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<document ID-DOI="http://doi.org/10.5281/zenodo.4759732" ID-GBIF-Dataset="3c7c2c9b-5ab5-45aa-92f5-f99f8456a1b4" ID-GBIF-Taxon="182165175" ID-ISSN="1854-0392" ID-Zenodo-Dep="4759732" checkinTime="1620959558361" checkinUser="felipe" docAuthor="Sivec, Ignac &amp; Stark, Bill P." docDate="2010" docId="03ECD40EFFC64E00D7A81364FBFFF9BC" docLanguage="en" docName="Illiesia.6.10.98-112.pdf" docOrigin="Illiesia 6 (10)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Phanoperla doisuthep Sivec &amp; Stark 2010, sp. n." docType="treatment" docVersion="7" lastPageId="1" lastPageNumber="99" masterDocId="FFD5AC76FFC64E01D4621503FFC2FFDF" masterDocTitle="Seven New Species Of Phanoperla Banks From Vietnam And Thailand (Plecoptera: Perlidae)" masterLastPageNumber="112" masterPageNumber="98" pageId="0" pageNumber="98" updateTime="1632022735424" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Seven New Species Of Phanoperla Banks From Vietnam And Thailand (Plecoptera: Perlidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sivec, Ignac</mods:namePart>
<mods:affiliation>Slovenian Museum of Natural History, Prešernova 20, P. O. Box 290, SLO- 1001 Ljubljana, Slovenia E-mail: isivec @ pms-lj. si</mods:affiliation>
<mods:nameIdentifier type="email">isivec@pms-lj.si</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>10</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>98</mods:start>
<mods:end>112</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4759732</mods:identifier>
<mods:identifier type="GBIF-Dataset">3c7c2c9b-5ab5-45aa-92f5-f99f8456a1b4</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4759732</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182165175" LSID="urn:lsid:plazi:treatment:03ECD40EFFC64E00D7A81364FBFFF9BC" httpUri="http://treatment.plazi.org/id/03ECD40EFFC64E00D7A81364FBFFF9BC" lastPageId="1" lastPageNumber="99" pageId="0" pageNumber="98">
<subSubSection box="[970,1308,1639,1664]" pageId="0" pageNumber="98" type="nomenclature">
<paragraph blockId="0.[970,1308,1639,1664]" box="[970,1308,1639,1664]" pageId="0" pageNumber="98">
<heading bold="true" box="[970,1308,1639,1664]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="98" reason="2">
<emphasis bold="true" box="[970,1308,1639,1664]" pageId="0" pageNumber="98">
<taxonomicName authority="Sivec &amp; Stark, 2010" authorityName="Sivec &amp; Stark" authorityYear="2010" box="[970,1230,1639,1663]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="98" phylum="Arthropoda" rank="species" species="doisuthep" status="sp. n.">
<emphasis bold="true" box="[970,1230,1639,1663]" italics="true" pageId="0" pageNumber="98">Phanoperla doisuthep</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1243,1308,1640,1664]" pageId="0" pageNumber="98" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1081,1197,1675,1700]" pageId="0" pageNumber="98" type="description">
<paragraph blockId="0.[1081,1197,1675,1700]" box="[1081,1197,1675,1700]" pageId="0" pageNumber="98">
(
<figureCitation box="[1089,1188,1675,1700]" captionStart="Figs" captionStartId="1.[189,241,900,924]" captionTargetBox="[252,1418,235,834]" captionTargetId="figure-50@1.[689,992,252,734]" captionTargetPageId="1" captionText="Figs. 1-3. Phanoperla doisuthep. 1. Male terminalia, dorsal. 2. Fully everted aedeagus, lateral. 3. Aedeagal sac apex." figureDoi="http://doi.org/10.5281/zenodo.4759734" httpUri="https://zenodo.org/record/4759734/files/figure.png" pageId="0" pageNumber="98">Figs. 1-3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="1" lastPageNumber="99" pageId="0" pageNumber="98" type="materials_examined">
<paragraph blockId="0.[835,1443,1745,1913]" lastBlockId="1.[189,798,1042,1919]" lastPageId="1" lastPageNumber="99" pageId="0" pageNumber="98">
<emphasis bold="true" box="[835,1073,1746,1770]" pageId="0" pageNumber="98">Material examined</emphasis>
.
<materialsCitation ID-GBIF-Occurrence="3111998338" collectingDate="2002-06-06" collectingDateMax="2002-06-07" collectingDateMin="2002-06-06" collectionCode="PMSL" country="Thailand" county="Doi Suthep-Pui National Park" elevation="1000" latitude="98.916664" location="Namtok Sai Yoi" longLatPrecision="936" longitude="18.8" municipality="Namtok Sai Yoi" pageId="0" pageNumber="98" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai Province" typeStatus="holotype">
<typeStatus box="[1095,1207,1747,1771]" pageId="0" pageNumber="98">Holotype</typeStatus>
<specimenCount box="[1223,1243,1745,1770]" pageId="0" pageNumber="98" type="male"></specimenCount>
from
<collectingCountry box="[1331,1439,1747,1771]" name="Thailand" pageId="0" pageNumber="98">Thailand</collectingCountry>
,
<collectingRegion box="[835,1083,1782,1806]" country="Thailand" name="Chiang Mai" pageId="0" pageNumber="98">Chiang Mai Province</collectingRegion>
,
<collectingCounty box="[1093,1439,1782,1806]" pageId="0" pageNumber="98">Doi Suthep-Pui National Park</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:03ECD40EFFC64E00D7A81364FBFFF9BC:8E9A33C3FFC64E01D7211219FC3EF8ED" box="[835,1020,1818,1842]" country="Thailand" county="Doi Suthep-Pui National Park" latitude="98.916664" longLatPrecision="936" longitude="18.8" municipality="Namtok Sai Yoi" name="Namtok Sai Yoi" pageId="0" pageNumber="98" stateProvince="Chiang Mai Province">Namtok Sai Yoi</location>
, 98° 55 N, 18° 48 E,
<quantity box="[1255,1338,1818,1842]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="98" unit="m" value="1000.0">
<elevation box="[1255,1338,1818,1842]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="98" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date pageId="0" pageNumber="98" value="2002-06-06" valueMax="2002-06-07" valueMin="2002-06-06">
<collectingDate pageId="0" pageNumber="98" value="2002-06-06" valueMax="2002-06-07" valueMin="2002-06-06">6-7 June 2002</collectingDate>
</date>
, Chiang Mai University Team (
<collectionCode box="[1354,1431,1854,1878]" country="Slovenia" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="0" pageNumber="98">PMSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111998320" collectingDate="2002-05-06" collectingDateMax="2002-05-26" collectingDateMin="2002-05-06" collectionCode="PMSL" collectorName="University Team" country="Thailand" county="National Park" elevation="800" lastPageId="1" lastPageNumber="99" latitude="98.916664" location="Doi Suthep-Pui" longLatPrecision="936" longitude="18.816668" municipality="Huai Kaew" pageId="0" pageNumber="98" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
<typeStatus box="[835,954,1889,1913]" pageId="0" pageNumber="98">Paratypes</typeStatus>
:
<collectingRegion box="[973,1238,1889,1913]" country="Thailand" name="Chiang Mai" pageId="0" pageNumber="98">Chiang Mai Province</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:03ECD40EFFC64E00D7A81364FBFFF9BC:8E9A33C3FFC64E01D0871262FA61F8A6" box="[1253,1443,1889,1913]" country="Thailand" name="Doi Suthep-Pui" pageId="0" pageNumber="98" stateProvince="Chiang Mai">Doi Suthep-Pui</location>
<collectingCounty box="[189,358,1042,1066]" pageId="1" pageNumber="99">National Park</collectingCounty>
,
<collectingMunicipality box="[371,506,1042,1066]" pageId="1" pageNumber="99">Huai Kaew</collectingMunicipality>
above
<location LSID="urn:lsid:plazi:treatment:03ECD40EFFC64E00D7A81364FBFFF9BC:8E9A33C3FFC74E00D6331111FD28FBF5" box="[593,746,1042,1066]" county="National Park" latitude="98.916664" longLatPrecision="936" longitude="18.816668" municipality="Huai Kaew" name="Monthratarn" pageId="1" pageNumber="99" stateProvince="Chiang Mai">Monthratarn</location>
, 98° 55 N, 18° 49 E,
<quantity box="[386,458,1077,1101]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="99" unit="m" value="800.0">
<elevation box="[386,458,1077,1101]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="99" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date box="[471,645,1077,1102]" pageId="1" pageNumber="99" value="2002-05-06" valueMax="2002-05-26" valueMin="2002-05-06">
<collectingDate box="[471,645,1077,1102]" pageId="1" pageNumber="99" value="2002-05-06" valueMax="2002-05-26" valueMin="2002-05-06">6-26 May 2002</collectingDate>
</date>
,
<collectingRegion box="[657,796,1077,1101]" country="Thailand" name="Chiang Mai" pageId="1" pageNumber="99">Chiang Mai</collectingRegion>
<collectorName box="[189,387,1113,1137]" pageId="1" pageNumber="99">University Team</collectorName>
<specimenCount box="[394,427,1112,1137]" pageId="1" pageNumber="99" type="male">1♂</specimenCount>
(
<collectionCode box="[444,520,1113,1137]" country="Slovenia" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="1" pageNumber="99">PMSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111998337" collectingDate="2003-05-04" collectingDateMax="2003-05-05" collectingDateMin="2003-05-04" collectionCode="PMSL" collectorName="I. Sivec" elevation="1280" latitude="101.0" location="Waterwheel Falls" longLatPrecision="942" longitude="16.983334" municipality="Phu Hin Rongkla National Park" pageId="1" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Phitsanulok">
<collectingRegion box="[538,792,1113,1137]" country="Thailand" name="Phitsanulok" pageId="1" pageNumber="99">Phitsanulok Province</collectingRegion>
,
<collectingMunicipality box="[189,572,1148,1172]" pageId="1" pageNumber="99">Phu Hin Rongkla National Park</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03ECD40EFFC64E00D7A81364FBFFF9BC:8E9A33C3FFC74E00D62A117FFCDAFB4B" box="[584,792,1148,1172]" latitude="101.0" longLatPrecision="942" longitude="16.983334" municipality="Phu Hin Rongkla National Park" name="Waterwheel Falls" pageId="1" pageNumber="99" stateProvince="Phitsanulok">Waterwheel Falls</location>
, 101° 00 N, 16° 59 E,
<quantity box="[442,528,1184,1208]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="1" pageNumber="99" unit="m" value="1280.0">
<elevation box="[442,528,1184,1208]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="1" pageNumber="99" unit="m" value="1280.0">1280 m</elevation>
</quantity>
,
<date box="[540,701,1184,1208]" pageId="1" pageNumber="99" value="2003-05-04" valueMax="2003-05-05" valueMin="2003-05-04">
<collectingDate box="[540,701,1184,1208]" pageId="1" pageNumber="99" value="2003-05-04" valueMax="2003-05-05" valueMin="2003-05-04">4-5 May 2003</collectingDate>
</date>
,
<collectorName box="[712,796,1184,1208]" pageId="1" pageNumber="99">I. Sivec</collectorName>
(
<collectionCode box="[197,272,1220,1244]" country="Slovenia" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="1" pageNumber="99">PMSL</collectionCode>
)
<specimenCount box="[285,319,1218,1244]" pageId="1" pageNumber="99" type="male">1♂</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4759734" ID-Zenodo-Dep="4759734" httpUri="https://zenodo.org/record/4759734/files/figure.png" pageId="1" pageNumber="99" startId="1.[189,241,900,924]" targetBox="[252,1418,235,834]" targetPageId="1">
<paragraph blockId="1.[189,1442,900,959]" pageId="1" pageNumber="99">
Figs. 1-3.
<taxonomicName authorityName="Sivec &amp; Stark" authorityYear="2010" box="[301,532,900,924]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="99" phylum="Arthropoda" rank="species" species="doisuthep">
<emphasis box="[301,532,900,924]" italics="true" pageId="1" pageNumber="99">Phanoperla doisuthep</emphasis>
</taxonomicName>
. 1. Male terminalia, dorsal. 2. Fully everted aedeagus, lateral. 3. Aedeagal sac apex.
</paragraph>
</caption>
<subSubSection pageId="1" pageNumber="99" type="description">
<paragraph blockId="1.[189,798,1042,1919]" pageId="1" pageNumber="99">
<emphasis bold="true" box="[189,364,1255,1279]" pageId="1" pageNumber="99">Adult habitus.</emphasis>
Biocellate. Ocelli large, close together and bordered by dark pigment on inner margins. Head yellow brown with slightly darker M-line and calluses; antennal bases yellow and flagellum brown. Pronotum pale with brown rugosities. Legs uniformly pale brown with small brown oval spot in upper third of tibiae; apical tarsal segment only slightly brownish. Cerci uniformly pale.
</paragraph>
<paragraph blockId="1.[189,798,1042,1919]" lastBlockId="1.[835,1444,1042,1635]" pageId="1" pageNumber="99">
<emphasis bold="true" box="[189,258,1540,1564]" pageId="1" pageNumber="99">Male.</emphasis>
Forewing length
<quantity box="[463,546,1540,1564]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="99" unit="mm" value="10.0">10 mm</quantity>
. Tergum 8 with small mesal lobe. Tergum 9 with large lateral and median sensilla basiconica patches set on low mounds (
<figureCitation captionStart="Figs" captionStartId="1.[189,241,900,924]" captionTargetBox="[252,1418,235,834]" captionTargetId="figure-50@1.[689,992,252,734]" captionTargetPageId="1" captionText="Figs. 1-3. Phanoperla doisuthep. 1. Male terminalia, dorsal. 2. Fully everted aedeagus, lateral. 3. Aedeagal sac apex." figureDoi="http://doi.org/10.5281/zenodo.4759734" httpUri="https://zenodo.org/record/4759734/files/figure.png" pageId="1" pageNumber="99">Fig. 1</figureCitation>
). Hemitergal processes wide at base, tapered to an acute apex and curved slightly laterad. Aedeagal tube poorly sclerotized but with a pair of small ventrobasal knobs and a low dorsoapical membranous lobe (
<figureCitation box="[435,511,1788,1813]" captionStart="Figs" captionStartId="1.[189,241,900,924]" captionTargetBox="[252,1418,235,834]" captionTargetId="figure-50@1.[689,992,252,734]" captionTargetPageId="1" captionText="Figs. 1-3. Phanoperla doisuthep. 1. Male terminalia, dorsal. 2. Fully everted aedeagus, lateral. 3. Aedeagal sac apex." figureDoi="http://doi.org/10.5281/zenodo.4759734" httpUri="https://zenodo.org/record/4759734/files/figure.png" pageId="1" pageNumber="99">Fig. 2</figureCitation>
); aedeagal sac curved strongly ventrad and bearing a rounded membranous lobe near midlength; sac armature restricted to a dorsobasal patch of small to moderate spines that extend onto lateral surface, and a more prominent subapical and apical patch which includes a prominent field of large cultriform spines (
<figureCitation box="[1360,1426,1113,1137]" captionStart="Figs" captionStartId="1.[189,241,900,924]" captionTargetBox="[252,1418,235,834]" captionTargetId="figure-50@1.[689,992,252,734]" captionTargetPageId="1" captionText="Figs. 1-3. Phanoperla doisuthep. 1. Male terminalia, dorsal. 2. Fully everted aedeagus, lateral. 3. Aedeagal sac apex." figureDoi="http://doi.org/10.5281/zenodo.4759734" httpUri="https://zenodo.org/record/4759734/files/figure.png" pageId="1" pageNumber="99">Fig. 3</figureCitation>
).
</paragraph>
<paragraph blockId="1.[835,1444,1042,1635]" box="[835,1062,1148,1173]" pageId="1" pageNumber="99">
<emphasis bold="true" box="[835,929,1149,1173]" pageId="1" pageNumber="99">Female.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="1.[835,1444,1042,1635]" box="[835,1042,1184,1208]" pageId="1" pageNumber="99">
<emphasis bold="true" box="[835,910,1184,1208]" pageId="1" pageNumber="99">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="99" type="etymology">
<paragraph blockId="1.[835,1444,1042,1635]" pageId="1" pageNumber="99">
<emphasis bold="true" box="[835,973,1220,1244]" pageId="1" pageNumber="99">Etymology.</emphasis>
The species name, used as a noun in apposition, is based on the
<typeStatus box="[1155,1207,1256,1280]" pageId="1" pageNumber="99">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="99" type="diagnosis">
<paragraph blockId="1.[835,1444,1042,1635]" pageId="1" pageNumber="99">
<emphasis bold="true" box="[835,964,1291,1315]" pageId="1" pageNumber="99">Diagnosis.</emphasis>
The aedeagus and male terminalia of
<taxonomicName authorityName="Sivec &amp; Stark" authorityYear="2010" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="99" phylum="Arthropoda" rank="species" species="doisuthep">
<emphasis italics="true" pageId="1" pageNumber="99">P. doisuthep</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName authority="Zwick" authorityName="Zwick" box="[1203,1439,1327,1351]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="99" phylum="Arthropoda" rank="species" species="himalayana">
<emphasis box="[1203,1356,1327,1351]" italics="true" pageId="1" pageNumber="99">P. himalayana</emphasis>
Zwick
</taxonomicName>
, a species known from
<collectingCountry box="[1130,1217,1363,1387]" name="Bhutan" pageId="1" pageNumber="99">Bhutan</collectingCountry>
,
<collectingCountry box="[1235,1297,1363,1387]" name="India" pageId="1" pageNumber="99">India</collectingCountry>
and
<collectingCountry box="[1371,1443,1363,1387]" name="Nepal" pageId="1" pageNumber="99">Nepal</collectingCountry>
(
<bibRefCitation author="Zwick, P." box="[843,981,1398,1422]" pageId="1" pageNumber="99" pagination="87 - 126" refId="ref7213" refString="Zwick, P. 1982. A revision of the Oriental stonefly genus Phanoperla (Plecoptera: Perlidae). Systematic Entomology, 7: 87 - 126." type="journal article" year="1982">Zwick 1982</bibRefCitation>
). It differs in having a poorly developed ventral sclerite and in lacking the ventroapical lobes on the aedeagal tube which occur distal to the sclerite in
<taxonomicName box="[873,1035,1505,1529]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="99" phylum="Arthropoda" rank="species" species="himalayana">
<emphasis box="[873,1035,1505,1529]" italics="true" pageId="1" pageNumber="99">P. himalayana</emphasis>
</taxonomicName>
. In addition,
<taxonomicName authorityName="Sivec &amp; Stark" authorityYear="2010" box="[1219,1360,1505,1529]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="99" phylum="Arthropoda" rank="species" species="doisuthep">
<emphasis box="[1219,1360,1505,1529]" italics="true" pageId="1" pageNumber="99">P. doisuthep</emphasis>
</taxonomicName>
has a prominent ventral lobe on the sac that is absent in
<taxonomicName class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="99" phylum="Arthropoda" rank="species" species="himalayana">
<emphasis italics="true" pageId="1" pageNumber="99">P. himalayana</emphasis>
</taxonomicName>
and the distribution of large apical spines on the sac also differ.
</paragraph>
</subSubSection>
</treatment>
</document>