mo.72.16.1-54.pdf" docOrigin="The Coleopterists Bulletin 72" docSource="http://dx.doi.org/10.1649/0010-065x-72.mo4.1" docStyle="DocumentStyle:A431C8F32F3D86E670C18C13DA9B5F2B.4:TheColeopBull.2000-2009.monograph" docStyleId="A431C8F32F3D86E670C18C13DA9B5F2B" docStyleName="TheColeopBull.2000-2009.monograph" docStyleVersion="4" docTitle="Phlaeopterus fusconiger Motschulsky 1853" docType="treatment" docVersion="3" lastPageNumber="33" masterDocId="FFAE6800FFF0BC44FFDAFFA6FFBDCE36" masterDocTitle="Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="32" updateTime="1699007718751" updateUser="plazi">
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<mods:title id="60103535332C7ABEF5BF6862C95AD462">Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="4D730283F802C690BD176419DE5F01A3">Mullen, Logan J.</mods:namePart>
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<mods:namePart id="90121D0BF6F1AC084469E6446A6609AE">Campbell, J. M.</mods:namePart>
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<mods:namePart id="843D3705263EFB81D34249F744B2E273">Sikes, Derek S.</mods:namePart>
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<mods:title id="39E06FB3C9153622CD2AC1AEEF5A5831">The Coleopterists Bulletin</mods:title>
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<mods:part id="EE30CD80C356E7581770717D9952AB9A">
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<mods:date id="4B6EB52E9F185524EC31021B48F11EC7">2018</mods:date>
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<mods:number id="6A0FDB278925D06935C9592E7085A724">2018-12-28</mods:number>
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<mods:number id="D8DD29C028BBE11D807DD75AC05115D1">72</mods:number>
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<paragraph id="8B81A16EFFEFBC5BFD62F9BEFB34C81B" blockId="31.[696,1161,1560,1608]" box="[696,1161,1560,1581]" pageId="31" pageNumber="32">
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<heading id="D0C91602FFEFBC5BFD62F9BEFB34C81B" box="[696,1161,1560,1581]" centered="true" fontSize="10" level="2" pageId="31" pageNumber="32" reason="2">
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<emphasis id="B94A7D7CFFEFBC5BFD62F9BEFB34C81B" bold="true" box="[696,1161,1560,1581]" pageId="31" pageNumber="32">
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8.
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<taxonomicName id="4C3EDAEDFFEFBC5BFD08F9BEFB34C81B" ID-CoL="76ZZY" authority="Motschulsky, 1853" authorityName="Motschulsky" authorityYear="1853" box="[722,1161,1560,1581]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="fusconiger">
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<emphasis id="B94A7D7CFFEFBC5BFD08F9BEFC79C81B" bold="true" box="[722,964,1560,1581]" italics="true" pageId="31" pageNumber="32">Phlaeopterus fusconiger</emphasis>
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<bibRefCitation id="EFAFDC9FFFEFBC5BFC16F9BEFB34C81B" author="Motschulsky" box="[972,1161,1560,1581]" pageId="31" pageNumber="32" pagination="77 - 80" refId="ref29403" refString="Motschulsky, V. I. 1853. Nouveaut'es. Etudes Entomologiques 1: 77 - 80." type="journal article" year="1853">Motschulsky, 1853</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C324F2E5FFEFBC5BFCD3F995FB84C87E" box="[777,1081,1587,1608]" pageId="31" pageNumber="32" type="description">
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<paragraph id="8B81A16EFFEFBC5BFCD3F995FB84C87E" blockId="31.[696,1161,1560,1608]" box="[777,1081,1587,1608]" pageId="31" pageNumber="32">
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(
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<figureCitation id="1305BDEBFFEFBC5BFCCAF995FCDBC87E" box="[784,870,1587,1608]" captionStart="Fig" captionStartId="11.[175,206,1726,1745]" captionTargetBox="[156,1167,196,1708]" captionTargetId="figure-1@11.[154,1169,194,1710]" captionTargetPageId="11" captionText="Fig. 2. Phlaeopterus species, habitus. A) P. elongatus, male (UAMObs:Ento:232543), B) P. filicornis, female" figureDoi="http://doi.org/10.5281/zenodo.5102845" httpUri="https://zenodo.org/record/5102845/files/figure.png" pageId="31" pageNumber="32">Figs. 2D</figureCitation>
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,
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||
<figureCitation id="1305BDEBFFEFBC5BFCA8F995FC32C87E" box="[882,911,1587,1608]" captionStart="Fig" captionStartId="17.[175,206,1027,1046]" captionText="Fig. 9. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. frosti, B) P. fusconiger. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." pageId="31" pageNumber="32">9B</figureCitation>
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,
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<figureCitation id="1305BDEBFFEFBC5BFC41F995FC7BC87E" box="[923,966,1587,1608]" captionStart="Fig" captionStartId="23.[175,206,1547,1566]" captionTargetBox="[168,1155,197,1527]" captionTargetId="figure-55@23.[164,1159,194,1532]" captionTargetPageId="23" captionText="Fig. 17. Phlaeopterus species. Pronota, dorsal view, of A) P. fusconiger, B) P. frosti, C) P. filicornis, D) P. hatchi, E) P. elongatus. Abdominal segments 4–6, dorsal view, of F) P. houkae, G) P. obsoletus, H) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102867" httpUri="https://zenodo.org/record/5102867/files/figure.png" pageId="31" pageNumber="32">17A</figureCitation>
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,
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<figureCitation id="1305BDEBFFEFBC5BFC08F995FC41C87E" box="[978,1020,1587,1608]" captionStart="Fig" captionStartId="24.[197,228,482,501]" captionTargetBox="[177,1188,196,465]" captionTargetId="figure-576@24.[175,1190,194,467]" captionTargetPageId="24" captionText="Fig. 18. Mesotibiae, dorsal view, of A) Phlaeopterus frosti (UAMObs:Ento:235497), and B) Phlaeopterus fusconiger (UAMObs:Ento:235751). Red highlights are added to emphasize the difference in the length of the subglabrous region (lacking dense pubescence but with longer spine-like setae) at the apex of the tibia." figureDoi="http://doi.org/10.5281/zenodo.5102869" httpUri="https://zenodo.org/record/5102869/files/figure.png" pageId="31" pageNumber="32">18B</figureCitation>
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,
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<figureCitation id="1305BDEBFFEFBC5BFBD2F995FB8CC87E" box="[1032,1073,1587,1608]" captionStart="Fig" captionStartId="29.[175,206,1486,1505]" captionTargetBox="[156,1167,196,1468]" captionTargetId="figure-90@29.[154,1169,194,1470]" captionTargetPageId="29" captionText="Fig. 24. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. occidentalis, B) P. olympicus, C) P. loganensis, D) P. fusconiger." figureDoi="http://doi.org/10.5281/zenodo.5102879" httpUri="https://zenodo.org/record/5102879/files/figure.png" pageId="31" pageNumber="32">24D</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C324F2E5FFEFBC64FD74F9FBFE82CF6D" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="reference_group">
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<paragraph id="8B81A16EFFEFBC5BFD74F9FBFB29C8F4" blockId="31.[686,1172,1629,1730]" pageId="31" pageNumber="32">
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<treatmentCitationGroup id="AB2E8640FFEFBC5BFD74F9FBFB29C8F4" pageId="31" pageNumber="32">
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<treatmentCitation id="0A9F877FFFEFBC5BFD74F9FBFB29C844" author="Motschulsky" box="[686,1172,1629,1650]" page="78" pageId="31" pageNumber="32" year="1853">
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<taxonomicName id="4C3EDAEDFFEFBC5BFD74F9FBFB29C844" ID-CoL="76ZZY" authority="Motschulsky 1853: 78" authorityName="Motschulsky" authorityPageNumber="78" authorityYear="1853" box="[686,1172,1629,1650]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="fusconiger">
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<emphasis id="B94A7D7CFFEFBC5BFD74F9FBFC22C844" box="[686,927,1629,1650]" italics="true" pageId="31" pageNumber="32">Phlaeopterus fusconiger</emphasis>
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<bibRefCitation id="EFAFDC9FFFEFBC5BFC77F9FBFB29C844" author="Motschulsky" box="[941,1172,1629,1650]" pageId="31" pageNumber="32" pagination="77 - 80" refId="ref29403" refString="Motschulsky, V. I. 1853. Nouveaut'es. Etudes Entomologiques 1: 77 - 80." type="journal article" year="1853">Motschulsky 1853: 78</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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[original description].
|
||
<treatmentCitation id="0A9F877FFFEFBC5BFC7BF9DEFBF4C8BB" author="Casey" box="[929,1097,1656,1677]" page="320" pageId="31" pageNumber="32" year="1885">
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<bibRefCitation id="EFAFDC9FFFEFBC5BFC7BF9DEFBF4C8BB" author="Casey" box="[929,1097,1656,1677]" pageId="31" pageNumber="32" pagination="281 - 606" refId="ref28625" refString="Casey, T. L. 1885. New genera and species of Californian Coleoptera. Bulletin of the California Academy of Sciences 7: 281 - 606." type="journal article" year="1885">Casey 1885: 320</bibRefCitation>
|
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</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A9F877FFFEFBC5BFB8FF9DEFCC9C891" author="Blackwelder" page="303" pageId="31" pageNumber="32" year="1952">
|
||
<bibRefCitation id="EFAFDC9FFFEFBC5BFB8FF9DEFCC9C891" author="Blackwelder" pageId="31" pageNumber="32" pagination="1 - 483" refId="ref28304" refString="Blackwelder, R. E. 1952. The generic names of the beetle family Staphylinidae, with an essay on genotypy. Bulletin of the United States National Museum 200: 1 - 483." type="journal article" year="1952">Blackwelder 1952: 303</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A9F877FFFEFBC5BFC5AF934FBA3C891" author="Moore" box="[896,1054,1682,1703]" page="52" pageId="31" pageNumber="32" year="1966">
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||
<bibRefCitation id="EFAFDC9FFFEFBC5BFC5AF934FBA3C891" author="Moore" box="[896,1054,1682,1703]" pageId="31" pageNumber="32" pagination="47 - 56" refId="ref29235" refString="Moore, I. 1966. Notes on the Nearctic Anthophagini with a key to the genera (Coleoptera: Staphylinidae). The Coleopterists Bulletin 20 (2): 47 - 56." type="journal article" year="1966">Moore 1966: 52</bibRefCitation>
|
||
</treatmentCitation>
|
||
; Moore and Legner 1979: 193; Campbell and Davies 1991: 5;
|
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</treatmentCitationGroup>
|
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</paragraph>
|
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<paragraph id="8B81A16EFFD0BC64FF1DFF64FE4ACEC7" blockId="32.[173,659,194,347]" pageId="32" pageNumber="33">
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<bibRefCitation id="EFAFDC9FFFD0BC64FF1DFF64FE38CEE1" author="Herman" box="[199,389,194,215]" pageId="32" pageNumber="33" pagination="1 - 650" refId="ref28893" refString="Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta, Coleoptera): 1758 to the end of the Second Millennium. I. Introduction, history, biogeographical sketches, and Omaliine group. Bulletin of the American Museum of Natural History 265: 1 - 650." type="journal article" year="2001">Herman 2001: 366</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAFDC9FFFD0BC64FE48FF64FD33CEE1" author="Bousquet" box="[402,654,194,215]" pageId="32" pageNumber="33" refId="ref28340" refString="Bousquet, Y., P. Bouchard, A. E. Davies, and D. S. Sikes. 2013. Checklist of Beetles (Coleoptera) of Canada and Alaska. Second Edition. Pensoft, Sofia, Bulgaria." type="book" year="2013">
|
||
Bousquet
|
||
<emphasis id="B94A7D7CFFD0BC64FE2DFF64FD99CEE1" box="[503,548,194,215]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
|
||
. 2013: 89
|
||
</bibRefCitation>
|
||
; Shavrin and Mullen 2015: 121.
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFD0BC64FF77FF51FE4ACF10" blockId="32.[173,659,194,347]" pageId="32" pageNumber="33">
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<treatmentCitationGroup id="AB2E8640FFD0BC64FF77FF51FE4ACF10" pageId="32" pageNumber="33">
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<treatmentCitation id="0A9F877FFFD0BC64FF77FF51FD8ECF3A" author="Maklin & [New species and notes & In & Dritter Nachtrag zur & Amerikanischen Laender des Russischen Reiches & Mannerheim & Bulletin de la Societe Imperiale des Naturalistes de Moscou" box="[173,563,247,268]" page="193" pageId="32" pageNumber="33" year="1853">
|
||
<taxonomicName id="4C3EDAEDFFD0BC64FF77FF51FD8ECF3A" authority="Maklin 1853: 193" authorityName="Maklin" authorityPageNumber="193" authorityYear="1853" box="[173,563,247,268]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fusconigra">
|
||
<emphasis id="B94A7D7CFFD0BC64FF77FF51FED4CF3A" box="[173,361,247,268]" italics="true" pageId="32" pageNumber="33">Lesteva fusconigra</emphasis>
|
||
<bibRefCitation id="EFAFDC9FFFD0BC64FEA2FF51FD8ECF3A" author="Maklin & [New species and notes & In & Dritter Nachtrag zur & Amerikanischen Laender des Russischen Reiches & Mannerheim & Bulletin de la Societe Imperiale des Naturalistes de Moscou" box="[376,563,247,268]" pageId="32" pageNumber="33" pagination="95 - 273" refId="ref29178" refString="Maklin, F. G. 1853. [New species and notes]. In: Dritter Nachtrag zur Kaefer-Fauna der Nord- Amerikanischen Laender des Russischen Reiches (C. von Mannerheim, editor). Bulletin de la Societe Imperiale des Naturalistes de Moscou 26 (3 & 4): 95 - 273." type="journal article" year="1853">Mäklin 1853: 193</bibRefCitation>
|
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</taxonomicName>
|
||
</treatmentCitation>
|
||
[original description].
|
||
<treatmentCitation id="0A9F877FFFD0BC64FE9CFEB7FE4ECF10" author="Fauvel" box="[326,499,273,294]" page="248" pageId="32" pageNumber="33" year="1878">
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<bibRefCitation id="EFAFDC9FFFD0BC64FE9CFEB7FE4ECF10" author="Fauvel" box="[326,499,273,294]" pageId="32" pageNumber="33" pagination="167 - 269" refId="ref28777" refString="Fauvel, A. 1878. Les Staphylinides de l' Am'erique du nord. Bulletin de la Soci'et'e Linn'eenne de Normandie 3 (2): 167 - 269." type="journal article" year="1878">Fauvel 1878: 248</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFD0BC64FF77FE8AFE82CF6D" blockId="32.[173,659,194,347]" pageId="32" pageNumber="33">
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<treatmentCitationGroup id="AB2E8640FFD0BC64FF77FE8AFE82CF6D" pageId="32" pageNumber="33">
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<treatmentCitation id="0A9F877FFFD0BC64FF77FE8AFD91CF77" author="Casey" box="[173,556,300,321]" page="402" pageId="32" pageNumber="33" year="1893">
|
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<taxonomicName id="4C3EDAEDFFD0BC64FF77FE8AFD91CF77" authority="Casey 1893: 402" authorityName="Casey" authorityPageNumber="402" authorityYear="1893" box="[173,556,300,321]" class="Insecta" family="Staphylinidae" genus="Tilea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fusconigra">
|
||
<emphasis id="B94A7D7CFFD0BC64FF77FE8AFEE5CF77" box="[173,344,300,321]" italics="true" pageId="32" pageNumber="33">Tilea fusconigra</emphasis>
|
||
<bibRefCitation id="EFAFDC9FFFD0BC64FEB7FE8AFD91CF77" author="Casey" box="[365,556,300,321]" pageId="32" pageNumber="33" pagination="281 - 624" refId="ref28683" refString="Casey, T. L. 1893. Coleopterological Notices V. Annals of the New York Academy of Sciences 7 (1): 281 - 624." type="journal article" year="1893">Casey 1893: 402</bibRefCitation>
|
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</taxonomicName>
|
||
</treatmentCitation>
|
||
[original description].
|
||
</treatmentCitationGroup>
|
||
</paragraph>
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||
</subSubSection>
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<paragraph id="8B81A16EFFD0BC64FF1EFEDAFD37CFA7" blockId="32.[173,660,380,1730]" box="[196,650,380,401]" pageId="32" pageNumber="33">
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<materialsCitation id="3B56AB33FFD0BC64FF1EFEDAFD3ACFA7" box="[196,647,380,401]" country="United States of America" location="Unalaska Island" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Alaska" typeStatus="holotype">
|
||
<emphasis id="B94A7D7CFFD0BC64FF1EFEDAFEEBCFA7" bold="true" box="[196,342,380,401]" pageId="32" pageNumber="33">
|
||
<typeStatus id="54851FCCFFD0BC64FF1EFEDAFF4BCFA7" box="[196,246,380,401]" pageId="32" pageNumber="33">Type</typeStatus>
|
||
Locality.
|
||
</emphasis>
|
||
<location id="8EE1F7B5FFD0BC64FE87FEDAFE44CFA7" LSID="urn:lsid:plazi:treatment:03971078FFEFBC64FD62F9BEFC87CB81:8EE1F7B5FFD0BC64FE87FEDAFE44CFA7" box="[349,505,380,401]" country="United States of America" name="Unalaska Island" pageId="32" pageNumber="33" stateProvince="Alaska">Unalaska Island</location>
|
||
,
|
||
<collectingRegion id="49FA6F8CFFD0BC64FDDFFEDAFDF4CFA7" box="[517,585,380,401]" country="United States of America" name="Alaska" pageId="32" pageNumber="33">Alaska</collectingRegion>
|
||
,
|
||
<collectingCountry id="F329E1FEFFD0BC64FD8EFEDAFD3ACFA7" box="[596,647,380,401]" name="United States of America" pageId="32" pageNumber="33">USA</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD0BC64FF1EFE30FE4DC8BB" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8B81A16EFFD0BC64FF1EFE30FE4DC8BB" blockId="32.[173,660,380,1730]" pageId="32" pageNumber="33">
|
||
<emphasis id="B94A7D7CFFD0BC64FF1EFE30FE07CF9D" bold="true" box="[196,442,406,427]" pageId="32" pageNumber="33">Redescription. Habitus:</emphasis>
|
||
Length
|
||
<quantity id="4CC60C8BFFD0BC64FDCFFE30FD2DCF9D" box="[533,656,406,428]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.2" metricValueMin="5.8" pageId="32" pageNumber="33" unit="mm" value="6.5" valueMax="7.2" valueMin="5.8">5.8–7.2 mm</quantity>
|
||
. Dark brown to reddish brown; legs, elytral epipleura, and tarsi often lighter; apices of tibiae, tarsi, and palpi even lighter reddish brown; elytra usually slightly lighter than pronotum (
|
||
<figureCitation id="1305BDEBFFD0BC64FE36FDA6FD83CC23" box="[492,574,512,533]" captionStart="Fig" captionStartId="11.[175,206,1726,1745]" captionTargetBox="[156,1167,196,1708]" captionTargetId="figure-1@11.[154,1169,194,1710]" captionTargetPageId="11" captionText="Fig. 2. Phlaeopterus species, habitus. A) P. elongatus, male (UAMObs:Ento:232543), B) P. filicornis, female" figureDoi="http://doi.org/10.5281/zenodo.5102845" httpUri="https://zenodo.org/record/5102845/files/figure.png" pageId="32" pageNumber="33">Fig. 2D</figureCitation>
|
||
).
|
||
<emphasis id="B94A7D7CFFD0BC64FD8FFDA6FD2FCC23" bold="true" box="[597,658,512,533]" pageId="32" pageNumber="33">Head:</emphasis>
|
||
Slightly broader than long, ratio of width across eyes to head length = 4:3. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5–10 at least 2 times longer than wide; antennomeres 4–11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3
|
||
<superScript id="7C4B0C26FFD0BC64FD5EFCB9FD2ECD1B" attach="none" box="[644,659,799,813]" fontSize="6" pageId="32" pageNumber="33">rd</superScript>
|
||
palpomere 1.6–1.8 times longer than palpomere 2.
|
||
<emphasis id="B94A7D7CFFD0BC64FF77FCFCFF42CD59" bold="true" box="[173,255,858,879]" pageId="32" pageNumber="33">Thorax:</emphasis>
|
||
Pronotum moderately wide (
|
||
<figureCitation id="1305BDEBFFD0BC64FDF0FCFCFD35CD59" box="[554,648,858,879]" captionStart="Fig" captionStartId="23.[175,206,1547,1566]" captionTargetBox="[168,1155,197,1527]" captionTargetId="figure-55@23.[164,1159,194,1532]" captionTargetPageId="23" captionText="Fig. 17. Phlaeopterus species. Pronota, dorsal view, of A) P. fusconiger, B) P. frosti, C) P. filicornis, D) P. hatchi, E) P. elongatus. Abdominal segments 4–6, dorsal view, of F) P. houkae, G) P. obsoletus, H) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102867" httpUri="https://zenodo.org/record/5102867/files/figure.png" pageId="32" pageNumber="33">Fig. 17A</figureCitation>
|
||
), length to width ratio = 0.68–0.72; ratio of pronotal width to head width = 1.54–1.67; maximum width subequal to elytral width at humeral angles; punctures separated by distance equal to slightly greater than diameter of a puncture; lateral margins explanate posterad but not anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.1–2.2 times longer than pronotum; apical margins convex or subtruncate. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth, tooth more strongly notched on posterior margin than on anterior margin; longitudinal carina along midline of mesosternum distinct on anterior half, reduced on posterior half.
|
||
<emphasis id="B94A7D7CFFD0BC64FEE1FAA5FECFCB2E" bold="true" box="[315,370,1283,1304]" pageId="32" pageNumber="33">Legs:</emphasis>
|
||
Apices of all tibiae glabrous, subglabrous apex of mesotibia subequal to length of basal 3 mesotarsomeres (
|
||
<figureCitation id="1305BDEBFFD0BC64FE78FA9EFE47CB7B" box="[418,506,1336,1357]" captionStart="Fig" captionStartId="24.[197,228,482,501]" captionTargetBox="[177,1188,196,465]" captionTargetId="figure-576@24.[175,1190,194,467]" captionTargetPageId="24" captionText="Fig. 18. Mesotibiae, dorsal view, of A) Phlaeopterus frosti (UAMObs:Ento:235497), and B) Phlaeopterus fusconiger (UAMObs:Ento:235751). Red highlights are added to emphasize the difference in the length of the subglabrous region (lacking dense pubescence but with longer spine-like setae) at the apex of the tibia." figureDoi="http://doi.org/10.5281/zenodo.5102869" httpUri="https://zenodo.org/record/5102869/files/figure.png" pageId="32" pageNumber="33">Fig. 18B</figureCitation>
|
||
), length of glabrous apical region of metatibia as ratio to metatibial length in males = 7.4–12.2, in females = 6.6–8.6. Metatrochanter without tooth on apical margin.
|
||
<emphasis id="B94A7D7CFFD0BC64FF77FA05FEAACB8E" bold="true" box="[173,279,1443,1464]" pageId="32" pageNumber="33">Abdomen:</emphasis>
|
||
Wing-folding spicules on tergites IV and V; shape of wing-folding patches on tergite V broadly oval and narrowly separate.
|
||
<emphasis id="B94A7D7CFFD0BC64FDF3FA7EFD2ECBDB" bold="true" box="[553,659,1496,1517]" pageId="32" pageNumber="33">Aedeagus:</emphasis>
|
||
Length
|
||
<quantity id="4CC60C8BFFD0BC64FF24FA55FE2DC83E" box="[254,400,1522,1544]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" metricValueMax="1.43" metricValueMin="1.23" pageId="32" pageNumber="33" unit="mm" value="1.33" valueMax="1.43" valueMin="1.23">1.23–1.43 mm</quantity>
|
||
long. Median lobe with margins concavely constricted near midline (
|
||
<figureCitation id="1305BDEBFFD0BC64FDB7F9ABFF67C80B" captionStart="Fig" captionStartId="29.[175,206,1486,1505]" captionTargetBox="[156,1167,196,1468]" captionTargetId="figure-90@29.[154,1169,194,1470]" captionTargetPageId="29" captionText="Fig. 24. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. occidentalis, B) P. olympicus, C) P. loganensis, D) P. fusconiger." figureDoi="http://doi.org/10.5281/zenodo.5102879" httpUri="https://zenodo.org/record/5102879/files/figure.png" pageId="32" pageNumber="33">Fig. 24D</figureCitation>
|
||
). Parameres convex, converging past apex of median lobe. Internal sac rectangular; lightly sclerotized, uniformly covered with microspinules; lacking subapical transverse fold.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD0BC64FF1FF934FC62CC7C" pageId="32" pageNumber="33" type="materials_examined">
|
||
<paragraph id="8B81A16EFFD0BC64FF1FF934FBD4CF3A" blockId="32.[173,660,380,1730]" lastBlockId="32.[707,1194,194,1463]" pageId="32" pageNumber="33">
|
||
<emphasis id="B94A7D7CFFD0BC64FF1FF934FED7C891" bold="true" box="[197,362,1682,1703]" pageId="32" pageNumber="33">Type Specimen.</emphasis>
|
||
<typeStatus id="54851FCCFFD0BC64FEACF934FE73C891" box="[374,462,1682,1703]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
|
||
female (UAMObs: Ento:235794) labeled as follows: Unalaschka,
|
||
<taxonomicName id="4C3EDAEDFFD0BC64FD19FF64FB14CEE1" authority="Motsch., Unalaschka" authorityName="Motsch., Unalaschka" box="[707,1193,194,215]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fusconiger">Phlaeopterus fusconiger Motsch., Unalaschka</taxonomicName>
|
||
[Unalaska Island]/
|
||
<taxonomicName id="4C3EDAEDFFD0BC64FCAFFF7AFB1BCEC7" authority="Mots." authorityName="Motschulsky" authorityYear="1853" box="[885,1190,220,241]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fusconiger">Phlaeopterus fusconiger Mots.</taxonomicName>
|
||
,
|
||
<typeStatus id="54851FCCFFD0BC64FD19FF51FCFBCF3A" box="[707,838,247,268]" pageId="32" pageNumber="33" type="holotype">HOLOTYPE</typeStatus>
|
||
, exam. 1984 J.M. Campbell.
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFD0BC64FD01FEB7FC62CC7C" blockId="32.[707,1194,194,1463]" pageId="32" pageNumber="33">
|
||
<materialsCitation id="3B56AB33FFD0BC64FD01FEB7FC67CC7C" collectionCode="V" collectorName="Shavrin & A. Shavrin & Although" pageId="32" pageNumber="33" specimenCount="4" typeStatus="holotype">
|
||
As of 1984, the
|
||
<typeStatus id="54851FCCFFD0BC64FCA8FEB7FC78CF10" box="[882,965,273,294]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
was confirmed by JMC to be located in the collection of the ZMM. The specimen is glued to a transparent plate and badly damaged with both elytra and apical five antennomeres of the right antenna, right middle and hind legs, and apical four metatarsomeres missing. In 2015, Alexey
|
||
<collectionCode id="ED2F39ABFFD0BC64FC8DFE17FCD5CFF0" box="[855,872,433,454]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="32" pageNumber="33" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="26CBC4B8FFD0BC64FCA3FE17FC7ECFF0" box="[889,963,433,454]" pageId="32" pageNumber="33">Shavrin</collectorName>
|
||
(
|
||
<collectorName id="26CBC4B8FFD0BC64FC0DFE17FBF5CFF0" box="[983,1096,433,454]" pageId="32" pageNumber="33">A. Shavrin</collectorName>
|
||
, personal communication) visited the collection and could not locate the
|
||
<typeStatus id="54851FCCFFD0BC64FCEDFE40FC30CFCD" box="[823,909,486,507]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
.
|
||
<collectorName id="26CBC4B8FFD0BC64FC7AFE40FC47CFCD" box="[928,1018,486,507]" pageId="32" pageNumber="33">Although</collectorName>
|
||
the
|
||
<typeStatus id="54851FCCFFD0BC64FBECFE40FB34CFCD" box="[1078,1161,486,507]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
is possibly missing, this species is currently in no danger of being confused with similar species, thus a
|
||
<typeStatus id="54851FCCFFD0BC64FCDEFD93FCF2CC7C" box="[772,847,565,586]" pageId="32" pageNumber="33" type="neotype">neotype</typeStatus>
|
||
is not needed
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD0BC64FD01FDF6FB17CD59" pageId="32" pageNumber="33" type="distribution">
|
||
<paragraph id="8B81A16EFFD0BC64FD01FDF6FB3CCCDC" blockId="32.[707,1194,194,1463]" pageId="32" pageNumber="33">
|
||
<emphasis id="B94A7D7CFFD0BC64FD01FDF6FCE1CC53" bold="true" box="[731,860,592,613]" pageId="32" pageNumber="33">Distribution.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFD0BC64FCB8FDF6FBF6CC53" authorityName="Motschulsky" authorityYear="1853" box="[866,1099,592,613]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fusconiger">
|
||
<emphasis id="B94A7D7CFFD0BC64FCB8FDF6FBF6CC53" box="[866,1099,592,613]" italics="true" pageId="32" pageNumber="33">Phlaeopterus fusconiger</emphasis>
|
||
</taxonomicName>
|
||
is widely distributed along the northwest coast of North America, from the Olympic Mountains of
|
||
<collectingRegion id="49FA6F8CFFD0BC64FBB7FD23FCBDCC83" country="United States of America" name="Washington" pageId="32" pageNumber="33">Washington</collectingRegion>
|
||
north through the Pacific Coast Range, on Vancouver Island and the Haida Gwaii, to the Aleutian Islands and Nome,
|
||
<collectingRegion id="49FA6F8CFFD0BC64FC02FD73FBA7CCDC" box="[984,1050,725,746]" country="United States of America" name="Alaska" pageId="32" pageNumber="33">Alaska</collectingRegion>
|
||
(
|
||
<figureCitation id="1305BDEBFFD0BC64FBF3FD73FBC9CCDC" box="[1065,1140,725,746]" captionStart="Fig" captionStartId="17.[175,206,1027,1046]" captionText="Fig. 9. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. frosti, B) P. fusconiger. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." pageId="32" pageNumber="33">Fig. 9B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFD0BC64FD01FD56FB17CD59" blockId="32.[707,1194,194,1463]" pageId="32" pageNumber="33">
|
||
<emphasis id="B94A7D7CFFD0BC64FD01FD56FCF7CD33" bold="true" box="[731,842,752,773]" pageId="32" pageNumber="33">Bionomics.</emphasis>
|
||
Adults have been collected at near sea level in
|
||
<collectingRegion id="49FA6F8CFFD0BC64FC9FFCACFC39CD29" box="[837,900,778,799]" country="United States of America" name="Alaska" pageId="32" pageNumber="33">Alaska</collectingRegion>
|
||
up to
|
||
<quantity id="4CC60C8BFFD0BC64FC0FFCACFB95CD29" box="[981,1064,778,799]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="m" value="2000.0">2,000 m</quantity>
|
||
in mainland
|
||
<collectingRegion id="49FA6F8CFFD0BC64FD19FC83FC8DCD0C" box="[707,816,805,826]" country="United States of America" name="Washington" pageId="32" pageNumber="33">Washington</collectingRegion>
|
||
and
|
||
<collectingRegion id="49FA6F8CFFD0BC64FCABFC83FBA5CD0C" box="[881,1048,805,826]" country="Canada" name="British Columbia" pageId="32" pageNumber="33">British Columbia</collectingRegion>
|
||
during June– August.The Nome record (UAMObs:Ento:234463) is the northernmost record for any
|
||
<taxonomicName id="4C3EDAEDFFD0BC64FC3EFCFCFB17CD59" box="[996,1194,858,879]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFD0BC64FC3EFCFCFBE0CD59" box="[996,1117,858,879]" italics="true" pageId="32" pageNumber="33">Phlaeopterus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD0BC64FD01FCD2FC87CB81" pageId="32" pageNumber="33" type="discussion">
|
||
<paragraph id="8B81A16EFFD0BC64FD01FCD2FC87CB81" blockId="32.[707,1194,194,1463]" pageId="32" pageNumber="33">
|
||
<emphasis id="B94A7D7CFFD0BC64FD01FCD2FC80CDBF" bold="true" box="[731,829,884,905]" pageId="32" pageNumber="33">Remarks.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFD0BC64FC9CFCD2FB8FCDBF" authorityName="Motschulsky" authorityYear="1853" box="[838,1074,884,905]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fusconiger">
|
||
<emphasis id="B94A7D7CFFD0BC64FC9CFCD2FB8FCDBF" box="[838,1074,884,905]" italics="true" pageId="32" pageNumber="33">Phlaeopterus fusconiger</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all other
|
||
<taxonomicName id="4C3EDAEDFFD0BC64FC67FC29FB35CD92" box="[957,1160,911,932]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFD0BC64FC67FC29FB86CD92" box="[957,1083,911,932]" italics="true" pageId="32" pageNumber="33">Phlaeopterus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by the shape of the pronotum, the glabrous region at the apex of the tibiae, and the aedeagus. This species is very similar to
|
||
<taxonomicName id="4C3EDAEDFFD0BC64FC8EFC79FC20CDC2" authorityName="Hatch" authorityYear="1957" box="[852,925,991,1012]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="frosti">
|
||
<emphasis id="B94A7D7CFFD0BC64FC8EFC79FC20CDC2" box="[852,925,991,1012]" italics="true" pageId="32" pageNumber="33">P. frosti</emphasis>
|
||
</taxonomicName>
|
||
but can be distinguished by the shorter glabrous region of the mesotibia, slightly lighter color, more strongly developed anterior margins of the mesosternal notch, and the aedeagus with the internal sac shorter, broader, less heavily sclerotized, and with the apex less acutely narrowed. These two species are sympatric in the coastal range from the Olympic Mountains of
|
||
<collectingRegion id="49FA6F8CFFD0BC64FC27FB3FFBCDCA98" box="[1021,1136,1177,1198]" country="United States of America" name="Washington" pageId="32" pageNumber="33">Washington</collectingRegion>
|
||
north to the Kenai Mountains of
|
||
<collectingRegion id="49FA6F8CFFD0BC64FC0AFB15FBABCAFE" box="[976,1046,1203,1224]" country="United States of America" name="Alaska" pageId="32" pageNumber="33">Alaska</collectingRegion>
|
||
. We have confirmed the surprising occurrence of
|
||
<taxonomicName id="4C3EDAEDFFD0BC64FBCFFB68FB2CCAD5" authorityName="Motschulsky" authorityYear="1853" box="[1045,1169,1230,1251]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fusconiger">
|
||
<emphasis id="B94A7D7CFFD0BC64FBCFFB68FB2CCAD5" box="[1045,1169,1230,1251]" italics="true" pageId="32" pageNumber="33">P. fusconiger</emphasis>
|
||
</taxonomicName>
|
||
in Nome,
|
||
<collectingRegion id="49FA6F8CFFD0BC64FCD2FB4EFCF6CACB" box="[776,843,1256,1277]" country="United States of America" name="Alaska" pageId="32" pageNumber="33">Alaska</collectingRegion>
|
||
by examination of a single specimen collected in Nome by George Ball in 1958 (UAMObs:Ento:234463). However, we are aware of no additional specimens of
|
||
<taxonomicName id="4C3EDAEDFFD0BC64FC2BFA9EFBCFCB7B" authorityName="Motschulsky" authorityYear="1853" box="[1009,1138,1336,1357]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94A7D7CFFD0BC64FC2BFA9EFBCFCB7B" box="[1009,1138,1336,1357]" italics="true" pageId="32" pageNumber="33">Phlaeopterus</emphasis>
|
||
</taxonomicName>
|
||
from Nome, or anywhere on the Seward Peninsula. This remote population would be interesting to confirm and compare genetically to other populations of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |