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<mods:title id="2EED643F2AE0249F47333B8E163D1F9E">Description, redescription and revision of sixteen putatively closely related species of Echinoderes (Kinorhyncha: Cyclorhagida), with the proposition of a new species group - the Echinoderes dujardinii group</mods:title>
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<mods:namePart id="CD7CA3D48F5355A57A1D8574BB15174D">Sørensen, Martin V.</mods:namePart>
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<mods:affiliation id="7F72C953406047EC5AD9D3ED35277307">Natural History Museum of Denmark, University of Copenhagen, DK- 2100 Copenhagen, Denmark. Department of Invertebrate Zoology, Smithsonian National Museum of Natural History, Washington, DC 20560, USA. Department of Biological Science, College of Natural and Life Sciences, Daegu University, Gyeongsan 38453, Korea. Crescent International School, Bario, Govindpur, Dhanbad 828109, Jharkhand, India. Isparta University of Applied Sciences, Department of Aquaculture, 32260 Isparta, Turkey. Department of Biology, University of Copenhagen, DK- 2100 Copenhagen, Denmark. Silifke Vocational School Aquaculture Program, Mersin University, 33940 Mersin, Turkey. Faculty of Arts and Science, Kyushu University, Fukuoka 819 - 0395, Japan.</mods:affiliation>
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<mods:affiliation id="17F9039A5B67801229D6543638E7EC25">Natural History Museum of Denmark, University of Copenhagen, DK- 2100 Copenhagen, Denmark. Department of Invertebrate Zoology, Smithsonian National Museum of Natural History, Washington, DC 20560, USA. Department of Biological Science, College of Natural and Life Sciences, Daegu University, Gyeongsan 38453, Korea. Crescent International School, Bario, Govindpur, Dhanbad 828109, Jharkhand, India. Isparta University of Applied Sciences, Department of Aquaculture, 32260 Isparta, Turkey. Department of Biology, University of Copenhagen, DK- 2100 Copenhagen, Denmark. Silifke Vocational School Aquaculture Program, Mersin University, 33940 Mersin, Turkey. Faculty of Arts and Science, Kyushu University, Fukuoka 819 - 0395, Japan.</mods:affiliation>
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<mods:namePart id="00293B46A6CC56D3978E86B6D19671A9">Durucan, Furkan</mods:namePart>
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<mods:affiliation id="42EB12CC8A5ABEDEB0C478ADFFD910BB">Department of Biology, University of Copenhagen, DK- 2100 Copenhagen, Denmark.</mods:affiliation>
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<mods:affiliation id="18D81B370BD37027F2292F6A30A887C3">Department of Invertebrate Zoology, Smithsonian National Museum of Natural History, Washington, DC 20560, USA.</mods:affiliation>
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<treatment id="03C79270FFB0573AB111F93E19DBFCC7" ID-DOI="http://doi.org/10.5281/zenodo.4420027" ID-GBIF-Taxon="176434266" ID-Zenodo-Dep="4420027" LSID="urn:lsid:plazi:treatment:03C79270FFB0573AB111F93E19DBFCC7" httpUri="http://treatment.plazi.org/id/03C79270FFB0573AB111F93E19DBFCC7" lastPageId="38" lastPageNumber="39" pageId="31" pageNumber="32">
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<heading id="D099940AFFB05703B111F93E19A9F96A" box="[601,986,1761,1788]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="32" reason="2">
<taxonomicName id="4C6E58E5FFB05703B111F93E19A9F96A" authority="Lang, 1949" authorityName="Lang" authorityYear="1949" box="[601,986,1761,1788]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="31" pageNumber="32" phylum="Cephalorhyncha" rank="species" species="pilosus">
<emphasis id="B91AFF74FFB05703B111F93E193EF96A" bold="true" box="[601,845,1761,1788]" italics="true" pageId="31" pageNumber="32">Echinoderes pilosus</emphasis>
<bibRefCitation id="EFFF5E97FFB05703B01CF93D19A9F96A" author="Lang K." box="[852,986,1762,1788]" pageId="31" pageNumber="32" pagination="1 - 22" refId="ref56538" refString="Lang K. 1949. Echinoderida. In: Odhner N. H. (ed.) Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, Vol. 4: 1 - 22. P. A. Norstedt &amp; Soner, Stockholm." type="journal article" year="1949">Lang, 1949</bibRefCitation>
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<paragraph id="8BD12366FFB05703B1D9F8DA19D1F889" blockId="31.[601,986,1761,1823]" box="[657,930,1797,1823]" pageId="31" pageNumber="32">
<figureCitation id="13553FE3FFB05703B1D9F8DA197AF889" box="[657,777,1797,1823]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="33.[189,232,1742,1768]" captionStartId-1="34.[189,232,1705,1731]" captionTargetBox-0="[263,1324,268,1704]" captionTargetBox-1="[287,1300,270,1661]" captionTargetId-0="figure-11@33.[260,1328,265,1706]" captionTargetId-1="figure-13@34.[284,1304,265,1665]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionText-0="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." captionText-1="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi-0="http://doi.org/10.5281/zenodo.4418999" figureDoi-1="http://doi.org/10.5281/zenodo.4419001" httpUri-0="https://zenodo.org/record/4418999/files/figure.png" httpUri-1="https://zenodo.org/record/4419001/files/figure.png" pageId="31" pageNumber="32">Figs 910</figureCitation>
;
<tableCitation id="C6EC16DDFFB05703B05CF8DA19D1F889" box="[788,930,1797,1823]" captionStart-0="Table 9" captionStart-1="Table 10" captionStartId-0="35.[189,258,265,291]" captionStartId-1="37.[189,258,265,291]" captionTargetPageId-0="35" captionTargetPageId-1="37" captionText-0="Table 9. Measurements from light microscopy for lectotype and nine paralectotypes of Echinoderes pilosus Lang, 1949 from Grytviken, South Georgia Island (in µm), including number of measured specimens (n) and standard deviation (SD)." captionText-1="Table 10. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes pilosus Lang, 1949." pageId="31" pageNumber="32">Tables 910</tableCitation>
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<heading id="D099940AFFB05703B3F5F8991BDCF8F6" bold="true" box="[189,431,1862,1888]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B91AFF74FFB05703B3F5F8991BDCF8F6" bold="true" box="[189,431,1862,1888]" pageId="31" pageNumber="32">Emended diagnosis</emphasis>
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<taxonomicName id="4C6E58E5FFB05703B3F5F8AA1B3CF819" authorityName="Dujardinii Claparede" authorityYear="1863" box="[189,335,1909,1935]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="31" pageNumber="32" phylum="Cephalorhyncha" rank="genus">
<emphasis id="B91AFF74FFB05703B3F5F8AA1B3CF819" box="[189,335,1909,1935]" italics="true" pageId="31" pageNumber="32">Echinoderes</emphasis>
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with middorsal spines on segments 4 to 8, and lateroventral spines on segments 6 to 9; all middorsal spines slender and flexible and projecting well beyond the posterior margin of their respective segments. Long and slender tubes present in ventrolateral positions on segment 2, lateroventral positions on segment 5, and in laterodorsal positions on 10. Incomplete midventral, intracuticular fissure present on anterior half of segment 2. Two middorsal, longitudinally aligned glandular cell outlets
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on segment 1. Minute glandular cell outlets
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subdorsal positions on segment 2, and in laterodorsal positions on segments 8 and 9. Tergal extensions of segment 11 short and pointed; margin of sternal extensions straight, with ventrolateral seta-like tuft of extended fringe tips. Females with female papillae with tubular substructure in ventrolateral positions on segment 7 and ventromedial positions on segment 8; males with ventrolateral sensory spots on segment 10.
</paragraph>
</subSubSection>
<subSubSection id="C37470EDFF8F573CB3F5FDC21946FCCE" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BD12366FF8F573CB3F5FDC21BD9FDA1" blockId="32.[189,426,541,567]" box="[189,426,541,567]" pageId="32" pageNumber="33">
<heading id="D099940AFF8F573CB3F5FDC21BD9FDA1" bold="true" box="[189,426,541,567]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B91AFF74FF8F573CB3F5FDC21BD9FDA1" bold="true" box="[189,426,541,567]" pageId="32" pageNumber="33">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD12366FF8F573CB3A2FD93184BFDF1" blockId="32.[189,1399,588,685]" box="[234,568,588,615]" pageId="32" pageNumber="33">
<emphasis id="B91AFF74FF8F573CB3A2FD931B15FDF0" bold="true" box="[234,358,588,614]" pageId="32" pageNumber="33">
<typeStatus id="54D59DC4FF8F573CB3A2FD931B15FDF0" box="[234,358,588,614]" pageId="32" pageNumber="33" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated)
</paragraph>
<paragraph id="8BD12366FF8F573CB3F5FDB01F41FD3B" blockId="32.[189,1399,588,685]" pageId="32" pageNumber="33">
<materialsCitation id="3B06293BFF8F573CB3F5FDB01F41FD3B" ID-GBIF-Occurrence="3009108346" collectingDate="1902-05-22" collectingDateMax="1902-05-23" collectingDateMin="1902-05-22" collectionCode="SMNH" country="South Georgia and the South Sandwich Islands" county="SOUTH GEORGIA ISLANDS" latitude="-54.283054" location="Grytviken" longLatPrecision="17" longitude="-36.508057" municipality="Swedish Antarctic Expedition in Grytviken" pageId="32" pageNumber="33" specimenCode="SMNH 3930" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<collectingCounty id="62B05BEAFF8F573CB3F5FDB0185FFD1F" box="[189,556,623,649]" pageId="32" pageNumber="33">SOUTH GEORGIA ISLANDS</collectingCounty>
<specimenCount id="9D68E8EFFF8F573CB10CFDAF1828FD1C" box="[580,603,624,650]" pageId="32" pageNumber="33" type="female"></specimenCount>
;
<location id="8EB175BDFF8F573CB123FDB01895FD1F" LSID="urn:lsid:plazi:treatment:03C79270FFB0573AB111F93E19DBFCC7:8EB175BDFF8F573CB123FDB01895FD1F" box="[619,742,623,649]" country="South Georgia and the South Sandwich Islands" county="SOUTH GEORGIA ISLANDS" latitude="-54.283054" longLatPrecision="17" longitude="-36.508057" municipality="Swedish Antarctic Expedition in Grytviken" name="Grytviken" pageId="32" pageNumber="33">Grytviken</location>
;
<geoCoordinate id="EE5A45A1FF8F573CB1B9FDB119F2FD1F" box="[753,897,622,650]" degrees="54" direction="south" minutes="16" orientation="latitude" pageId="32" pageNumber="33" precision="15" seconds="59" value="-54.283054">54°1659″ S</geoCoordinate>
,
<geoCoordinate id="EE5A45A1FF8F573CB0C6FDB11E5AFD1C" box="[910,1065,622,650]" degrees="36" direction="west" minutes="30" orientation="longitude" pageId="32" pageNumber="33" precision="15" seconds="29" value="-36.508057">36°3029″ W</geoCoordinate>
;
<date id="FFD005A6FF8F573CB77DFDB01F73FD1F" box="[1077,1280,623,650]" pageId="32" pageNumber="33" value="1902-05-22" valueMax="1902-05-23" valueMin="1902-05-22">
<collectingDate id="EF94FC4EFF8F573CB77DFDB01F73FD1F" box="[1077,1280,623,650]" pageId="32" pageNumber="33" value="1902-05-22" valueMax="1902-05-23" valueMin="1902-05-22">2223 May 1902</collectingDate>
</date>
; collected during the Swedish Antarctic Expedition in Grytviken;
<specimenCode id="DBC88B1DFF8F573CB00EFD4C1993FD3B" box="[838,992,659,685]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="32" pageNumber="33">SMNH 3930</specimenCode>
. Specimen mounted for LM.
</materialsCitation>
</paragraph>
<paragraph id="8BD12366FF8F573CB3A2FD0B1BD1FD78" blockId="32.[189,1398,724,858]" box="[234,418,724,750]" pageId="32" pageNumber="33">
<emphasis id="B91AFF74FF8F573CB3A2FD0B1BD1FD78" bold="true" box="[234,418,724,750]" pageId="32" pageNumber="33">
<typeStatus id="54D59DC4FF8F573CB3A2FD0B1BD1FD78" box="[234,418,724,750]" pageId="32" pageNumber="33" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD12366FF8F573CB3F5FD281946FCCE" blockId="32.[189,1398,724,858]" pageId="32" pageNumber="33">
<materialsCitation id="3B06293BFF8F573CB3F5FD281943FCCE" ID-GBIF-Occurrence="3009108329" collectingDate="1902-05-22" collectingDateMax="1902-05-23" collectingDateMin="1902-05-22" collectionCode="SMNH" country="South Georgia and the South Sandwich Islands" county="SOUTH GEORGE ISLANDS" latitude="-54.283054" location="SOUTH GEORGE ISLANDS" longLatPrecision="17" longitude="-36.508057" municipality="Swedish Antarctic Expedition in Grytviken" pageId="32" pageNumber="33" specimenCode="SMNH-1007, SMNH-9256, 9282" specimenCount="58" specimenCount-female="24" specimenCount-male="31" typeStatus="paralectotype">
<collectingCounty id="62B05BEAFF8F573CB3F5FD281851FC87" box="[189,546,759,785]" pageId="32" pageNumber="33">SOUTH GEORGE ISLANDS</collectingCounty>
<specimenCount id="9D68E8EFFF8F573CB174FD2818FDFC84" box="[572,654,759,786]" pageId="32" pageNumber="33" type="female">15 ♀♀</specimenCount>
,
<specimenCount id="9D68E8EFFF8F573CB1D4FD281899FC82" box="[668,746,759,788]" pageId="32" pageNumber="33" type="male">18 ♂♂</specimenCount>
,
<specimenCount id="9D68E8EFFF8F573CB1B5FD28197FFC87" box="[765,780,759,785]" pageId="32" pageNumber="33">3</specimenCount>
uncertain sex; same collection data as for lectotype;
<specimenCode id="DBC88B1DFF8F573CB3F5FCC51B2AFCA2" box="[189,345,794,820]" collectionCode="SMNH-" pageId="32" pageNumber="33">SMNH-1007</specimenCode>
,
<specimenCode id="DBC88B1DFF8F573CB22DFCC51B8DFCA3" box="[357,510,794,821]" collectionCode="SMNH-" pageId="32" pageNumber="33">SMNH-9256</specimenCode>
to
<specimenCode id="DBC88B1DFF8F573CB16BFCC51812FCA2" box="[547,609,794,820]" collectionCode="SMNH-" pageId="32" pageNumber="33">9282</specimenCode>
.
<specimenCode id="DBC88B1DFF8F573CB124FCC5197BFCA2" box="[620,776,794,820]" collectionCode="SMNH-" pageId="32" pageNumber="33">SMNH-1007</specimenCode>
,
<specimenCode id="DBC88B1DFF8F573CB05BFCC519DEFCA3" box="[787,941,794,821]" collectionCode="SMNH-" pageId="32" pageNumber="33">SMNH-9256</specimenCode>
to
<specimenCode id="DBC88B1DFF8F573CB09AFCC51E63FCA2" box="[978,1040,794,820]" collectionCode="SMNH-" pageId="32" pageNumber="33">9282</specimenCode>
.
<specimenCount id="9D68E8EFFF8F573CB753FCC51E2EFCA3" box="[1051,1117,794,821]" pageId="32" pageNumber="33" type="female">9 ♀♀</specimenCount>
,
<specimenCount id="9D68E8EFFF8F573CB723FCC51EC8FCA1" box="[1131,1211,794,823]" pageId="32" pageNumber="33" type="male">13 ♂♂</specimenCount>
and 3 uncertain sex mounted for LM; 6 ♀♀, 5 ♂♂ mounted for SEM
</materialsCitation>
;
</paragraph>
</subSubSection>
<subSubSection id="C37470EDFF8F573CB3F5FCA018A5FB2C" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BD12366FF8F573CB3F5FCA01B5CFC0F" blockId="32.[189,303,895,921]" box="[189,303,895,921]" pageId="32" pageNumber="33">
<heading id="D099940AFF8F573CB3F5FCA01B5CFC0F" bold="true" box="[189,303,895,921]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B91AFF74FF8F573CB3F5FCA01B5CFC0F" bold="true" box="[189,303,895,921]" pageId="32" pageNumber="33">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD12366FF8F573CB3F5FC721B01FB98" blockId="32.[188,1399,941,1038]" pageId="32" pageNumber="33">
<bibRefCitation id="EFFF5E97FF8F573CB3F5FC721B07FC5E" author="Higgins R. P." box="[189,372,941,968]" pageId="32" pageNumber="33" pagination="399 - 405" refId="ref56187" refString="Higgins R. P. 1986. Redescription of Echinoderes pilosus (Kinorhyncha: Cyclorhagida). Proceedings of the Biological Society of Washington 99: 399 - 405. Available from https: // www. biodiversitylibrary. org / part / 45462 # / summary [accessed 27 Nov. 2020]." type="journal article" year="1986">Higgins (1986)</bibRefCitation>
mentions that he accessed the
<typeStatus id="54D59DC4FF8F573CB1B2FC71195EFC5E" box="[762,813,942,968]" pageId="32" pageNumber="33">type</typeStatus>
material of
<taxonomicName id="4C6E58E5FF8F573CB082FC711EBAFC5E" authority="in SMNH" authorityName="in SMNH" box="[970,1225,941,968]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="32" pageNumber="33" phylum="Cephalorhyncha" rank="species" species="pilosus">
<emphasis id="B91AFF74FF8F573CB082FC711E31FC51" box="[970,1090,941,967]" italics="true" pageId="32" pageNumber="33">E. pilosus</emphasis>
in SMNH
</taxonomicName>
, mounted and observed two paratypic specimens. However, such mounted specimens were found in neither the SMNH nor the USNM.
</paragraph>
<paragraph id="8BD12366FF8F573CB3F5FBEA18A5FB2C" blockId="32.[189,1399,1077,1210]" pageId="32" pageNumber="33">
After examining the
<typeStatus id="54D59DC4FF8F573CB2F5FBE91857FBC6" box="[445,548,1078,1104]" pageId="32" pageNumber="33" type="syntype">syntypes</typeStatus>
and providing the present redescription we choose, in the interest of assuring future taxonomic stability, to designate one of the female
<typeStatus id="54D59DC4FF8F573CB08FFB861E51FBE5" box="[967,1058,1113,1139]" pageId="32" pageNumber="33" type="syntype">syntype</typeStatus>
as
<typeStatus id="54D59DC4FF8F573CB701FB871EC6FBE4" box="[1097,1205,1112,1138]" pageId="32" pageNumber="33" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="4C6E58E5FF8F573CB7ADFB871B63FB00" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="32" pageNumber="33" phylum="Cephalorhyncha" rank="species" species="pilosus">
<emphasis id="B91AFF74FF8F573CB7ADFB871B63FB00" italics="true" pageId="32" pageNumber="33">Echinoderes pilosus</emphasis>
</taxonomicName>
with the catalogue number SMNH-3930, and consequently designate the remaining specimens as
<typeStatus id="54D59DC4FF8F573CB396FB401BF4FB2F" box="[222,391,1183,1209]" pageId="32" pageNumber="33" type="paralectotype">paralectotypes</typeStatus>
(SMNH-1007, 92569282).
</paragraph>
</subSubSection>
<subSubSection id="C37470EDFF8F573AB3F5FB3F19DBFCC7" lastPageId="38" lastPageNumber="39" pageId="32" pageNumber="33" type="description">
<paragraph id="8BD12366FF8F573CB3F5FB3F1B3DFB6C" blockId="32.[189,334,1248,1274]" box="[189,334,1248,1274]" pageId="32" pageNumber="33">
<heading id="D099940AFF8F573CB3F5FB3F1B3DFB6C" bold="true" box="[189,334,1248,1274]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B91AFF74FF8F573CB3F5FB3F1B3DFB6C" bold="true" box="[189,334,1248,1274]" pageId="32" pageNumber="33">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD12366FF8F573CB3F5FAD0197CFA04" blockId="32.[188,1401,1295,1427]" pageId="32" pageNumber="33">
The species was described by
<bibRefCitation id="EFFF5E97FF8F573CB154FAD018DCFABF" author="Lang K." box="[540,687,1295,1321]" pageId="32" pageNumber="33" pagination="1 - 22" refId="ref56538" refString="Lang K. 1949. Echinoderida. In: Odhner N. H. (ed.) Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, Vol. 4: 1 - 22. P. A. Norstedt &amp; Soner, Stockholm." type="journal article" year="1949">Lang (1949)</bibRefCitation>
, and more recently redescribed by
<bibRefCitation id="EFFF5E97FF8F573CB70DFAD01E88FABC" author="Higgins R. P." box="[1093,1275,1295,1322]" pageId="32" pageNumber="33" pagination="399 - 405" refId="ref56187" refString="Higgins R. P. 1986. Redescription of Echinoderes pilosus (Kinorhyncha: Cyclorhagida). Proceedings of the Biological Society of Washington 99: 399 - 405. Available from https: // www. biodiversitylibrary. org / part / 45462 # / summary [accessed 27 Nov. 2020]." type="journal article" year="1986">Higgins (1986)</bibRefCitation>
. However, since information from SEM not previously has been available for the species, and since the present examinations with LM and SEM resulted in substantial new information about the morphology of the species, we are here providing a full redescription.
</paragraph>
<paragraph id="8BD12366FF8F573CB3F5FA651E65F931" blockId="32.[188,1400,1466,1704]" pageId="32" pageNumber="33">
Adults with head, neck and eleven trunk segments. The anterior part of the trunk is relatively stout, and maximum width is reached around segments 6 and 7 (
<figureCitation id="13553FE3FF8F573CB074FA0219F1FA61" box="[828,898,1501,1527]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="32" pageNumber="33">Fig. 9</figureCitation>
AB). From segment 8 the segments taper significantly, giving the species an appearance of a pointed hind end (
<figureCitation id="13553FE3FF8F573CB746F9DF1E26F98D" box="[1038,1109,1536,1563]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="32" pageNumber="33">Fig. 9</figureCitation>
AB). The trunk cuticle appears thin in SEM specimens (
<figureCitation id="13553FE3FF8F573CB10DF9FC18C0F9A8" box="[581,691,1571,1598]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="32" pageNumber="33">Fig. 10A</figureCitation>
), and tends to collapse posterior to the stronger pachycycli. Secondary pectinate fringe present near anterior segment margin on segments 2 to 10. For complete overview of measurements and dimensions, see
<tableCitation id="C6EC16DDFF8F573CB1A3F9B51931F912" box="[747,834,1642,1668]" captionStart="Table 9" captionStartId="35.[189,258,265,291]" captionTargetPageId="35" captionText="Table 9. Measurements from light microscopy for lectotype and nine paralectotypes of Echinoderes pilosus Lang, 1949 from Grytviken, South Georgia Island (in µm), including number of measured specimens (n) and standard deviation (SD)." pageId="32" pageNumber="33">Table 9</tableCitation>
. Distribution of cuticular structures, i.e., sensory spots, glandular cell outlets, spines and tubes, is summarized in
<tableCitation id="C6EC16DDFF8F573CB0E1F9521E60F931" box="[937,1043,1677,1703]" captionStart="Table 10" captionStartId="37.[189,258,265,291]" captionTargetPageId="37" captionText="Table 10. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes pilosus Lang, 1949." pageId="32" pageNumber="33">Table 10</tableCitation>
.
</paragraph>
<paragraph id="8BD12366FF8F573CB3F5F9101BA2F8C4" blockId="32.[189,1399,1743,1875]" pageId="32" pageNumber="33">The head consists of a retractable mouth cone and an introvert. Inner oral styles could not be examined. Nine outer oral styles are present; nine tufts with ca. six long, rigid and distally bifurcated tips in each, attach basally on the mouth cone, posterior to each outer oral style. The exact arrangement of scalids could not be examined.</paragraph>
<paragraph id="8BD12366FF8F573CB3F5F8A51F04F821" blockId="32.[189,1399,1914,1975]" pageId="32" pageNumber="33">
The neck has 16 placids, measuring
<quantity id="4C968E83FF8F573CB134F8A518BFF802" box="[636,716,1914,1940]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="32" pageNumber="33" unit="mm" value="19.0">19 µm</quantity>
in length. The midventral placid is broadest, measuring
<quantity id="4C968E83FF8F573CB3F5F8421B7FF821" box="[189,268,1949,1975]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="32" pageNumber="33" unit="mm" value="17.0">17 µm</quantity>
in width at its base, whereas all other are narrower, measuring
<quantity id="4C968E83FF8F573CB744F8421E28F821" box="[1036,1115,1949,1975]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="33" unit="mm" value="10.0">10 µm</quantity>
in width at their bases.
</paragraph>
<caption id="DF1173EEFF8E573DB3F5F9111928F848" ID-DOI="http://doi.org/10.5281/zenodo.4418999" ID-Zenodo-Dep="4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="33" pageNumber="34" startId="33.[189,232,1742,1768]" targetBox="[263,1324,268,1704]" targetPageId="33">
<paragraph id="8BD12366FF8E573DB3F5F9111928F848" blockId="33.[188,1399,1742,2015]" pageId="33" pageNumber="34">
<emphasis id="B91AFF74FF8E573DB3F5F9111B7DF97E" bold="true" box="[189,270,1742,1768]" pageId="33" pageNumber="34">Fig. 9.</emphasis>
Light micrographs showing overviews and details of
<taxonomicName id="4C6E58E5FF8E573DB0EAF9111F5AF97E" authority="Lang, 1949" authorityName="Lang" authorityYear="1949" box="[930,1321,1742,1769]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="33" pageNumber="34" phylum="Cephalorhyncha" rank="species" species="pilosus">
<emphasis id="B91AFF74FF8E573DB0EAF9111EE2F97E" box="[930,1169,1742,1768]" italics="true" pageId="33" pageNumber="34">Echinoderes pilosus</emphasis>
<bibRefCitation id="EFFF5E97FF8E573DB7D4F9101F5AF97E" author="Lang K." box="[1180,1321,1742,1769]" pageId="33" pageNumber="34" pagination="1 - 22" refId="ref56538" refString="Lang K. 1949. Echinoderida. In: Odhner N. H. (ed.) Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, Vol. 4: 1 - 22. P. A. Norstedt &amp; Soner, Stockholm." type="journal article" year="1949">Lang, 1949</bibRefCitation>
</taxonomicName>
.
<emphasis id="B91AFF74FF8E573DB670F9111A85F89D" bold="true" pageId="33" pageNumber="34">AB, GL</emphasis>
. Lectotype, ♀ (SMNH-3930a).
<emphasis id="B91AFF74FF8E573DB130F92E18ABF89D" bold="true" box="[632,728,1777,1803]" pageId="33" pageNumber="34">CF, M</emphasis>
. Paralectotype, ♂ (SMNH-3930b).
<emphasis id="B91AFF74FF8E573DB7CEF92E1EE8F89D" bold="true" box="[1158,1179,1777,1803]" pageId="33" pageNumber="34">A</emphasis>
. Dorsal overview.
<emphasis id="B91AFF74FF8E573DB3F7F8CA1AA3F8B9" bold="true" box="[191,208,1813,1839]" pageId="33" pageNumber="34">B</emphasis>
. Ventral overview.
<emphasis id="B91AFF74FF8E573DB2FDF8CB1BB9F8B8" bold="true" box="[437,458,1812,1838]" pageId="33" pageNumber="34">C</emphasis>
. Segments 1 to 2, dorsal view.
<emphasis id="B91AFF74FF8E573DB074F8CA1922F8B9" bold="true" box="[828,849,1813,1839]" pageId="33" pageNumber="34">D</emphasis>
. Segments 1 to 2, ventral view.
<emphasis id="B91AFF74FF8E573DB784F8CA1EACF8B9" bold="true" box="[1228,1247,1813,1839]" pageId="33" pageNumber="34">E</emphasis>
. Segments 1 to 3, dorsal view.
<emphasis id="B91AFF74FF8E573DB2DBF8E71BD7F8C4" bold="true" box="[403,420,1848,1874]" pageId="33" pageNumber="34">F</emphasis>
. Segments 2 to 4, ventral view.
<emphasis id="B91AFF74FF8E573DB06DF8E81948F8C7" bold="true" box="[805,827,1847,1873]" pageId="33" pageNumber="34">G</emphasis>
. Segments 6 to 7, ventral view, showing female sexual dimorphism.
<emphasis id="B91AFF74FF8E573DB2E1F8841BB3F8E3" bold="true" box="[425,448,1883,1909]" pageId="33" pageNumber="34">H</emphasis>
. Segments 3 to 5, dorsal view.
<emphasis id="B91AFF74FF8E573DB06EF8841942F8E3" bold="true" box="[806,817,1883,1909]" pageId="33" pageNumber="34">I</emphasis>
. Segments 8 to 9, dorsal view.
<emphasis id="B91AFF74FF8E573DB7D1F8841EDBF8E3" bold="true" box="[1177,1192,1883,1909]" pageId="33" pageNumber="34">J</emphasis>
. Segments 7 to 8, ventral view, showing female sexual dimorphism.
<emphasis id="B91AFF74FF8E573DB052F8A1195CF80E" bold="true" box="[794,815,1918,1944]" pageId="33" pageNumber="34">K</emphasis>
. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.
<emphasis id="B91AFF74FF8E573DB2B6F87E1861F82D" bold="true" box="[510,530,1953,1979]" pageId="33" pageNumber="34">L</emphasis>
. Segments 8 to 9, dorsal view but deeper focal plane than (I).
<emphasis id="B91AFF74FF8E573DB792F87E1E85F82D" bold="true" box="[1242,1270,1953,1979]" pageId="33" pageNumber="34">M</emphasis>
. Segments 10 to 11, ventral view, showing male sexual dimorphism.
</paragraph>
</caption>
<caption id="DF1173EEFF8D573EB3F5F9761F44F84B" ID-DOI="http://doi.org/10.5281/zenodo.4419001" ID-Zenodo-Dep="4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="34" pageNumber="35" startId="34.[189,232,1705,1731]" targetBox="[287,1300,270,1661]" targetPageId="34">
<paragraph id="8BD12366FF8D573EB3F5F9761F44F84B" blockId="34.[188,1400,1705,2014]" pageId="34" pageNumber="35">
<emphasis id="B91AFF74FF8D573EB3F5F9761B6FF952" bold="true" box="[189,284,1705,1732]" pageId="34" pageNumber="35">Fig. 10.</emphasis>
Scanning electron micrographs showing overviews and details of
<taxonomicName id="4C6E58E5FF8D573EB775F9751A88F971" authority="Lang, 1949" authorityName="Lang" authorityYear="1949" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="34" pageNumber="35" phylum="Cephalorhyncha" rank="species" species="pilosus">
<emphasis id="B91AFF74FF8D573EB775F9751F58F952" box="[1085,1323,1706,1732]" italics="true" pageId="34" pageNumber="35">Echinoderes pilosus</emphasis>
<bibRefCitation id="EFFF5E97FF8D573EB67DF9751A88F971" author="Lang K." pageId="34" pageNumber="35" pagination="1 - 22" refId="ref56538" refString="Lang K. 1949. Echinoderida. In: Odhner N. H. (ed.) Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, Vol. 4: 1 - 22. P. A. Norstedt &amp; Soner, Stockholm." type="journal article" year="1949">Lang, 1949</bibRefCitation>
</taxonomicName>
.
<emphasis id="B91AFF74FF8D573EB24DF9121B2AF971" bold="true" box="[261,345,1741,1767]" pageId="34" pageNumber="35">AJ, L</emphasis>
. ♀ (SMNH-9262, SMNH-9281).
<emphasis id="B91AFF74FF8D573EB1ACF9121888F971" bold="true" box="[740,763,1741,1767]" pageId="34" pageNumber="35">K</emphasis>
.♂ (SMNH-9262).
<emphasis id="B91AFF74FF8D573EB091F912199DF971" bold="true" box="[985,1006,1741,1767]" pageId="34" pageNumber="35">A</emphasis>
. Lateral overview.
<emphasis id="B91AFF74FF8D573EB785F9121E92F971" bold="true" box="[1229,1249,1741,1767]" pageId="34" pageNumber="35">B</emphasis>
. Segments 1 to 2, dorsal view.
<emphasis id="B91AFF74FF8D573EB2D1F92F1BDFF89C" bold="true" box="[409,428,1776,1802]" pageId="34" pageNumber="35">C</emphasis>
. Segments 1 to 2, ventral view.
<emphasis id="B91AFF74FF8D573EB07EF92F193AF89C" bold="true" box="[822,841,1776,1802]" pageId="34" pageNumber="35">D</emphasis>
. Segments 3 to 4, dorsal view.
<emphasis id="B91AFF74FF8D573EB782F92F1EA8F89C" bold="true" box="[1226,1243,1776,1802]" pageId="34" pageNumber="35">E</emphasis>
. Segments 5 to 6, lateral view.
<emphasis id="B91AFF74FF8D573EB2D2F8CC1BD9F8BB" bold="true" box="[410,426,1811,1837]" pageId="34" pageNumber="35">F</emphasis>
. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm.
<emphasis id="B91AFF74FF8D573EB0ACF8E91988F8C6" bold="true" box="[996,1019,1846,1872]" pageId="34" pageNumber="35">G</emphasis>
. Segments 8 to 9, dorsal view.
<emphasis id="B91AFF74FF8D573EB3F5F8851AA7F8E2" bold="true" box="[189,212,1882,1908]" pageId="34" pageNumber="35">H</emphasis>
. Segments 8 to 9, laterodorsal view.
<emphasis id="B91AFF74FF8D573EB1E7F88518C9F8E2" bold="true" box="[687,698,1882,1908]" pageId="34" pageNumber="35">I</emphasis>
. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm.
<emphasis id="B91AFF74FF8D573EB7AFF8A21E85F801" bold="true" box="[1255,1270,1917,1943]" pageId="34" pageNumber="35">J</emphasis>
. Segments 10 to 11 dorsal view, showing female sexual dimorphism.
<emphasis id="B91AFF74FF8D573EB025F87F19F0F82C" bold="true" box="[877,899,1952,1978]" pageId="34" pageNumber="35">K</emphasis>
. Segments 10 to 11 ventral view, showing male sexual dimorphism.
<emphasis id="B91AFF74FF8D573EB2A4F81C1873F84B" bold="true" box="[492,512,1987,2013]" pageId="34" pageNumber="35">L</emphasis>
. Segments 10 to 11 ventral view, showing female sexual dimorphism.
</paragraph>
</caption>
<caption id="DF1173EEFF8C573FB3F5FED618CAFEFC" pageId="35" pageNumber="36" targetBox="[516,1053,416,1953]" targetIsTable="true" targetPageId="35">
<paragraph id="8BD12366FF8C573FB3F5FED618CAFEFC" blockId="35.[189,1399,265,362]" pageId="35" pageNumber="36">
<emphasis id="B91AFF74FF8C573FB3F5FED61B51FEB5" bold="true" box="[189,290,265,291]" pageId="35" pageNumber="36">Table 9.</emphasis>
Measurements from light microscopy for lectotype and nine paralectotypes of
<taxonomicName id="4C6E58E5FF8C573FB7ADFED61BD4FED0" authority="Lang, 1949" authorityName="Lang" authorityYear="1949" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="35" pageNumber="36" phylum="Cephalorhyncha" rank="species" species="pilosus">
<emphasis id="B91AFF74FF8C573FB7ADFED61B63FED0" italics="true" pageId="35" pageNumber="36">Echinoderes pilosus</emphasis>
<bibRefCitation id="EFFF5E97FF8C573FB254FEF21BD4FED0" author="Lang K." box="[284,423,300,327]" pageId="35" pageNumber="36" pagination="1 - 22" refId="ref56538" refString="Lang K. 1949. Echinoderida. In: Odhner N. H. (ed.) Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, Vol. 4: 1 - 22. P. A. Norstedt &amp; Soner, Stockholm." type="journal article" year="1949">Lang, 1949</bibRefCitation>
</taxonomicName>
from Grytviken, South Georgia Island (in µm), including number of measured specimens (
<emphasis id="B91AFF74FF8C573FB200FE8F1B24FEFF" box="[328,343,336,361]" italics="true" pageId="35" pageNumber="36">n</emphasis>
) and standard deviation (SD).
</paragraph>
</caption>
<paragraph id="8BD12366FF8C573FB14CFE7F1E67F837" pageId="35" pageNumber="36">
<table id="F96ED1C6FF8CA8E3B14CFE7F1E6EF837" box="[516,1053,416,1953]" colsContinueFrom="30.[516,1053,430,1973]" colsContinueIn="63.[516,1065,392,1988]" gridcols="5" gridrows="5" pageId="35" pageNumber="36">
<tr id="355E2124FF8CA8E3B14CFE7F1E6EFE25" box="[516,1053,416,435]" gridrow="0" pageId="35" pageNumber="36">
<th id="768F4858FF8CA8E3B14CFE7F1814FE25" box="[516,615,416,435]" gridcol="0" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B91AFF74FF8C573FB14CFE7F1811FE25" bold="true" box="[516,610,416,435]" pageId="35" pageNumber="36">Character</emphasis>
</th>
<th id="768F4858FF8CA8E3B1CEFE7F18E8FE25" box="[646,667,416,435]" gridcol="1" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B91AFF74FF8C573FB1C3FE7F18E4FE25" bold="true" box="[651,663,416,435]" italics="true" pageId="35" pageNumber="36">n</emphasis>
</th>
<th id="768F4858FF8CA8E3B186FE7F1945FE25" box="[718,822,416,435]" gridcol="2" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B91AFF74FF8C573FB1ADFE7F196CFE25" bold="true" box="[741,799,416,435]" pageId="35" pageNumber="36">Range</emphasis>
</th>
<th id="768F4858FF8CA8E3B03CFE7F19DFFE25" box="[884,940,416,435]" gridcol="3" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B91AFF74FF8C573FB03EFE7F19D9FE25" bold="true" box="[886,938,416,435]" pageId="35" pageNumber="36">Mean</emphasis>
</th>
<th id="768F4858FF8CA8E3B0AEFE7F1E6EFE25" box="[998,1053,416,435]" gridcol="4" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B91AFF74FF8C573FB0BCFE7F1E7CFE25" bold="true" box="[1012,1039,416,435]" pageId="35" pageNumber="36">SD</emphasis>
</th>
</tr>
<tr id="355E2124FF8CA8E3B14CFE2A1E6EFB02" box="[516,1053,501,1172]" gridrow="1" pageId="35" pageNumber="36">
<th id="768F4858FF8CA8E3B14CFE2A1814FB02" box="[516,615,501,1172]" gridcol="0" gridrow="1" pageId="35" pageNumber="36">TL MSW-8 MSW-8/TL SW-10 SW-10/TL S1 S2 S3 S4 S5 S6 S7 S8 S9 S10 S11</th>
<td id="768F4858FF8CA8E3B1CEFE2A18E8FB02" box="[646,667,501,1172]" gridcol="1" gridrow="1" pageId="35" pageNumber="36">14 14 11 15 11 14 14 15 15 15 17 18 17 16 15 15</td>
<td id="768F4858FF8CA8E3B186FE2A1945FB02" box="[718,822,501,1172]" gridcol="2" gridrow="1" pageId="35" pageNumber="36">343440 7990 19.025.1% 6578 15.420.0% 3654 2741 2633 2737 3138 3343 3549 4052 4255 5161 3040</td>
<td id="768F4858FF8CA8E3B03CFE2A19DFFB02" box="[884,940,501,1172]" gridcol="3" gridrow="1" pageId="35" pageNumber="36">389 86 22.5% 71 18.6% 44 34 30 33 35 38 42 46 49 55 34</td>
<td id="768F4858FF8CA8E3B0AEFE2A1E6EFB02" box="[998,1053,501,1172]" gridcol="4" gridrow="1" pageId="35" pageNumber="36">28.83 3.09 1.86% 4.24% 1.44% 5,70 4,43 1,99 2,93 1,93 2,76 3,60 3,46 3,07 3,70 3,40</td>
</tr>
<tr id="355E2124FF8CA8E3B14CFB081E6EFA0E" box="[516,1053,1239,1432]" gridrow="2" pageId="35" pageNumber="36">
<th id="768F4858FF8CA8E3B14CFB081814FA0E" box="[516,615,1239,1432]" gridcol="0" gridrow="2" pageId="35" pageNumber="36">MD4 (ac) MD5 (ac) MD6 (ac) MD7 (ac) MD8 (ac)</th>
<td id="768F4858FF8CA8E3B1CEFB0818E8FA0E" box="[646,667,1239,1432]" gridcol="1" gridrow="2" pageId="35" pageNumber="36">20 19 19 22 22</td>
<td id="768F4858FF8CA8E3B186FB081945FA0E" box="[718,822,1239,1432]" gridcol="2" gridrow="2" pageId="35" pageNumber="36">2947 3348 3857 4261 6080</td>
<td id="768F4858FF8CA8E3B03CFB0819DFFA0E" box="[884,940,1239,1432]" gridcol="3" gridrow="2" pageId="35" pageNumber="36">35 39 44 49 70</td>
<td id="768F4858FF8CA8E3B0AEFB081E6EFA0E" box="[998,1053,1239,1432]" gridcol="4" gridrow="2" pageId="35" pageNumber="36">3,66 4,00 4,22 4,65 5,65</td>
</tr>
<tr id="355E2124FF8CA8E3B14CFA031E6EF962" box="[516,1053,1500,1780]" gridrow="3" pageId="35" pageNumber="36">
<th id="768F4858FF8CA8E3B14CFA031814F962" box="[516,615,1500,1780]" gridcol="0" gridrow="3" pageId="35" pageNumber="36">LV2 (tu) LV5 (tu) LV6 (ac) LV7 (ac) LV8 (ac) LV9 (ac) ML10 (tu)</th>
<td id="768F4858FF8CA8E3B1CEFA0318E8F962" box="[646,667,1500,1780]" gridcol="1" gridrow="3" pageId="35" pageNumber="36">21 23 24 25 23 22 23</td>
<td id="768F4858FF8CA8E3B186FA031945F962" box="[718,822,1500,1780]" gridcol="2" gridrow="3" pageId="35" pageNumber="36">2444 2434 2637 2842 3643 3854 2838</td>
<td id="768F4858FF8CA8E3B03CFA0319DFF962" box="[884,940,1500,1780]" gridcol="3" gridrow="3" pageId="35" pageNumber="36">31 30 31 37 41 44 32</td>
<td id="768F4858FF8CA8E3B0AEFA031E6EF962" box="[998,1053,1500,1780]" gridcol="4" gridrow="3" pageId="35" pageNumber="36">4,00 2,97 2,83 3,08 1,75 3,33 2,80</td>
</tr>
<tr id="355E2124FF8CA8E3B14CF8E81E6EF837" box="[516,1053,1847,1953]" gridrow="4" pageId="35" pageNumber="36">
<th id="768F4858FF8CA8E3B14CF8E81814F837" box="[516,615,1847,1953]" gridcol="0" gridrow="4" pageId="35" pageNumber="36">LTS LTS/TL LTAS</th>
<td id="768F4858FF8CA8E3B1CEF8E818E8F837" box="[646,667,1847,1953]" gridcol="1" gridrow="4" pageId="35" pageNumber="36">20 12 9</td>
<td id="768F4858FF8CA8E3B186F8E81945F837" box="[718,822,1847,1953]" gridcol="2" gridrow="4" pageId="35" pageNumber="36">163205 39.956.4% 4648</td>
<td id="768F4858FF8CA8E3B03CF8E819DFF837" box="[884,940,1847,1953]" gridcol="3" gridrow="4" pageId="35" pageNumber="36">186 48.0% 52</td>
<td id="768F4858FF8CA8E3B0AEF8E81E6EF837" box="[998,1053,1847,1953]" gridcol="4" gridrow="4" pageId="35" pageNumber="36">9,80 4.58% 3.91</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD12366FF8B5738B3F5FED6184BFED0" blockId="36.[189,1398,265,327]" pageId="36" pageNumber="37">The trichoscalid plates are well-developed, subdorsal and laterodorsal ones narrow and elongated, and ventromedial ones broadly oval.</paragraph>
<paragraph id="8BD12366FF8B5738B3F5FEB118A8FD7E" blockId="36.[188,1400,366,744]" pageId="36" pageNumber="37">
Segment 1 consists of a complete cuticular ring. Sensory spots are located on the anterior segment half in subdorsal, laterodorsal, and sublateral positions, and medially on segment, in ventromedial positions (
<figureCitation id="13553FE3FF8B5738B38DFE6B1B56FE58" box="[197,293,436,462]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Figs 9C</figureCitation>
,
<figureCitation id="13553FE3FF8B5738B27AFE6B1B3CFE58" box="[306,335,436,462]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">10</figureCitation>
BC); sensory spots on this all following segments are small, and rounded, with numerous micropapillae. Two longitudinally arranged glandular cell outlets
<typeStatus id="54D59DC4FF8B5738B0A4FE071E6CFE64" box="[1004,1055,472,498]" pageId="36" pageNumber="37">type</typeStatus>
1 are present in middorsal position (
<figureCitation id="13553FE3FF8B5738B262FE251BFAFD83" box="[298,393,506,533]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Figs 9C</figureCitation>
,
<figureCitation id="13553FE3FF8B5738B2DDFE241BB9FD83" box="[405,458,507,533]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">10B</figureCitation>
); anteriormost outlet appears narrower and more elongate than the posterior one; an additional pair of glandular cell outlets
<typeStatus id="54D59DC4FF8B5738B1E6FDC11892FDAE" box="[686,737,542,568]" pageId="36" pageNumber="37">type</typeStatus>
1 is present in ventrolateral positions (
<figureCitation id="13553FE3FF8B5738B7E0FDC11F60FDAE" box="[1192,1299,542,568]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">Fig. 10C</figureCitation>
). Dorsal and lateral sides, with scattered cuticular hairs emerging through rounded perforation sites; posterior half of ventral side with similar hair covering, but anterior part with large, W-shaped hairless area (
<figureCitation id="13553FE3FF8B5738B38EFD581B68FD37" box="[198,283,647,673]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">Fig. 10</figureCitation>
BC). The posterior segment margin is straight along the dorsal and lateral sides, and slightly convex ventrally, terminating into a pectinate fringe with short and very flexible fringe tips. Pachycyclus thin to medium thickness, and not interrupted.
</paragraph>
<paragraph id="8BD12366FF8B5738B3F5FCD019B6FB1F" blockId="36.[187,1400,783,1162]" pageId="36" pageNumber="37">
Segment 2 consists of a complete cuticular ring, but with indication of an intracuticular, partial, midventral fissure (
<figureCitation id="13553FE3FF8B5738B2D3FCED1B80FCDA" box="[411,499,818,844]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Figs 9F</figureCitation>
,
<figureCitation id="13553FE3FF8B5738B2B5FCED1841FCDA" box="[509,562,818,844]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">10C</figureCitation>
). Sensory spots are located in middorsal (but slightly laterally displaced), laterodorsal (twin pair) (
<figureCitation id="13553FE3FF8B5738B2ABFC8A1837FCE6" box="[483,580,853,880]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Figs 9E</figureCitation>
,
<figureCitation id="13553FE3FF8B5738B11CFC8918F6FCE6" box="[596,645,854,880]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">10B</figureCitation>
) and ventromedial positions (
<figureCitation id="13553FE3FF8B5738B0A7FC8A1E2EFCE6" box="[1007,1117,853,880]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">Fig. 10C</figureCitation>
). Minute glandular cell outlets
<typeStatus id="54D59DC4FF8B5738B25AFCA61B36FC05" box="[274,325,889,915]" pageId="36" pageNumber="37">type</typeStatus>
2 present in subdorsal positions (
<figureCitation id="13553FE3FF8B5738B184FCA6195FFC05" box="[716,812,889,915]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Figs 9C</figureCitation>
,
<figureCitation id="13553FE3FF8B5738B070FCA6191FFC05" box="[824,876,889,915]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">10B</figureCitation>
), and tubes present in ventrolateral positions (
<figureCitation id="13553FE3FF8B5738B38DFC431B55FC20" box="[197,294,924,950]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Figs 9D</figureCitation>
,
<figureCitation id="13553FE3FF8B5738B27AFC431B14FC20" box="[306,359,924,950]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">10C</figureCitation>
). Glandular cell outlets
<typeStatus id="54D59DC4FF8B5738B134FC4318DCFC20" box="[636,687,924,950]" pageId="36" pageNumber="37">type</typeStatus>
1 present in middorsal and ventromedial positions (
<figureCitation id="13553FE3FF8B5738B643FC431F21FC20" box="[1291,1362,924,950]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Figs 9</figureCitation>
C D, 10B). The segment is densely covered with bracteate hairs, but interrupted by hairless areas posterior to ventrolateral tubes, and in paraventral areas. Pachycyclus of the anterior segment margin is of medium thickness and not interrupted. The posterior segment margin is straight along dorsal and lateral sides, and extend into V-shaped flap on the ventral side; pectinate fringe from middorsal to midlateral positions with short fringe tips, as on segment 1; fringe tips from midlateral to ventromedial positions longer and very flexible, and then very short between ventromedial positions.
</paragraph>
<paragraph id="8BD12366FF8B5738B3F5FB6E1F49FA08" blockId="36.[187,1400,1200,1438]" pageId="36" pageNumber="37">
Segment 3, and remaining segments, consisting of one tergal and two sternal plates. Pachycyclus of the anterior segment margin of medium thickness, and interrupted only at tergosternal junctions. Sensory spots present in subdorsal, laterodorsal (
<figureCitation id="13553FE3FF8B5738B1D3FB281888FA87" box="[667,763,1271,1297]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Figs 9E</figureCitation>
,
<figureCitation id="13553FE3FF8B5738B043FB28194EFA87" box="[779,829,1271,1297]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">10D</figureCitation>
) and sublateral positions. Glandular cell outlets
<typeStatus id="54D59DC4FF8B5738B3F5FAC41A9CFAA3" box="[189,239,1307,1333]" pageId="36" pageNumber="37">type</typeStatus>
1 present in middorsal (
<figureCitation id="13553FE3FF8B5738B14EFAC518F2FAA3" box="[518,641,1306,1333]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Fig. 9E, H</figureCitation>
) and ventromedial positions. Bracteate cuticular hairs are densely covering the segment from middorsal to ventromedial positions (except in V-shaped areas in laterodorsal positions anterior on segment); paraventral areas that are devoid of any hair-like structures. Pectinate fringe of posterior margin as on preceding segment, but with ventromedial fringe tips being slightly longer.
</paragraph>
<paragraph id="8BD12366FF8B5738B3F5FA1A18B7F925" blockId="36.[187,1399,1477,1715]" pageId="36" pageNumber="37">
Segment 4 with acicular spine in middorsal position, reaching attachment point of middorsal spine on following segment (
<figureCitation id="13553FE3FF8B5738B298FA37184CF994" box="[464,575,1512,1538]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">Fig. 10D</figureCitation>
); this and all other spines on following five segments are slender and flexible. Sensory spots present in subdorsal (
<figureCitation id="13553FE3FF8B5738B1A8F9D41934F9B0" box="[736,839,1547,1574]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Figs 9H</figureCitation>
,
<figureCitation id="13553FE3FF8B5738B01FF9D319FFF9B0" box="[855,908,1548,1574]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">10D</figureCitation>
), midlateral and ventromedial positions; midlateral and ventromedial sensory spots considerably smaller than most other sensory spots in the species, but they occur consistently in all examined specimens. Glandular cell outlets
<typeStatus id="54D59DC4FF8B5738B7FAF98D1E96F9FA" box="[1202,1253,1618,1644]" pageId="36" pageNumber="37">type</typeStatus>
1 present in subdorsal (
<figureCitation id="13553FE3FF8B5738B274F9AA1BE5F919" box="[316,406,1653,1679]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Fig. 9H</figureCitation>
) and ventromedial positions. Pectinate fringe of posterior segment margin, pachycycli and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BD12366FF8B5738B3F5F9051E8EF817" blockId="36.[189,1399,1754,1921]" pageId="36" pageNumber="37">
Segment 5 with acicular spine in middorsal position, reaching attachment point of middorsal spine on following segment, and long, slender tubes in lateroventral positions (
<figureCitation id="13553FE3FF8B5738B70FF9221EC5F881" box="[1095,1206,1789,1815]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">Fig. 10E</figureCitation>
). Sensory spots present in subdorsal, midlateral (
<figureCitation id="13553FE3FF8B5738B11DF8FF18CEF8AC" box="[597,701,1824,1850]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="36" pageNumber="37">Figs 9H</figureCitation>
,
<figureCitation id="13553FE3FF8B5738B187F8FF1973F8AC" box="[719,768,1824,1850]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">10E</figureCitation>
) and ventromedial positions; ventromedial sensory spots considerably smaller than most other sensory spots in the species. Glandular cell outlets
<typeStatus id="54D59DC4FF8B5738B66DF89B1F2BF8C8" box="[1317,1368,1860,1886]" pageId="36" pageNumber="37">type</typeStatus>
1, pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BD12366FF8B5739B3F5F8771E5EFB0C" blockId="36.[189,1400,1960,2022]" lastBlockId="37.[189,1399,975,1178]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
Segment 6 with acicular spine in middorsal position, reaching attachment point of middorsal spine on following segment, and in lateroventral positions (
<figureCitation id="13553FE3FF8B5738B050F814191EF873" box="[792,877,1995,2021]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="36" pageNumber="37">Fig. 10</figureCitation>
EF), reaching beyond the posterior margin of the segment. Females with female papillae in ventrolateral positions; openings of papillae with fine fringes around their margins (
<figureCitation id="13553FE3FF8A5739B11FFC2C18CEFB9B" box="[599,701,1011,1037]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">Fig. 10F</figureCitation>
inset); intracuticular substructures each form an indistinct tubular structure (
<figureCitation id="13553FE3FF8A5739B2C6FBC91B9FFBA6" box="[398,492,1046,1072]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="37" pageNumber="38">Fig. 9G</figureCitation>
). Sensory spots present in paradorsal, midlateral and ventromedial (
<figureCitation id="13553FE3FF8A5739B640FBC91F1DFBA6" box="[1288,1390,1046,1072]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">Fig. 10F</figureCitation>
) positions. Pectinate fringe of posterior segment margin with uniform long fringe tips on tergal plate and on most lateral halves of sternal plates; fringe tips shorter between ventromedial positions. Glandular cell outlets
<typeStatus id="54D59DC4FF8A5739B20EFB5F1B0AFB0C" box="[326,377,1152,1178]" pageId="37" pageNumber="38">type</typeStatus>
1, pachycycli and cuticular hairs as on preceding segment.
</paragraph>
<caption id="DF1173EEFF8A5739B3F5FED6194BFED0" pageId="37" pageNumber="38" targetBox="[213,1359,385,895]" targetIsTable="true" targetPageId="37">
<paragraph id="8BD12366FF8A5739B3F5FED6194BFED0" blockId="37.[189,1398,265,327]" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B3F5FED61B42FEB5" bold="true" box="[189,305,265,291]" pageId="37" pageNumber="38">Table 10.</emphasis>
Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in
<taxonomicName id="4C6E58E5FF8A5739B2F0FEF31947FED0" authority="Lang, 1949" authorityName="Lang" authorityYear="1949" box="[440,820,300,327]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="37" pageNumber="38" phylum="Cephalorhyncha" rank="species" species="pilosus">
<emphasis id="B91AFF74FF8A5739B2F0FEF318D7FED0" box="[440,676,300,326]" italics="true" pageId="37" pageNumber="38">Echinoderes pilosus</emphasis>
<bibRefCitation id="EFFF5E97FF8A5739B1E3FEF21947FED0" author="Lang K." box="[683,820,300,327]" pageId="37" pageNumber="38" pagination="1 - 22" refId="ref56538" refString="Lang K. 1949. Echinoderida. In: Odhner N. H. (ed.) Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, Vol. 4: 1 - 22. P. A. Norstedt &amp; Soner, Stockholm." type="journal article" year="1949">Lang, 1949</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BD12366FF8A5739B39FFE5E1E32FCE8" pageId="37" pageNumber="38">
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<emphasis id="B91AFF74FF8A5739B39FFE5E1B50FE3B" bold="true" pageId="37" pageNumber="38">Position Segment</emphasis>
</th>
<th id="768F4858FF8AA8E3B207FE5E1BDFFE3B" box="[335,428,385,429]" gridcol="1" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B224FE511BE3FE37" bold="true" box="[364,400,398,417]" pageId="37" pageNumber="38">MD</emphasis>
</th>
<th id="768F4858FF8AA8E3B297FE5E1B88FE3B" box="[479,507,385,429]" gridcol="2" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B297FE511B88FE37" bold="true" box="[479,507,398,417]" pageId="37" pageNumber="38">PD</emphasis>
</th>
<th id="768F4858FF8AA8E3B161FE5E181DFE3B" box="[553,622,385,429]" gridcol="3" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B176FE51182AFE37" bold="true" box="[574,601,398,417]" pageId="37" pageNumber="38">SD</emphasis>
</th>
<th id="768F4858FF8AA8E3B1D2FE5E1893FE3B" box="[666,736,385,429]" gridcol="4" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B1E6FE5118BFFE37" bold="true" box="[686,716,398,417]" pageId="37" pageNumber="38">LD</emphasis>
</th>
<th id="768F4858FF8AA8E3B040FE5E1926FE3B" box="[776,853,385,429]" gridcol="5" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B054FE521930FE34" bold="true" box="[796,835,397,418]" pageId="37" pageNumber="38">ML</emphasis>
</th>
<th id="768F4858FF8AA8E3B0C8FE5E19E9FE3B" box="[896,922,385,429]" gridcol="6" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B0C8FE5119E9FE37" bold="true" box="[896,922,398,417]" pageId="37" pageNumber="38">SL</emphasis>
</th>
<th id="768F4858FF8AA8E3B089FE5E1E71FE3B" box="[961,1026,385,429]" gridcol="7" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B09CFE511981FE37" bold="true" box="[980,1010,398,417]" pageId="37" pageNumber="38">LA</emphasis>
</th>
<th id="768F4858FF8AA8E3B761FE5E1E36FE3B" box="[1065,1093,385,429]" gridcol="8" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B761FE511E36FE37" bold="true" box="[1065,1093,398,417]" pageId="37" pageNumber="38">LV</emphasis>
</th>
<th id="768F4858FF8AA8E3B724FE5E1ED8FE3B" box="[1132,1195,385,429]" gridcol="9" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B736FE511EEFFE37" bold="true" box="[1150,1180,398,417]" pageId="37" pageNumber="38">VL</emphasis>
</th>
<th id="768F4858FF8AA8E3B794FE5E1F3CFE3B" box="[1244,1359,385,429]" gridcol="10" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B91AFF74FF8A5739B64CFE511F5BFE37" bold="true" box="[1284,1320,398,417]" pageId="37" pageNumber="38">VM</emphasis>
</th>
</tr>
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<th id="768F4858FF8AA8E3B39DFE171B50FE48" box="[213,291,456,478]" gridcol="0" gridrow="1" pageId="37" pageNumber="38">1</th>
<td id="768F4858FF8AA8E3B207FE171BDFFE48" box="[335,428,456,478]" gridcol="1" gridrow="1" pageId="37" pageNumber="38">gco1, gco1</td>
<td id="768F4858FF8AA8E3B161FE17181DFE48" box="[553,622,456,478]" gridcol="3" gridrow="1" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B1D2FE171893FE48" box="[666,736,456,478]" gridcol="4" gridrow="1" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B0C8FE1719E9FE48" box="[896,922,456,478]" gridcol="6" gridrow="1" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B724FE171ED8FE48" box="[1132,1195,456,478]" gridcol="9" gridrow="1" pageId="37" pageNumber="38">gco1</td>
<td id="768F4858FF8AA8E3B794FE171F3CFE48" box="[1244,1359,456,478]" gridcol="10" gridrow="1" pageId="37" pageNumber="38">ss</td>
</tr>
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<th id="768F4858FF8AA8E3B39DFE2E1B50FD91" box="[213,291,497,519]" gridcol="0" gridrow="2" pageId="37" pageNumber="38">2</th>
<td id="768F4858FF8AA8E3B207FE2E1BDFFD91" box="[335,428,497,519]" gridcol="1" gridrow="2" pageId="37" pageNumber="38">gco1, ss</td>
<td id="768F4858FF8AA8E3B161FE2E181DFD91" box="[553,622,497,519]" gridcol="3" gridrow="2" pageId="37" pageNumber="38">gco2</td>
<td id="768F4858FF8AA8E3B1D2FE2E1893FD91" box="[666,736,497,519]" gridcol="4" gridrow="2" pageId="37" pageNumber="38">ss, ss</td>
<td id="768F4858FF8AA8E3B724FE2E1ED8FD91" box="[1132,1195,497,519]" gridcol="9" gridrow="2" pageId="37" pageNumber="38">tu</td>
<td id="768F4858FF8AA8E3B794FE2E1F3CFD91" box="[1244,1359,497,519]" gridcol="10" gridrow="2" pageId="37" pageNumber="38">gco1, ss</td>
</tr>
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<th id="768F4858FF8AA8E3B39DFDC41B50FDA7" box="[213,291,539,561]" gridcol="0" gridrow="3" pageId="37" pageNumber="38">3</th>
<td id="768F4858FF8AA8E3B207FDC41BDFFDA7" box="[335,428,539,561]" gridcol="1" gridrow="3" pageId="37" pageNumber="38">gco1</td>
<td id="768F4858FF8AA8E3B161FDC4181DFDA7" box="[553,622,539,561]" gridcol="3" gridrow="3" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B1D2FDC41893FDA7" box="[666,736,539,561]" gridcol="4" gridrow="3" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B0C8FDC419E9FDA7" box="[896,922,539,561]" gridcol="6" gridrow="3" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B794FDC41F3CFDA7" box="[1244,1359,539,561]" gridcol="10" gridrow="3" pageId="37" pageNumber="38">gco1</td>
</tr>
<tr id="355E2124FF8AA8E3B39DFD981F3CFDCD" box="[213,1359,583,603]" gridrow="4" pageId="37" pageNumber="38" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="768F4858FF8AA8E3B39DFD981B50FDCD" box="[213,291,583,603]" gridcol="0" gridrow="4" pageId="37" pageNumber="38">4</th>
<td id="768F4858FF8AA8E3B207FD981BDFFDCD" box="[335,428,583,603]" gridcol="1" gridrow="4" pageId="37" pageNumber="38">ac</td>
<td id="768F4858FF8AA8E3B161FD98181DFDCD" box="[553,622,583,603]" gridcol="3" gridrow="4" pageId="37" pageNumber="38">gco1, ss</td>
<td id="768F4858FF8AA8E3B040FD981926FDCD" box="[776,853,583,603]" gridcol="5" gridrow="4" pageId="37" pageNumber="38">ss*</td>
<td id="768F4858FF8AA8E3B794FD981F3CFDCD" box="[1244,1359,583,603]" gridcol="10" gridrow="4" pageId="37" pageNumber="38">ss*, gco1</td>
</tr>
<tr id="355E2124FF8AA8E3B39DFDAE1F3CFD12" box="[213,1359,625,644]" gridrow="5" pageId="37" pageNumber="38" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="768F4858FF8AA8E3B39DFDAE1B50FD12" box="[213,291,625,644]" gridcol="0" gridrow="5" pageId="37" pageNumber="38">5</th>
<td id="768F4858FF8AA8E3B207FDAE1BDFFD12" box="[335,428,625,644]" gridcol="1" gridrow="5" pageId="37" pageNumber="38">ac</td>
<td id="768F4858FF8AA8E3B161FDAE181DFD12" box="[553,622,625,644]" gridcol="3" gridrow="5" pageId="37" pageNumber="38">gco1, ss</td>
<td id="768F4858FF8AA8E3B040FDAE1926FD12" box="[776,853,625,644]" gridcol="5" gridrow="5" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B761FDAE1E36FD12" box="[1065,1093,625,644]" gridcol="8" gridrow="5" pageId="37" pageNumber="38">tu</td>
<td id="768F4858FF8AA8E3B794FDAE1F3CFD12" box="[1244,1359,625,644]" gridcol="10" gridrow="5" pageId="37" pageNumber="38">ss*, gco1</td>
</tr>
<tr id="355E2124FF8AA8E3B39DFD471F3CFD38" box="[213,1359,664,686]" gridrow="6" pageId="37" pageNumber="38" rowspan-4="1" rowspan-6="1" rowspan-7="1">
<th id="768F4858FF8AA8E3B39DFD471B50FD38" box="[213,291,664,686]" gridcol="0" gridrow="6" pageId="37" pageNumber="38">6</th>
<td id="768F4858FF8AA8E3B207FD471BDFFD38" box="[335,428,664,686]" gridcol="1" gridrow="6" pageId="37" pageNumber="38">ac</td>
<td id="768F4858FF8AA8E3B297FD471B88FD38" box="[479,507,664,686]" gridcol="2" gridrow="6" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B161FD47181DFD38" box="[553,622,664,686]" gridcol="3" gridrow="6" pageId="37" pageNumber="38">gco1</td>
<td id="768F4858FF8AA8E3B040FD471926FD38" box="[776,853,664,686]" gridcol="5" gridrow="6" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B761FD471E36FD38" box="[1065,1093,664,686]" gridcol="8" gridrow="6" pageId="37" pageNumber="38">ac</td>
<td id="768F4858FF8AA8E3B724FD471ED8FD38" box="[1132,1195,664,686]" gridcol="9" gridrow="6" pageId="37" pageNumber="38">fpa (♀)</td>
<td id="768F4858FF8AA8E3B794FD471F3CFD38" box="[1244,1359,664,686]" gridcol="10" gridrow="6" pageId="37" pageNumber="38">ss, gco1</td>
</tr>
<tr id="355E2124FF8AA8E3B39DFD1D1F3CFD4E" box="[213,1359,706,728]" gridrow="7" pageId="37" pageNumber="38" rowspan-4="1" rowspan-6="1" rowspan-7="1">
<th id="768F4858FF8AA8E3B39DFD1D1B50FD4E" box="[213,291,706,728]" gridcol="0" gridrow="7" pageId="37" pageNumber="38">7</th>
<td id="768F4858FF8AA8E3B207FD1D1BDFFD4E" box="[335,428,706,728]" gridcol="1" gridrow="7" pageId="37" pageNumber="38">ac</td>
<td id="768F4858FF8AA8E3B297FD1D1B88FD4E" box="[479,507,706,728]" gridcol="2" gridrow="7" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B161FD1D181DFD4E" box="[553,622,706,728]" gridcol="3" gridrow="7" pageId="37" pageNumber="38">gco1</td>
<td id="768F4858FF8AA8E3B040FD1D1926FD4E" box="[776,853,706,728]" gridcol="5" gridrow="7" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B761FD1D1E36FD4E" box="[1065,1093,706,728]" gridcol="8" gridrow="7" pageId="37" pageNumber="38">ac</td>
<td id="768F4858FF8AA8E3B724FD1D1ED8FD4E" box="[1132,1195,706,728]" gridcol="9" gridrow="7" pageId="37" pageNumber="38">fpa (♀)</td>
<td id="768F4858FF8AA8E3B794FD1D1F3CFD4E" box="[1244,1359,706,728]" gridcol="10" gridrow="7" pageId="37" pageNumber="38">ss, gco1</td>
</tr>
<tr id="355E2124FF8AA8E3B39DFD341F3CFC94" box="[213,1359,747,770]" gridrow="8" pageId="37" pageNumber="38" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="768F4858FF8AA8E3B39DFD341B50FC94" box="[213,291,747,770]" gridcol="0" gridrow="8" pageId="37" pageNumber="38">8</th>
<td id="768F4858FF8AA8E3B207FD341BDFFC94" box="[335,428,747,770]" gridcol="1" gridrow="8" pageId="37" pageNumber="38">ac</td>
<td id="768F4858FF8AA8E3B297FD341B88FC94" box="[479,507,747,770]" gridcol="2" gridrow="8" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B161FD34181DFC94" box="[553,622,747,770]" gridcol="3" gridrow="8" pageId="37" pageNumber="38">gco1</td>
<td id="768F4858FF8AA8E3B1D2FD341893FC94" box="[666,736,747,770]" gridcol="4" gridrow="8" pageId="37" pageNumber="38">gco2, ss</td>
<td id="768F4858FF8AA8E3B761FD341E36FC94" box="[1065,1093,747,770]" gridcol="8" gridrow="8" pageId="37" pageNumber="38">ac</td>
<td id="768F4858FF8AA8E3B724FD341ED8FC94" box="[1132,1195,747,770]" gridcol="9" gridrow="8" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B794FD341F3CFC94" box="[1244,1359,747,770]" gridcol="10" gridrow="8" pageId="37" pageNumber="38">gco1, fpa (♀)</td>
</tr>
<tr id="355E2124FF8AA8E3B39DFCCA1F3CFCBD" box="[213,1359,789,811]" gridrow="9" pageId="37" pageNumber="38" rowspan-1="1" rowspan-5="1" rowspan-6="1">
<th id="768F4858FF8AA8E3B39DFCCA1B50FCBD" box="[213,291,789,811]" gridcol="0" gridrow="9" pageId="37" pageNumber="38">9</th>
<td id="768F4858FF8AA8E3B297FCCA1B88FCBD" box="[479,507,789,811]" gridcol="2" gridrow="9" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B161FCCA181DFCBD" box="[553,622,789,811]" gridcol="3" gridrow="9" pageId="37" pageNumber="38">gco1, ss</td>
<td id="768F4858FF8AA8E3B1D2FCCA1893FCBD" box="[666,736,789,811]" gridcol="4" gridrow="9" pageId="37" pageNumber="38">gco2, ss</td>
<td id="768F4858FF8AA8E3B089FCCA1E71FCBD" box="[961,1026,789,811]" gridcol="7" gridrow="9" pageId="37" pageNumber="38">si</td>
<td id="768F4858FF8AA8E3B761FCCA1E36FCBD" box="[1065,1093,789,811]" gridcol="8" gridrow="9" pageId="37" pageNumber="38">ac</td>
<td id="768F4858FF8AA8E3B724FCCA1ED8FCBD" box="[1132,1195,789,811]" gridcol="9" gridrow="9" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B794FCCA1F3CFCBD" box="[1244,1359,789,811]" gridcol="10" gridrow="9" pageId="37" pageNumber="38">gco1</td>
</tr>
<tr id="355E2124FF8AA8E3B39DFCE01F3CFCC3" box="[213,1359,831,853]" gridrow="10" pageId="37" pageNumber="38" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="768F4858FF8AA8E3B39DFCE01B50FCC3" box="[213,291,831,853]" gridcol="0" gridrow="10" pageId="37" pageNumber="38">10</th>
<td id="768F4858FF8AA8E3B207FCE01BDFFCC3" box="[335,428,831,853]" gridcol="1" gridrow="10" pageId="37" pageNumber="38">gco1, gco1</td>
<td id="768F4858FF8AA8E3B297FCE01B88FCC3" box="[479,507,831,853]" gridcol="2" gridrow="10" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B1D2FCE01893FCC3" box="[666,736,831,853]" gridcol="4" gridrow="10" pageId="37" pageNumber="38">tu</td>
<td id="768F4858FF8AA8E3B724FCE01ED8FCC3" box="[1132,1195,831,853]" gridcol="9" gridrow="10" pageId="37" pageNumber="38">ss (Ƌ)</td>
<td id="768F4858FF8AA8E3B794FCE01F3CFCC3" box="[1244,1359,831,853]" gridcol="10" gridrow="10" pageId="37" pageNumber="38">gco1</td>
</tr>
<tr id="355E2124FF8AA8E3B39DFCB61F3CFCE9" box="[213,1359,873,895]" gridrow="11" pageId="37" pageNumber="38" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-9="1">
<th id="768F4858FF8AA8E3B39DFCB61B50FCE9" box="[213,291,873,895]" gridcol="0" gridrow="11" pageId="37" pageNumber="38">11</th>
<td id="768F4858FF8AA8E3B207FCB61BDFFCE9" box="[335,428,873,895]" gridcol="1" gridrow="11" pageId="37" pageNumber="38">gco1, gco1</td>
<td id="768F4858FF8AA8E3B161FCB6181DFCE9" box="[553,622,873,895]" gridcol="3" gridrow="11" pageId="37" pageNumber="38">ss</td>
<td id="768F4858FF8AA8E3B040FCB61926FCE9" box="[776,853,873,895]" gridcol="5" gridrow="11" pageId="37" pageNumber="38">3xpe (Ƌ)</td>
<td id="768F4858FF8AA8E3B089FCB61E71FCE9" box="[961,1026,873,895]" gridcol="7" gridrow="11" pageId="37" pageNumber="38">ltas (♀)</td>
<td id="768F4858FF8AA8E3B761FCB61E36FCE9" box="[1065,1093,873,895]" gridcol="8" gridrow="11" pageId="37" pageNumber="38">lts</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD12366FF8A5739B3F5FB1E19E3FAD3" blockId="37.[189,1400,1217,1349]" pageId="37" pageNumber="38">
Segment 7 with acicular spine in middorsal position, extending about halfway over the following segment, and in lateroventral positions, reaching beyond the posterior margin of the segment. Female papillae (
<figureCitation id="13553FE3FF8A5739B263FAD81BDBFAB4" box="[299,424,1287,1314]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="37" pageNumber="38">Figs 9G, J</figureCitation>
,
<figureCitation id="13553FE3FF8A5739B2FCFAD81B89FAB4" box="[436,506,1287,1314]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">10F, I</figureCitation>
), sensory spots, glandular cell outlets
<typeStatus id="54D59DC4FF8A5739B0F3FAD7199DFAB4" box="[955,1006,1288,1314]" pageId="37" pageNumber="38">type</typeStatus>
1, pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BD12366FF8A5739B3F5FAB318E6F936" blockId="37.[188,1400,1388,1696]" pageId="37" pageNumber="38">
Segment 8 with acicular spine in middorsal position, extending more than halfway over the following segment (
<figureCitation id="13553FE3FF8A5739B267FA501BEFFA3F" box="[303,412,1423,1449]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">Fig. 10G</figureCitation>
), and in lateroventral positions (
<figureCitation id="13553FE3FF8A5739B05AFA501916FA3F" box="[786,869,1423,1449]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="37" pageNumber="38">Fig. 9J</figureCitation>
), reaching beyond the posterior margin of the segment (
<figureCitation id="13553FE3FF8A5739B265FA6D1BFDFA5A" box="[301,398,1458,1484]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">Fig. 10I</figureCitation>
). Minute glandular cell outlets
<typeStatus id="54D59DC4FF8A5739B1B0FA6C1958FA5B" box="[760,811,1459,1485]" pageId="37" pageNumber="38">type</typeStatus>
2 present in laterodorsal positions (
<figureCitation id="13553FE3FF8A5739B781FA6D1F55FA5A" box="[1225,1318,1458,1484]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="37" pageNumber="38">Figs 9L</figureCitation>
,
<figureCitation id="13553FE3FF8A5739B67AFA6D1F1AFA5A" box="[1330,1385,1458,1484]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">10H</figureCitation>
). Sensory spots present in paradorsal, laterodorsal (posterior to glandular cell outlets) and ventrolateral positions (
<figureCitation id="13553FE3FF8A5739B271FA261BF9F985" box="[313,394,1529,1555]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">Fig. 10</figureCitation>
GI). Females with female papillae in ventromedial positions (
<figureCitation id="13553FE3FF8A5739B721FA261EB3F985" box="[1129,1216,1529,1555]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="37" pageNumber="38">Figs 9J</figureCitation>
,
<figureCitation id="13553FE3FF8A5739B784FA261E84F985" box="[1228,1271,1529,1555]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">10I</figureCitation>
); openings of papillae with fine fringes around their margins (
<figureCitation id="13553FE3FF8A5739B040F9C31916F9A0" box="[776,869,1564,1590]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">Fig. 10I</figureCitation>
inset); intracuticular substructures each form an indistinct tubular structure. Glandular cell outlets
<typeStatus id="54D59DC4FF8A5739B06BF99F1925F9CC" box="[803,854,1600,1626]" pageId="37" pageNumber="38">type</typeStatus>
1 as on preceding segment, but subdorsal ones are situated slightly closer to each other (
<figureCitation id="13553FE3FF8A5739B1EDF9BD188BF9EA" box="[677,760,1634,1660]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="37" pageNumber="38">Fig. 9I</figureCitation>
). Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BD12366FF8A5739B3F5F9181855F823" blockId="37.[189,1400,1735,1973]" pageId="37" pageNumber="38">
Segment 9 with acicular spines in lateroventral positions, extending about halfway over the following segment. Minute glandular cell outlets
<typeStatus id="54D59DC4FF8A5739B1C2F93418CEF893" box="[650,701,1771,1797]" pageId="37" pageNumber="38">type</typeStatus>
2 present in laterodorsal positions (
<figureCitation id="13553FE3FF8A5739B72AF9351EB2F892" box="[1122,1217,1770,1796]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="37" pageNumber="38">Figs 9L</figureCitation>
,
<figureCitation id="13553FE3FF8A5739B786F9351F77F892" box="[1230,1284,1770,1796]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">10H</figureCitation>
). Sensory spots present in paradorsal, subdorsal, laterodorsal (
<figureCitation id="13553FE3FF8A5739B062F8D2190CF8BE" box="[810,895,1805,1832]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="37" pageNumber="38">Fig. 10</figureCitation>
GH), and ventrolateral positions. Female papillae absent. Glandular cell outlets
<typeStatus id="54D59DC4FF8A5739B1C9F8EE18C7F8DD" box="[641,692,1841,1867]" pageId="37" pageNumber="38">type</typeStatus>
1 as on preceding segment, but with subdorsal ones situated even closer to each other, very close to paradorsal positions. Small, rounded nephridial sieve plates present in lateral accessory positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BD12366FF89573AB3F5FED61B95FD8C" blockId="38.[187,1399,265,538]" pageId="38" pageNumber="39">
Segment 10 with long, slender laterodorsal tubes near posterior segment margin (
<figureCitation id="13553FE3FF89573AB7D6FED61F7AFEB5" box="[1182,1289,265,291]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="38" pageNumber="39">Figs 9M</figureCitation>
,
<figureCitation id="13553FE3FF89573AB650FED61F1AFEB5" box="[1304,1385,265,291]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="38" pageNumber="39">10J, L</figureCitation>
). Sensory spots present in paradorsal positions (
<figureCitation id="13553FE3FF89573AB1AAFEF31935FED0" box="[738,838,300,326]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="38" pageNumber="39">Fig. 10J</figureCitation>
); males with additional pair of sensory spots in ventrolateral positions (
<figureCitation id="13553FE3FF89573AB29DFE901830FEFF" box="[469,579,335,361]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="38" pageNumber="39">Fig. 10K</figureCitation>
). Glandular cell outlets
<typeStatus id="54D59DC4FF89573AB017FE8F19E1FEFC" box="[863,914,336,362]" pageId="38" pageNumber="39">type</typeStatus>
1 present as two longitudinally arranged middorsal ones and in ventromedial positions. The posterior segment margin of the tergal plate is straight, whereas margins of sternal plates are concave and extend into a midventral point (
<figureCitation id="13553FE3FF89573AB733FE491E9BFE26" box="[1147,1256,406,432]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="38" pageNumber="39">Fig. 10K</figureCitation>
); fringe tips of pectinate fringe are as on preceding segment on tergal plate, but considerably shorter on sternal plates. Cuticular hairs as on preceding segments on tergal plates, but more sparse on sternal plates. Pachycycli as on preceding segment.
</paragraph>
<paragraph id="8BD12366FF89573AB3F5FD9E19DBFCC7" blockId="38.[187,1399,577,849]" pageId="38" pageNumber="39">
Segment 11 with lateral terminal spines (
<figureCitation id="13553FE3FF89573AB1D9FD9E1947FDCD" box="[657,820,577,603]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="38" pageNumber="39">Fig. 9B, K, M</figureCitation>
). Males with three pairs of penile spines (
<figureCitation id="13553FE3FF89573AB645FD9E1F01FDCD" box="[1293,1394,577,603]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="38" pageNumber="39">Figs 9M</figureCitation>
,
<figureCitation id="13553FE3FF89573AB3F3FDBB1A82FDE8" box="[187,241,612,638]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="38" pageNumber="39">10K</figureCitation>
); dorsal and ventral penile spines are stout but still flexible tubes, whereas the median ones are slightly thicker and more rigid; females with short, and relatively stout lateral terminal accessory spines (
<figureCitation id="13553FE3FF89573AB38DFD751B55FD53" box="[197,294,682,709]" captionStart="Fig" captionStartId="33.[189,232,1742,1768]" captionTargetBox="[263,1324,268,1704]" captionTargetId="figure-11@33.[260,1328,265,1706]" captionTargetPageId="33" captionText="Fig. 9. Light micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AB, GL. Lectotype, ♀ (SMNH-3930a). CF, M. Paralectotype, ♂ (SMNH-3930b). A. Dorsal overview. B. Ventral overview. C. Segments 1 to 2, dorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, dorsal view. F. Segments 2 to 4, ventral view. G. Segments 6 to 7, ventral view, showing female sexual dimorphism. H. Segments 3 to 5, dorsal view. I. Segments 8 to 9, dorsal view. J. Segments 7 to 8, ventral view, showing female sexual dimorphism. K. Segments 10 to 11, ventrolateral view, showing female sexual dimorphism.L. Segments 8 to 9, dorsal view but deeper focal plane than (I). M. Segments 10 to 11, ventral view, showing male sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4418999" httpUri="https://zenodo.org/record/4418999/files/figure.png" pageId="38" pageNumber="39">Figs 9K</figureCitation>
,
<figureCitation id="13553FE3FF89573AB279FD741B0DFD53" box="[305,382,683,709]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="38" pageNumber="39">10J, L</figureCitation>
). Sensory spots present in subdorsal positions medially on segment. Two longitudinally arranged glandular cell outlets
<typeStatus id="54D59DC4FF89573AB17AFD111816FD7E" box="[562,613,718,744]" pageId="38" pageNumber="39">type</typeStatus>
1 present in middorsal position, anterior on segment. The dorsal side of the segment is sparsely covered with thin, non-bracteate hair-like extensions; ventral side is almost hair-less. Tergal extensions are well-spaced, short and pointed; sternal extensions short, with ventrolateral seta-like tuft of extended fringe tips (
<figureCitation id="13553FE3FF89573AB044FCE81913FCC7" box="[780,864,823,849]" captionStart="Fig" captionStartId="34.[189,232,1705,1731]" captionTargetBox="[287,1300,270,1661]" captionTargetId="figure-13@34.[284,1304,265,1665]" captionTargetPageId="34" captionText="Fig. 10. Scanning electron micrographs showing overviews and details of Echinoderes pilosus Lang, 1949. AJ, L. ♀ (SMNH-9262, SMNH-9281). K.♂ (SMNH-9262). A. Lateral overview. B. Segments 1 to 2, dorsal view. C. Segments 1 to 2, ventral view. D. Segments 3 to 4, dorsal view. E. Segments 5 to 6, lateral view. F. Segments 6 to 7, ventral view, showing female sexual dimorphism; inset shows ventrolateral female papilla on segment 6, scale on inset = 1 µm. G. Segments 8 to 9, dorsal view. H. Segments 8 to 9, laterodorsal view. I. Segments 7 to 8, ventral view, showing female sexual dimorphism; inset shows ventromedial female papilla on segment 8, scale on inset = 2 µm. J. Segments 10 to 11 dorsal view, showing female sexual dimorphism. K. Segments 10 to 11 ventral view, showing male sexual dimorphism. L. Segments 10 to 11 ventral view, showing female sexual dimorphism." figureDoi="http://doi.org/10.5281/zenodo.4419001" httpUri="https://zenodo.org/record/4419001/files/figure.png" pageId="38" pageNumber="39">Fig. 10</figureCitation>
KL).
</paragraph>
</subSubSection>
</treatment>
</document>