137 lines
20 KiB
XML
137 lines
20 KiB
XML
<document id="B9C0C7EE43B89E507CFD1209A0755CB0" ID-CLB-Dataset="50790" ID-DOI="10.5281/zenodo.172376" ID-GBIF-Dataset="54e48000-e28d-42b8-993e-d39b4923683f" ID-ISSN="11755326" ID-Zenodo-Dep="172376" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459755737607" checkinUser="plazi" docAuthor="Johanson, Kjell Arne & Oláh, János" docDate="2006" docId="03F512008345FFF2FED2E11BFCEEAC8F" docLanguage="en" docName="zt01205p030.pdf" docOrigin="Zootaxa 1205" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Helicopsyche (Petrotrichia) sahadika Johanson & Oláh, 2006, new species" docType="treatment" docVersion="11" lastPageNumber="8" masterDocId="FFCC6A788340FFF5FFDAE437FFE5AE11" masterDocTitle="Eleven new species of Sericostomatoidea from Madagascar (Trichoptera: Helicopsychidae, Petrothrincidae, Sericostomatidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="6" updateTime="1698217669661" updateUser="ExternalLinkService">
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<mods:title id="7C38BF5DF3C348605F46AD45DA691C88">Eleven new species of Sericostomatoidea from Madagascar (Trichoptera: Helicopsychidae, Petrothrincidae, Sericostomatidae)</mods:title>
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<mods:namePart id="6C17E3B8DC5B1432E1A0244BCAA00D2A">Johanson, Kjell Arne</mods:namePart>
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<mods:namePart id="A8060EAE852297711AFF8C7EA0907593">Oláh, János</mods:namePart>
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<mods:title id="B277225DB6AC53C351D3AB0BFB04364B">Zootaxa</mods:title>
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<mods:part id="201083C5E33D1ED23E736FA151E0523D">
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<mods:date id="6E5E9ED5142F7FC6569BDA34EF291140">2006</mods:date>
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<treatment id="03F512008345FFF2FED2E11BFCEEAC8F" ID-DOI="http://doi.org/10.5281/zenodo.5049120" ID-GBIF-Taxon="119343617" ID-Zenodo-Dep="5049120" LSID="urn:lsid:plazi:treatment:03F512008345FFF2FED2E11BFCEEAC8F" httpUri="http://treatment.plazi.org/id/03F512008345FFF2FED2E11BFCEEAC8F" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
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<subSubSection id="C346F09D8345FFF0FED2E11BFE42AB78" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="8BE3A3168345FFF0FED2E11BFC8DAB57" blockId="5.[264,872,1324,1385]" box="[264,872,1324,1350]" pageId="5" pageNumber="6">
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<heading id="D0AB147A8345FFF0FED2E11BFC8DAB57" bold="true" box="[264,872,1324,1350]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
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<emphasis id="B9287F048345FFF0FED2E11BFC8DAB57" bold="true" box="[264,872,1324,1350]" pageId="5" pageNumber="6">
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<taxonomicName id="4C5CD8958345FFF0FED2E11BFD2CAB57" box="[264,713,1324,1350]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sahadika" status="sp. nov." subGenus="Petrotrichia">
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<emphasis id="B9287F048345FFF0FED2E11BFD2CAB57" bold="true" box="[264,713,1324,1350]" italics="true" pageId="5" pageNumber="6">Helicopsyche (Petrotrichia) sahadika</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21BC27F8345FFF0FD02E11BFC8DAB57" box="[728,872,1324,1350]" pageId="5" pageNumber="6" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE3A3168345FFF0FED2E178FE42AB78" blockId="5.[264,872,1324,1385]" box="[264,423,1359,1385]" pageId="5" pageNumber="6">
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<figureCitation id="1367BF938345FFF0FED2E178FE98AB78" box="[264,381,1359,1385]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="5" pageNumber="6">Figs 7–11</figureCitation>
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,
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<figureCitation id="1367BF938345FFF0FE50E178FE42AB78" box="[394,423,1359,1385]" captionStart="FIGURE 58" captionStartId="23.[264,368,1765,1789]" captionTargetBox="[418,1164,324,1727]" captionTargetId="figure@23.[396,1191,307,1742]" captionTargetPageId="23" captionText="FIGURE 58. Map of Madagascar showing distributions of Helicopsyche spp. (Helicopsychidae): H. ambodiva (circle); H. ninakosha and H. sahadika (rectangle); H. giboni (diamond); and H. anomana (triangle)." httpUri="https://zenodo.org/record/172389/files/figure.png" pageId="5" pageNumber="6">58</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C346F09D8345FFF2FED2E1A8FCEEAC8F" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
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<paragraph id="8BE3A3168345FFF0FED2E1A8FCACA848" blockId="5.[264,1324,1439,1905]" pageId="5" pageNumber="6">
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This new species is close to
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<emphasis id="B9287F048345FFF0FD81E1A8FD36ABA9" box="[603,723,1439,1464]" italics="true" pageId="5" pageNumber="6">ninakhosa</emphasis>
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, new species, sharing the character states of fused pronotal setal warts, and reduced ventral part of segment IX. However, males are easily distinguished from
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<emphasis id="B9287F048345FFF0FE37E1D8FD80A819" box="[493,613,1519,1544]" italics="true" pageId="5" pageNumber="6">ninakhosa</emphasis>
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by the absence of a prominently modified ventral branch of the gonocoxite, segment IX more rounded, Xth tergum longer, gonocoxites apically bent ventrally and having a long, curved basal branch.
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</paragraph>
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<paragraph id="8BE3A3168345FFF0FEE2E250FEAAA960" blockId="5.[264,1324,1439,1905]" pageId="5" pageNumber="6">
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Male head and thorax (
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<figureCitation id="1367BF938345FFF0FD94E250FD7CA890" box="[590,665,1639,1665]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="5" pageNumber="6">Fig. 7</figureCitation>
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): Scape about 3.5x as long as wide, nearly 1.5x longer than eye diameter. Maxillary palp (
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<figureCitation id="1367BF938345FFF0FD6CE2B8FCE5A8B8" box="[694,768,1679,1705]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="5" pageNumber="6">Fig. 7</figureCitation>
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) 2segmented, uniformly broad, 2nd segment slightly longer than first. Interantennal warts separate from each other. Cephalic setal warts small, rounded triangular. Postantennal setal warts small, flat, rounded. Postocular setal warts not conspicuous. Pronotum with broad setal warts completely fused medially to form a long, transverse wart. Mesoscutum and mesoscutellum each with one pair setal warts.
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</paragraph>
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<caption id="DF23F39E8346FFF3FED2E0CCFEBCAB46" httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="6" pageNumber="7" targetBox="[289,1314,326,1260]" targetPageId="6">
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<paragraph id="8BE3A3168346FFF3FED2E0CCFEBCAB46" blockId="6.[264,1323,1275,1367]" pageId="6" pageNumber="7">
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<emphasis id="B9287F048346FFF3FED2E0CCFE23AB02" bold="true" box="[264,454,1275,1299]" pageId="6" pageNumber="7">FIGURES 7–11.</emphasis>
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<taxonomicName id="4C5CD8958346FFF3FE0CE0CCFC93AB03" box="[470,886,1275,1298]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sahadika" status="sp. nov." subGenus="Petrotrichia">
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<emphasis id="B9287F048346FFF3FE0CE0CCFD80AB03" box="[470,613,1275,1298]" italics="true" pageId="6" pageNumber="7">Helicopsyche</emphasis>
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(
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<emphasis id="B9287F048346FFF3FDA7E0CCFD1BAB03" box="[637,766,1275,1298]" italics="true" pageId="6" pageNumber="7">Petrotrichia</emphasis>
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)
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<emphasis id="B9287F048346FFF3FCCCE0CCFC93AB03" box="[790,886,1275,1298]" italics="true" pageId="6" pageNumber="7">sahadika</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21BC27F8346FFF3FC51E0CCFBF3AB03" box="[907,1046,1275,1298]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
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, holotype. 7—head and thorax, dorsal; 8—right wing pair; 9—genitalia, lateral; 10—genitalia, dorsal; 11—genitalia, ventral.
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</paragraph>
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</caption>
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<paragraph id="8BE3A3168346FFF3FEE2E1BCFE6DA854" blockId="6.[264,1323,1419,1885]" pageId="6" pageNumber="7">
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Male wings (
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<figureCitation id="1367BF938346FFF3FE0BE1BCFDFCABB4" box="[465,537,1419,1445]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
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): Forewing length
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<quantity id="4CA40EF38346FFF3FD2DE1BCFCDBABB4" box="[759,830,1419,1445]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="mm" value="3.0">3 mm</quantity>
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, hind wing length 2.6 mm. Denuded wing membrane pale flavous with pale helvus veins. In forewing R5 originating distad of crossvein R3R4. M and Cu separate; a longitudinal groove with sensilla. Discoidal cell long, about 1/
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<date id="FFE285D68346FFF3FE7DE234FE22A80C" box="[423,455,1539,1565]" pageId="6" pageNumber="7">4x</date>
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the wing length. Abdominal reticulation absent. VIth sternal process well developed.
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</paragraph>
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<paragraph id="8BE3A3168346FFF2FEE2E264FB56AFEF" blockId="6.[264,1323,1419,1885]" lastBlockId="7.[264,1323,284,670]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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Male abdomen and genitalia (
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<figureCitation id="1367BF938346FFF3FD7AE264FCF8A87C" box="[672,797,1619,1645]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="6" pageNumber="7">Figs 9–11</figureCitation>
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): Segment IX strongly modified: extremely narrow, with clubshaped, anterior lobe located at dorsalmost part; the lateral and ventral regions reduced to narrow bands. Superior appendages fingerlike, downwardly directed in lateral (
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<figureCitation id="1367BF938346FFF3FEB8E2FCFE4EA8F4" box="[354,427,1739,1765]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
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), and slightly divergent in dorsal, aspects (
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<figureCitation id="1367BF938346FFF3FC70E2FCFBE6A8F4" box="[938,1027,1739,1765]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="6" pageNumber="7">Fig. 10</figureCitation>
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). Tergum X forms a pair of sigmoid elongate processes, tapering along its length; narrow in dorsal view (
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<figureCitation id="1367BF938346FFF3FB19E2C4FAF9A91C" box="[1219,1308,1779,1805]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="6" pageNumber="7">Fig. 10</figureCitation>
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); wide in lateral view (
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<figureCitation id="1367BF938346FFF3FE27E32CFDA6A924" box="[509,579,1819,1845]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
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), apex bifid and dorsally directed. Gonocoxite forms a complex of narrow strips, the distal end armed with a strong and elongate subapical spine (
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<figureCitation id="1367BF938346FFF2FB02E374FEC2AF27" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">Figs 9, 11</figureCitation>
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); apex of primary branch angled 90°, ventral branch oriented ventrally and divided into anterior and posterior lobes (
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<figureCitation id="1367BF938347FFF2FDAEE573FD27AF4F" box="[628,706,324,350]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
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). A pair of very long, slender basal branches of gonocoxite oriented dorsally at basis, curve posteriorly and slightly ventrally near apex, with distal 1/3. annulate. Phallus about as long as basal branch of gonocoxite, forming a wide tube, with undulating dorsal and ventral margins (
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<figureCitation id="1367BF938347FFF2FC1CE58BFBF0AFC7" box="[966,1045,444,470]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
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), subapical flank well sclerotized and, in lateral aspect, ending in a downwardly curving hook (
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<figureCitation id="1367BF938347FFF2FB87E5D3FB40AFEF" box="[1117,1189,484,510]" captionStart="FIGURES 7 – 11" captionStartId="6.[264,382,1275,1299]" captionTargetBox="[289,1314,326,1260]" captionTargetId="figure@6.[269,1328,318,1279]" captionTargetPageId="6" captionText="FIGURES 7 – 11. Helicopsyche (Petrotrichia) sahadika, new species, holotype. 7 — head and thorax, dorsal; 8 — right wing pair; 9 — genitalia, lateral; 10 — genitalia, dorsal; 11 — genitalia, ventral." httpUri="https://zenodo.org/record/172378/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
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).
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</paragraph>
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<paragraph id="8BE3A3168347FFF2FEE2E63BFC57AC5F" blockId="7.[264,1323,284,670]" pageId="7" pageNumber="8">
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Etymology. “
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<taxonomicName id="4C5CD8958347FFF2FE04E63BFDADAC34" box="[478,584,524,549]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sahadika">
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<emphasis id="B9287F048347FFF2FE04E63BFDADAC34" box="[478,584,524,549]" italics="true" pageId="7" pageNumber="8">sahadika</emphasis>
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</taxonomicName>
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”, from Sanscrit, meaning frame, referring to the unique framework of the narrow strip complex of the gonocoxite.
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</paragraph>
|
||
<paragraph id="8BE3A3168347FFF2FEE2E66BFCEEAC8F" blockId="7.[264,1323,284,670]" pageId="7" pageNumber="8">
|
||
<typeStatus id="54E71DB48347FFF2FEE2E66BFE40AC67" box="[312,421,604,630]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
|
||
male:
|
||
<collectingCountry id="F34BE3868347FFF2FE24E66BFD36AC67" box="[510,723,604,630]" name="Madagascar" pageId="7" pageNumber="8">MADAGASCAR</collectingCountry>
|
||
: Demongloy, Mananteina Fort, Dauphin District,
|
||
<date id="FFE285D68347FFF2FED2E6B3FE95AC8F" box="[264,368,644,670]" pageId="7" pageNumber="8" value="1948-12">xii. 1948</date>
|
||
[Hoogstraal] (MNHN, in alcohol).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |