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<document id="28EBF2D4B410C91753AC0C477EE96AB0" ID-DOI="10.5852/ejt.2024.942.2581" ID-ISSN="2118-9773" ID-Zenodo-Dep="12550288" ID-ZooBank="DF991BC4-1761-4C5D-96E5-EFC62F8F7D1A" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1719420232185" checkinUser="plazi" docAuthor="Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna &amp; Żyła, Dagmara" docDate="2024" docId="03EC292BFF89FF80FE625D96DC4AFB6C" docLanguage="en" docName="EJT.2024.942.1-76.pdf" docOrigin="European Journal of Taxonomy 942" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Neolindus parahermani Guzman, Tokareva &amp; Zyla 2024, sp. nov." docType="treatment" docUuid="2D2F08A4-E884-4C1C-8AED-B289223FDC9B" docUuidSource="ZooBank" docVersion="3" lastPageNumber="41" masterDocId="FFD55153FFAFFFA8FFDA5C0DD87EFFDF" masterDocTitle="Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="39" updateTime="1719496693333" updateUser="valdenar" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6EFEC8128B292F918D97FE7835926AFB">Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae)</mods:title>
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<mods:namePart id="B475DE7D9F1227A3D544977EA2BA645B">Guzman, Yoan Camilo</mods:namePart>
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<mods:affiliation id="097E1EC527092FE738C22B854380BC41">Museum and Institute of Zoology, Polish Academy of Sciences, Warsaw, Poland.</mods:affiliation>
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<mods:namePart id="B9F8A9D3198796DF6444EA341A4B8EA2">Tokareva, Alexandra</mods:namePart>
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<mods:affiliation id="7A2FA9075E15D513E37762BC6AAD5C8E">Museum and Institute of Zoology, Polish Academy of Sciences, Warsaw, Poland.</mods:affiliation>
<mods:nameIdentifier id="61D52889CF32671035406C28E16BC78C" type="email">atokareva@miiz.waw.pl</mods:nameIdentifier>
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<mods:namePart id="5D2F9F030B0583426108D2422C9E6415">Koszela, Katarzyna</mods:namePart>
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<mods:affiliation id="6D06A51581F354BFE087AE79380DB276">Museum and Institute of Zoology, Polish Academy of Sciences, Warsaw, Poland.</mods:affiliation>
<mods:nameIdentifier id="B8DC6C2E9E144BBF022FFF3DE36D50CF" type="email">kat.koszela@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EA6FD4B1AB17669DF91C1B69D35260D0">Żyła, Dagmara</mods:namePart>
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<mods:affiliation id="B38C7DF4510030DFC3D98BB571FA82C8">Museum of Nature Hamburg, Leibniz Institute for the Analysis of Biodiversity Change, Hamburg, Germany.</mods:affiliation>
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<mods:title id="EC98036E94B194BAD67E17FF08D9586F">European Journal of Taxonomy</mods:title>
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<mods:date id="C4FEC26139247F2A8FADEE8A5BE72FC7">2024</mods:date>
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<heading id="D0B22F51FF89FF8EFE625D96DC02FE69" box="[440,1148,411,440]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="39" reason="2">
<taxonomicName id="4C45E3BEFF89FF8EFE625D96DC64FE67" authority="Guzman, Tokareva &amp; Zyla" authorityName="Guzman, Tokareva &amp; Zyla" authorityYear="2024" box="[440,1050,411,440]" class="Insecta" family="Staphylinidae" genus="Neolindus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="parahermani" status="sp. nov.">
<emphasis id="B931442FFF89FF8EFE625D96DAA7FE69" bold="true" box="[440,729,411,438]" italics="true" pageId="38" pageNumber="39">Neolindus parahermani</emphasis>
Guzman, Tokareva &amp; Żyła
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<taxonomicNameLabel id="A202F954FF89FF8EFBFB5D91DC02FE69" box="[1057,1148,412,438]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="137E84B8FF89FF8EFD7F5DEFDA85FE23" box="[677,763,482,508]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[213,1380,351,1821]" captionTargetId="figure-12@39.[208,1380,265,1847]" captionTargetPageId="39" captionText="Fig. 13. Neolindus parahermani Guzman, Tokareva &amp; Żyła sp. nov., holotype, ♂ (NHMUK). A. Habitus, dorsal view. B−E. Abdominal segments, posterior margin schemes. B. Tergite VII. C. Sternite VII. D. Tergite VIII. E. Sternite VIII. F−I. Aedeagus. F. Parameral view. G. Parameral view scheme. H. Lateral view. I. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="38" pageNumber="39">Fig. 13</figureCitation>
; Supp. file 2
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<subSubSection id="C35FCBB6FF89FF8EFF675E2EDA0DFD09" pageId="38" pageNumber="39" type="diagnosis">
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<emphasis id="B931442FFF89FF8EFF675E2ED948FDE2" bold="true" box="[189,310,547,573]" pageId="38" pageNumber="39">Diagnosis</emphasis>
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<taxonomicName id="4C45E3BEFF89FF8EFF675E5FD9ABFDB3" authorityName="Guzman &amp; Tokareva &amp; Koszela &amp; Żyła" authorityYear="2024" box="[189,469,594,620]" class="Insecta" family="Staphylinidae" genus="Neolindus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="parahermani" status="sp. nov.">
<emphasis id="B931442FFF89FF8EFF675E5FD9ABFDB3" box="[189,469,594,620]" italics="true" pageId="38" pageNumber="39">Neolindus parahermani</emphasis>
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<taxonomicNameLabel id="A202F954FF89FF8EFE045E5EDA47FDB2" box="[478,569,595,621]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
resembles 
<taxonomicName id="4C45E3BEFF89FF8EFD655E5EDB3AFDB3" authorityName="Asenjo" authorityYear="2011" box="[703,836,594,620]" class="Insecta" family="Staphylinidae" genus="Neolindus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hermani">
<emphasis id="B931442FFF89FF8EFD655E5EDB3AFDB3" box="[703,836,594,620]" italics="true" pageId="38" pageNumber="39">N. hermani</emphasis>
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in habitus and genitalia morphology. However, they can be distinguished by the structure of pPMS of aedeagus. Antiparameral apical arms of pPMS in 
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<emphasis id="B931442FFF89FF8EFF015E94D9E9FD6D" box="[219,407,664,690]" italics="true" pageId="38" pageNumber="39">N. parahermani</emphasis>
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are bifid with two acute processes, while in 
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<emphasis id="B931442FFF89FF8EFC7B5E94DC58FD6D" box="[929,1062,664,690]" italics="true" pageId="38" pageNumber="39">N. hermani</emphasis>
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, they end with a rounded tip (
<bibRefCitation id="EFD4E5CCFF89FF8EFF1F5EB1D921FD09" author="Asenjo A." box="[197,351,700,726]" pageId="38" pageNumber="39" pagination="57 - 67" refId="ref41542" refString="Asenjo A. 2011. First record of Neolindus Scheerpeltz from French Guiana (Coleoptera, Staphylinidae, Paederinae), with a key to males. Zookeys 135: 57 - 67. https: // doi. org / 10.3897 / zookeys. 135.1740" type="journal article" year="2011">Asenjo 2011</bibRefCitation>
: figs 9–10; 
<figureCitation id="137E84B8FF89FF8EFE3C5EB1DA18FD09" box="[486,614,700,726]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[213,1380,351,1821]" captionTargetId="figure-12@39.[208,1380,265,1847]" captionTargetPageId="39" captionText="Fig. 13. Neolindus parahermani Guzman, Tokareva &amp; Żyła sp. nov., holotype, ♂ (NHMUK). A. Habitus, dorsal view. B−E. Abdominal segments, posterior margin schemes. B. Tergite VII. C. Sternite VII. D. Tergite VIII. E. Sternite VIII. F−I. Aedeagus. F. Parameral view. G. Parameral view scheme. H. Lateral view. I. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="38" pageNumber="39">Fig. 13F–I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35FCBB6FF89FF8EFF675EF0DBC4FC53" pageId="38" pageNumber="39" type="etymology">
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<emphasis id="B931442FFF89FF8EFF675EF0D93AFCC8" bold="true" box="[189,324,765,791]" pageId="38" pageNumber="39">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA983DFF89FF8EFF675F21DBC4FC53" blockId="38.[189,1398,812,909]" pageId="38" pageNumber="39">
The specific epithet is the combination of ‘ 
<emphasis id="B931442FFF89FF8EFD135F21DB7EFC9A" box="[713,768,812,837]" italics="true" pageId="38" pageNumber="39">para</emphasis>
’, a Latin prefix that means ‘similar’ or ‘equal’, and 
<taxonomicName id="4C45E3BEFF89FF8EFF675F42D957FCB6" authorityName="Asenjo" authorityYear="2011" box="[189,297,847,873]" class="Insecta" family="Staphylinidae" genus="Neolindus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hermani">
‘ 
<emphasis id="B931442FFF89FF8EFF1D5F42D957FCB6" box="[199,297,847,873]" italics="true" pageId="38" pageNumber="39">hermani</emphasis>
</taxonomicName>
’ as the name of the species described by 
<bibRefCitation id="EFD4E5CCFF89FF8EFCD05F42DBCAFCB6" author="Asenjo A." box="[778,948,847,873]" pageId="38" pageNumber="39" pagination="57 - 67" refId="ref41542" refString="Asenjo A. 2011. First record of Neolindus Scheerpeltz from French Guiana (Coleoptera, Staphylinidae, Paederinae), with a key to males. Zookeys 135: 57 - 67. https: // doi. org / 10.3897 / zookeys. 135.1740" type="journal article" year="2011">Asenjo (2011)</bibRefCitation>
, 
<emphasis id="B931442FFF89FF8EFC1A5F5DDC32FCB6" box="[960,1100,847,873]" italics="true" pageId="38" pageNumber="39">
<taxonomicName id="4C45E3BEFF89FF8EFC1A5F5DDC36FCB6" authorityName="Asenjo" authorityYear="2011" box="[960,1096,847,873]" class="Insecta" family="Staphylinidae" genus="Neolindus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hermani">N. hermani</taxonomicName>
.
</emphasis>
It indicates the similarity of morphology between aedeagi of the two species. An adjective.
</paragraph>
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<subSubSection id="C35FCBB6FF89FF8EFF675FB9DA89FBB9" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BFA983DFF89FF8EFF675FB9D912FC12" blockId="38.[189,364,947,974]" box="[189,364,947,974]" pageId="38" pageNumber="39">
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<emphasis id="B931442FFF89FF8EFF675FB9D912FC12" bold="true" box="[189,364,947,974]" pageId="38" pageNumber="39">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA983DFF89FF8EFF305FEFDA89FBB9" blockId="38.[189,1399,994,1126]" pageId="38" pageNumber="39">
<materialsCitation id="3B2D9260FF89FF8EFF305FEFDA89FBB9" collectingDate="2009-12" collectionCode="BMNH, NHMUK" country="French Guiana" elevation="225" latitude="4.6102777" location="FRENCH GUIANA" longLatPrecision="1" longitude="-52.278473" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B931442FFF89FF8EFF305FEFD922FC23" bold="true" box="[234,348,994,1020]" pageId="38" pageNumber="39">
<typeStatus id="54FE269FFF89FF8EFF305FEFD922FC23" box="[234,348,994,1020]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F352D8ADFF89FF8EFF675808D9D5FBC0" box="[189,427,1029,1055]" name="French Guiana" pageId="38" pageNumber="39">FRENCH GUIANA</collectingCountry>
• 
<specimenCount id="9D4353B4FF89FF8EFE1D580BD99EFBFF" box="[455,480,1030,1056]" count="1" pageId="38" pageNumber="39" type="male">♂</specimenCount>
; “ 
<taxonomicName id="4C45E3BEFF89FF8EFE205808DCE2FBC0" authority="Guzman, Tokareva &amp; Zyla 2024" authorityName="Guzman, Tokareva &amp; Zyla" authorityYear="2024" box="[506,1180,1029,1058]" class="Insecta" family="Staphylinidae" genus="Neolindus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="parahermani">
<emphasis id="B931442FFF89FF8EFE205808DB6DFBC0" box="[506,787,1029,1055]" italics="true" pageId="38" pageNumber="39">Neolindus parahermani</emphasis>
Guzman, Tokareva &amp; Żyła 2024
</taxonomicName>
<typeStatus id="54FE269FFF89FF8EFB7E5808DD41FBC0" box="[1188,1343,1029,1055]" pageId="38" pageNumber="39" type="holotype">HOLOTYPE</typeStatus>
[red label] \\ Réserve Trésor, cca 
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</quantity>
: 
<geoCoordinate id="EE71FEFAFF89FF8EFDBD5825DB7FFB9D" box="[615,769,1064,1090]" degrees="4" direction="north" minutes="36" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="37.0" value="4.6102777">4°36’37.0”N</geoCoordinate>
, 
<geoCoordinate id="EE71FEFAFF89FF8EFCD55825DBC3FB9C" box="[783,957,1064,1091]" degrees="52" direction="west" minutes="16" orientation="longitude" pageId="38" pageNumber="39" precision="1" seconds="42.5" value="-52.278473">52°16’42.5”W</geoCoordinate>
; 
<date id="FFFBBEFDFF89FF8EFC115824DC3FFB9D" box="[971,1089,1064,1091]" pageId="38" pageNumber="39" value="2009-12">
<collectingDate id="EFBF4715FF89FF8EFC115824DC3FFB9D" box="[971,1089,1064,1091]" pageId="38" pageNumber="39" value="2009-12">Dec 2009</collectingDate>
</date>
; Stephane Brule Leg.; ex. 
<collectingMethod id="5204E02AFF89FF8EFF675846D928FBB9" box="[189,342,1099,1126]" pageId="38" pageNumber="39">Window trap</collectingMethod>
\\ 
<collectionCode id="ED5400F8FF89FF8EFEAF5841D9A9FBB9" box="[373,471,1100,1126]" pageId="38" pageNumber="39">BMNH</collectionCode>
{E} 2010-62”; 
<collectionCode id="ED5400F8FF89FF8EFD5A5841DA8DFBB9" box="[640,755,1100,1126]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="38" pageNumber="39" type="Museum">NHMUK</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35FCBB6FF89FF80FF675880D94CFBC0" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" type="description">
<paragraph id="8BFA983DFF89FF8EFF675880D930FB78" blockId="38.[189,334,1165,1191]" box="[189,334,1165,1191]" pageId="38" pageNumber="39">
<heading id="D0B22F51FF89FF8EFF675880D930FB78" bold="true" box="[189,334,1165,1191]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B931442FFF89FF8EFF675880D930FB78" bold="true" box="[189,334,1165,1191]" pageId="38" pageNumber="39">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA983DFF89FF8EFF6758B1DA65FABD" blockId="38.[189,1400,1211,1379]" pageId="38" pageNumber="39">
<smallCapsWord id="8D1C0EE1FF89FF8EFF6758B1D910FB0B" baselines="1231,1231" box="[189,366,1212,1238]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="38" pageNumber="39">MEASUREMENTS</smallCapsWord>
. BL (13.63), H (0.96, 2.03), A (3.56), A1 (0.78, 0.16), A2 (0.19, 0.14), A3 (0.37, 0.13), A4 (0.27, 0.1), A5 (0.28, 0.11), A6 (0.33, 0.1), A7 (0.34, 0.1), A8 (0.31, 0.1), A9 (0.26, 0.1), A10 (0.2, 0.11), A11 (0.21, 0.1), NKW (0.85), GL (0.4), P (1.87, 2.3), E (2.63, 2.49), PC (0.7, 0.42), PF (1.35, 0.76), PT (0.81, 0.35), MSC (0.72, 0.59), MSF (1.58, 0.47), MST (1.14, 0.15), MTC (0.71, 0.64), MTF (1.72, 0.49), MTT (1.7, 0.22).
</paragraph>
<paragraph id="8BFA983DFF89FF8EFF675987D933FA18" blockId="38.[189,1399,1418,1479]" pageId="38" pageNumber="39">
<smallCapsWord id="8D1C0EE1FF89FF8EFF675987D929FA7D" baselines="1437,1437" box="[189,343,1418,1444]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colouration" pageId="38" pageNumber="39">COLOURATION</smallCapsWord>
. Head and pronotum black; legs dark brown; abdomen black; intersegmental membrane dark brown.
</paragraph>
<paragraph id="8BFA983DFF89FF8EFF6759E2DBEFF89A" blockId="38.[188,1400,1518,1861]" pageId="38" pageNumber="39">
<smallCapsWord id="8D1C0EE1FF89FF8EFF6759E2D882F9D8" baselines="1538,1538" box="[189,252,1519,1545]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="38" pageNumber="39">HEAD</smallCapsWord>
. Head capsule wider than long; anterior margin straight; posterior margin rounded; with emargination in front of neck; setation with 1 pair of PCS and 1 row of PMS. Epicranium with umbilicate, high density micropunctuation, setation with 2 pairs of FS and 1 pair of PFS, without OS. Gena with irregular surface, with depression from mandibular base to posterior margin of head, setation with 1 OT, MS, and POS; sockets of MS and POS umbilicate. Postgena with row of 3–5 setae in line from eye to gular suture; gular sutures not reaching posterior margin of head; anterior part of gula with 6–8 setae. Neck with foramen magnum occupying ¾ of occiput. Antenna thin, longer than head and pronotum combined (
<figureCitation id="137E84B8FF89FF8EFE645AE8DA53F920" box="[446,557,1765,1791]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[213,1380,351,1821]" captionTargetId="figure-12@39.[208,1380,265,1847]" captionTargetPageId="39" captionText="Fig. 13. Neolindus parahermani Guzman, Tokareva &amp; Żyła sp. nov., holotype, ♂ (NHMUK). A. Habitus, dorsal view. B−E. Abdominal segments, posterior margin schemes. B. Tergite VII. C. Sternite VII. D. Tergite VIII. E. Sternite VIII. F−I. Aedeagus. F. Parameral view. G. Parameral view scheme. H. Lateral view. I. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="38" pageNumber="39">Fig. 13A</figureCitation>
); antennomeres longer than wide, tomentose pubescence starting from antennomere 4; antennomere 1 as long as 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, with U-shaped emargination (
<figureCitation id="137E84B8FF89FF8EFD5B5B26DA90F89A" box="[641,750,1835,1861]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[213,1380,351,1821]" captionTargetId="figure-12@39.[208,1380,265,1847]" captionTargetPageId="39" captionText="Fig. 13. Neolindus parahermani Guzman, Tokareva &amp; Żyła sp. nov., holotype, ♂ (NHMUK). A. Habitus, dorsal view. B−E. Abdominal segments, posterior margin schemes. B. Tergite VII. C. Sternite VII. D. Tergite VIII. E. Sternite VIII. F−I. Aedeagus. F. Parameral view. G. Parameral view scheme. H. Lateral view. I. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="38" pageNumber="39">Fig. 13A</figureCitation>
; Supp. file 2).
</paragraph>
<paragraph id="8BFA983DFF89FF80FF675B60D9A8FE99" blockId="38.[189,1400,1901,1962]" lastBlockId="40.[188,1398,265,326]" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
<smallCapsWord id="8D1C0EE1FF89FF8EFF675B60D967F85A" baselines="1920,1920" box="[189,281,1901,1927]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="38" pageNumber="39">THORAX</smallCapsWord>
. Pronotum wider than long, with umbilicate micropunctures, 2 paramedial and 2 lateral rows of setae. Metasternal intercoxal process with pair of acute processes light brown at tips. Elytron longer than wide, surface of elytra with moderately dense umbilicate micropunctures in 8–10 longitudinal rows (
<figureCitation id="137E84B8FF87FF80FF1F5D21D94CFE99" box="[197,306,300,326]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[213,1380,351,1821]" captionTargetId="figure-12@39.[208,1380,265,1847]" captionTargetPageId="39" captionText="Fig. 13. Neolindus parahermani Guzman, Tokareva &amp; Żyła sp. nov., holotype, ♂ (NHMUK). A. Habitus, dorsal view. B−E. Abdominal segments, posterior margin schemes. B. Tergite VII. C. Sternite VII. D. Tergite VIII. E. Sternite VIII. F−I. Aedeagus. F. Parameral view. G. Parameral view scheme. H. Lateral view. I. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="40" pageNumber="41">Fig. 13A</figureCitation>
; Supp. file 2).
</paragraph>
<caption id="DF3AC8B5FF88FF8FFF675B69DC28F836" ID-DOI="http://doi.org/10.5281/zenodo.12550328" ID-Zenodo-Dep="12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,232,1892,1918]" targetBox="[213,1380,351,1821]" targetPageId="39">
<paragraph id="8BFA983DFF88FF8FFF675B69DC28F836" blockId="39.[187,1399,1892,2025]" pageId="39" pageNumber="40">
<emphasis id="B931442FFF88FF8FFF675B69D969F8A0" bold="true" box="[189,279,1892,1919]" pageId="39" pageNumber="40">Fig. 13.</emphasis>
<taxonomicName id="4C45E3BEFF88FF8FFEC65B68DB15F85E" authority="Guzman, Tokareva &amp; Zyla" authorityName="Guzman, Tokareva &amp; Zyla" authorityYear="2024" box="[284,875,1893,1921]" class="Insecta" family="Staphylinidae" genus="Neolindus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="parahermani" status="sp. nov.">
<emphasis id="B931442FFF88FF8FFEC65B68DA4CF8A0" box="[284,562,1893,1919]" italics="true" pageId="39" pageNumber="40">Neolindus parahermani</emphasis>
Guzman, Tokareva &amp; Żyła
</taxonomicName>
<taxonomicNameLabel id="A202F954FF88FF8FFCAA5B68DBB5F8A0" box="[880,971,1893,1919]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMUK). 
<emphasis id="B931442FFF88FF8FFB285B69DD79F8A1" bold="true" box="[1266,1287,1892,1918]" pageId="39" pageNumber="40">A</emphasis>
. Habitus, dorsal view. 
<emphasis id="B931442FFF88FF8FFE805B85D9ECF87D" bold="true" box="[346,402,1928,1954]" pageId="39" pageNumber="40">B−E</emphasis>
. Abdominal segments, posterior margin schemes. 
<emphasis id="B931442FFF88FF8FFC205B85DC70F87D" bold="true" box="[1018,1038,1928,1954]" pageId="39" pageNumber="40">B</emphasis>
. Tergite VII. 
<emphasis id="B931442FFF88FF8FFB625B85DCB3F87D" bold="true" box="[1208,1229,1928,1954]" pageId="39" pageNumber="40">C</emphasis>
. Sternite VII. 
<emphasis id="B931442FFF88FF8FFF675BA6D8ACF81A" bold="true" box="[189,210,1963,1989]" pageId="39" pageNumber="40">D</emphasis>
. Tergite VIII. 
<emphasis id="B931442FFF88FF8FFE565BA6D9DEF81A" bold="true" box="[396,416,1963,1989]" pageId="39" pageNumber="40">E</emphasis>
. Sternite VIII. 
<emphasis id="B931442FFF88FF8FFDB95BA6DAEFF81A" bold="true" box="[611,657,1963,1989]" pageId="39" pageNumber="40">F−I</emphasis>
. Aedeagus. 
<emphasis id="B931442FFF88FF8FFCF65BA6DB40F81A" bold="true" box="[812,830,1963,1989]" pageId="39" pageNumber="40">F</emphasis>
. Parameral view. 
<emphasis id="B931442FFF88FF8FFBF85BA6DC47F81A" bold="true" box="[1058,1081,1963,1989]" pageId="39" pageNumber="40">G</emphasis>
. Parameral view scheme. 
<emphasis id="B931442FFF88FF8FFF675BC3D8AAF837" bold="true" box="[189,212,1998,2024]" pageId="39" pageNumber="40">H</emphasis>
. Lateral view. 
<emphasis id="B931442FFF88FF8FFE595BC3D9F0F837" bold="true" box="[387,398,1998,2024]" pageId="39" pageNumber="40">I</emphasis>
. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BFA983DFF87FF80FF675D63DCD2FDCA" blockId="40.[189,1399,365,533]" pageId="40" pageNumber="41">
<smallCapsWord id="8D1C0EE1FF87FF80FF675D63D888FE59" baselines="385,385" box="[189,246,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="40" pageNumber="41">LEGS</smallCapsWord>
. Protibia with 5–6 well-developed combs of setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 as long as 2, mesotarsomere 3 shorter than 1 and as long as 4, mesotarsomere 5 as long as mesotarsomere 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 longer than metatarsomere 2, metatarsomere 3 shorter than metatarsomere 2, metatarsomere 4 as long metatarsomere 3, metatarsomere 5 as long as metatarsome 1.
</paragraph>
<paragraph id="8BFA983DFF87FF80FF675E31D94CFBC0" blockId="40.[188,1400,572,1055]" pageId="40" pageNumber="41">
<smallCapsWord id="8D1C0EE1FF87FF80FF675E31D952FD8B" baselines="591,591" box="[189,300,572,598]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="40" pageNumber="41">ABDOMEN</smallCapsWord>
. 
<emphasis id="B931442FFF87FF80FEE45E31D904FD89" box="[318,378,572,598]" italics="true" pageId="40" pageNumber="41">Male</emphasis>
: tergites with low-density random micropunctuation. Posterior margin of tergite VII straight (
<figureCitation id="137E84B8FF87FF80FEF35E52D9E6FDA6" box="[297,408,607,633]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[213,1380,351,1821]" captionTargetId="figure-12@39.[208,1380,265,1847]" captionTargetPageId="39" captionText="Fig. 13. Neolindus parahermani Guzman, Tokareva &amp; Żyła sp. nov., holotype, ♂ (NHMUK). A. Habitus, dorsal view. B−E. Abdominal segments, posterior margin schemes. B. Tergite VII. C. Sternite VII. D. Tergite VIII. E. Sternite VIII. F−I. Aedeagus. F. Parameral view. G. Parameral view scheme. H. Lateral view. I. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="40" pageNumber="41">Fig. 13B</figureCitation>
). Posterior margin of sternite VII with wide midline emargination delimited with 2 long teeth-like processes with acute ends (
<figureCitation id="137E84B8FF87FF80FD0B5E8FDB41FD43" box="[721,831,642,668]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[213,1380,351,1821]" captionTargetId="figure-12@39.[208,1380,265,1847]" captionTargetPageId="39" captionText="Fig. 13. Neolindus parahermani Guzman, Tokareva &amp; Żyła sp. nov., holotype, ♂ (NHMUK). A. Habitus, dorsal view. B−E. Abdominal segments, posterior margin schemes. B. Tergite VII. C. Sternite VII. D. Tergite VIII. E. Sternite VIII. F−I. Aedeagus. F. Parameral view. G. Parameral view scheme. H. Lateral view. I. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="40" pageNumber="41">Fig. 13C</figureCitation>
). Posterior margin of tergite VIII trilobed, with rounded middle lobe and triangle acute lateral lobes (
<figureCitation id="137E84B8FF87FF80FCE35EA8DBD9FD1F" box="[825,935,677,704]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[213,1380,351,1821]" captionTargetId="figure-12@39.[208,1380,265,1847]" captionTargetPageId="39" captionText="Fig. 13. Neolindus parahermani Guzman, Tokareva &amp; Żyła sp. nov., holotype, ♂ (NHMUK). A. Habitus, dorsal view. B−E. Abdominal segments, posterior margin schemes. B. Tergite VII. C. Sternite VII. D. Tergite VIII. E. Sternite VIII. F−I. Aedeagus. F. Parameral view. G. Parameral view scheme. H. Lateral view. I. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="40" pageNumber="41">Fig. 13D</figureCitation>
). Posterior margin of sternite VIII with broad midline emargination about ½ of segment length and 1 pair of longitudinal depressions on sides of midline emargination (
<figureCitation id="137E84B8FF87FF80FE1F5EE1DA51FCD9" box="[453,559,748,774]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[213,1380,351,1821]" captionTargetId="figure-12@39.[208,1380,265,1847]" captionTargetPageId="39" captionText="Fig. 13. Neolindus parahermani Guzman, Tokareva &amp; Żyła sp. nov., holotype, ♂ (NHMUK). A. Habitus, dorsal view. B−E. Abdominal segments, posterior margin schemes. B. Tergite VII. C. Sternite VII. D. Tergite VIII. E. Sternite VIII. F−I. Aedeagus. F. Parameral view. G. Parameral view scheme. H. Lateral view. I. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="40" pageNumber="41">Fig. 13E</figureCitation>
). Posterior margin of tergite IX with midlongitudinal deep emargination in ½ of segment length; aedeagus, in parameral view, median lobe slightly widened at apical part, apex with deep V-shaped emargination and with narrow flattened lateral processes adjacent to pair of lateral elongated ‘windows’ in membrane. pPMS elongated, complex, curved around longitudinal axis, with 2 apical arms almost fully fused and pointing distad and 1 apical arm short and bent laterad; bases of pPMS triangular, anchored in median foramen with internal sclerotised ring. pLS thin, slightly longer than pPMS and APS, with net-like microsculpture, forming elongate flat processes with lateral rounded lobes. APS spoon-shaped, with slightly emarginate top, shorter than pPMS (
<figureCitation id="137E84B8FF87FF80FBAF5FEFDC82FC23" box="[1141,1276,994,1020]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[213,1380,351,1821]" captionTargetId="figure-12@39.[208,1380,265,1847]" captionTargetPageId="39" captionText="Fig. 13. Neolindus parahermani Guzman, Tokareva &amp; Żyła sp. nov., holotype, ♂ (NHMUK). A. Habitus, dorsal view. B−E. Abdominal segments, posterior margin schemes. B. Tergite VII. C. Sternite VII. D. Tergite VIII. E. Sternite VIII. F−I. Aedeagus. F. Parameral view. G. Parameral view scheme. H. Lateral view. I. Lateral view scheme. Scale bars: A = 1 mm; B–I = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12550328" httpUri="https://zenodo.org/record/12550328/files/figure.png" pageId="40" pageNumber="41">Fig. 13F–I</figureCitation>
). 
<emphasis id="B931442FFF87FF80FACC5FEFDD10FC23" box="[1302,1390,994,1020]" italics="true" pageId="40" pageNumber="41">Female</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35FCBB6FF87FF80FF67584ADC4AFB6C" pageId="40" pageNumber="41" type="distribution">
<paragraph id="8BFA983DFF87FF80FF67584AD928FBBE" blockId="40.[189,342,1095,1121]" box="[189,342,1095,1121]" pageId="40" pageNumber="41">
<heading id="D0B22F51FF87FF80FF67584AD928FBBE" bold="true" box="[189,342,1095,1121]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="B931442FFF87FF80FF67584AD928FBBE" bold="true" box="[189,342,1095,1121]" pageId="40" pageNumber="41">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA983DFF87FF80FF675878DC4AFB6C" blockId="40.[188,1398,1141,1203]" pageId="40" pageNumber="41">
This species is only known from the 
<typeStatus id="54FE269FFF87FF80FDBE587BDAE9FB4F" box="[612,663,1142,1168]" pageId="40" pageNumber="41">type</typeStatus>
locality in 
<collectingCountry id="F352D8ADFF87FF80FCCD5878DBBAFB50" box="[791,964,1141,1167]" name="French Guiana" pageId="40" pageNumber="41">French Guiana</collectingCountry>
(Roura, Réserve Naturelle Régionale Trésor). It was collected at a low altitude of 
<quantity id="4CBD35D8FF87FF80FD185895DB73FB6C" box="[706,781,1176,1203]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="40" pageNumber="41" unit="m" value="225.0">225 m</quantity>
a.s.l. using window trap.
</paragraph>
</subSubSection>
</treatment>
</document>