304 lines
34 KiB
XML
304 lines
34 KiB
XML
<document id="9AA7B1824B7F75D69E5B4B96C33543A7" ID-DOI="10.1080/00222930400008736" ID-ISSN="1464-5262" ID-Zenodo-Dep="5214684" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629257395829" checkinUser="felipe" docAuthor="Chen, I-Shiung & Kottelat, Maurice" docDate="2005" docId="03C7915F37116D0F82C60345FD1E7576" docLanguage="en" docName="JNatHist.39.17.1407-1429.pdf" docOrigin="Journal of Natural History 39 (17)" docSource="http://dx.doi.org/10.1080/00222930400008736" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Rhinogbius variolatus Chen & Kottelat 2005, new species" docType="treatment" docVersion="3" lastPageNumber="1422" masterDocId="FFFEE927371D6D1E837A0657FFFB7740" masterDocTitle="Four new freshwater gobies of the genus Rhinogobius (Teleostei: Gobiidae) from northern Vietnam" masterLastPageNumber="1429" masterPageNumber="1407" pageNumber="1417" updateTime="1699065453420" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods id="C4F5B59ED5235FE10AB1995FFC506A14" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="444336A7EB493DF405ABCBCA4B9E3F10">
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<mods:title id="3BCD89C6FEF0D7000229B0A9620B0B0F">Four new freshwater gobies of the genus Rhinogobius (Teleostei: Gobiidae) from northern Vietnam</mods:title>
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</mods:titleInfo>
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<mods:name id="89AAEA9CA29418B7FC83B2A428478E42" type="personal">
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<mods:role id="2C38B6AD94CFCEEF360C8403F6001C40">
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<mods:roleTerm id="CDFE21738F3758F164A65BF884D6C9A0">Author</mods:roleTerm>
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<mods:namePart id="2FF1B954B9AB23CF44F520C926B091BF">Chen, I-Shiung</mods:namePart>
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</mods:name>
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<mods:name id="617C1C0BCB55C10DE93A037F447C2F11" type="personal">
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<mods:roleTerm id="E51C080BA20A5A1298410E879D069C9F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9443ABA3188168E2573BA66E5831C832">Kottelat, Maurice</mods:namePart>
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<mods:typeOfResource id="1B6B37009675506B08835AFED580A913">text</mods:typeOfResource>
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<mods:title id="2430BA224EEDF6DED244A3ABDB01F959">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="1B91EC4D0F6C14B65664D354718C837B">
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<mods:date id="3074222E350AE2725CFE5233655BBE92">2005</mods:date>
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<mods:detail id="46BED163F617341E35107D58234E109E" type="pubDate">
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<mods:number id="191ED61F84AB794A8E0C2A3D91DBC6C2">2005-09-30</mods:number>
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</mods:detail>
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<mods:detail id="555D3728DD1245B0CA14B8F417ADCE60" type="volume">
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<mods:number id="E8ECD69EBD49A413D57690DD89130353">39</mods:number>
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</mods:detail>
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<mods:detail id="5A818D813869F63322BD23F782C7F13A" type="issue">
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<mods:number id="1CB82E441A12ECE22E2237344DBCF210">17</mods:number>
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</mods:detail>
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<mods:extent id="7D1A70EDF739B38ED626D86F64F2E872" unit="page">
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<mods:start id="3204D97D5AF1C2D4EA941CEA1B83704A">1407</mods:start>
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<mods:end id="91393FA7F701EDE88E517D73AE6F44DF">1429</mods:end>
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</mods:extent>
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<mods:location id="2836BD1A216E26EE6B9B679042DAC769">
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<mods:url id="C6816DDF3148268D45AF6559EEAE05A6">http://dx.doi.org/10.1080/00222930400008736</mods:url>
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<mods:classification id="09889C847E5B9B4D25F70FE0EEED5C1A">journal article</mods:classification>
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<mods:identifier id="20D6AA25E09AC59E3E24DA469C21D3FB" type="DOI">10.1080/00222930400008736</mods:identifier>
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<mods:identifier id="D0AB408D8D5C6D8D9BF4A5E682A1A8EA" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="B03A22ED19D3EF7CF674FA2B41AFF6EE" type="Zenodo-Dep">5214684</mods:identifier>
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<treatment id="03C7915F37116D0F82C60345FD1E7576" LSID="urn:lsid:plazi:treatment:03C7915F37116D0F82C60345FD1E7576" httpUri="http://treatment.plazi.org/id/03C7915F37116D0F82C60345FD1E7576" lastPageId="17" lastPageNumber="1422" pageId="12" pageNumber="1417">
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<subSubSection id="C37473C237116D1282C60345FC94726A" box="[444,879,1298,1322]" pageId="12" pageNumber="1417" type="nomenclature">
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<paragraph id="8BD1204937116D1282C60345FC94726A" blockId="12.[444,879,1298,1361]" box="[444,879,1298,1322]" pageId="12" pageNumber="1417">
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<heading id="D099972537116D1282C60345FC94726A" box="[444,879,1298,1322]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="1417" reason="2">
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<taxonomicName id="4C6E5BCA37116D1282C60345FD26726A" authority="Chen & Kottelat, 2005" authorityName="Chen & Kottelat" authorityYear="2005" box="[444,733,1298,1322]" class="Actinopterygii" family="Gobiidae" genus="Rhinogbius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="1417" phylum="Chordata" rank="species" species="variolatus" status="sp. nov.">
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<emphasis id="B91AFC5B37116D1282C60345FD26726A" bold="true" box="[444,733,1298,1322]" italics="true" pageId="12" pageNumber="1417">Rhinogbius variolatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A229412037116D12819C0345FC94726A" box="[742,879,1298,1322]" pageId="12" pageNumber="1417" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37473C237116D0F814A036EFF277603" lastPageId="17" lastPageNumber="1422" pageId="12" pageNumber="1417" type="description">
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<paragraph id="8BD1204937116D12814A036EFD007211" blockId="12.[444,879,1298,1361]" box="[560,763,1337,1361]" pageId="12" pageNumber="1417">
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(
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<figureCitation id="13553CCC37116D128142036EFD5D7211" box="[568,678,1337,1361]" captionStart="Figure 4" captionStartId="11.[125,185,1659,1678]" captionTargetBox="[208,1091,192,1622]" captionTargetId="figure-0@11.[202,1095,187,1623]" captionTargetPageId="11" captionText="Figure 4. Head lateral-line system of Rhinogobius variolatus, paratype, CMK 14982, 25.9 mm SL. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5214692" httpUri="https://zenodo.org/record/5214692/files/figure.png" pageId="12" pageNumber="1417">Figures 4</figureCitation>
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,
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<figureCitation id="13553CCC37116D1281CE036EFD3F7211" box="[692,708,1337,1361]" captionStart="Figure 8" captionStartId="17.[125,185,1636,1655]" captionTargetBox="[176,1128,634,1598]" captionTargetId="figure-126@17.[170,1128,630,1600]" captionTargetPageId="17" captionText="Figure 8. Rhinogobius variolatus, holotype, ZRC 49202, 26.8 mm SL, cephalic colour pattern of male. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5214700" httpUri="https://zenodo.org/record/5214700/files/figure.png" pageId="12" pageNumber="1417">8</figureCitation>
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,
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<figureCitation id="13553CCC37116D1281A9036EFD097211" box="[723,754,1337,1361]" captionStart="Figure 12" captionStartId="20.[138,198,697,716]" captionTargetBox="[183,1140,133,661]" captionTargetId="figure-67@20.[183,1140,133,661]" captionTargetPageId="20" captionText="Figure 12. Rhinogobius variolatus. (A) Male, holotype, ZRC 49202, 26.8 mm SL; (B) female, paratype, ZRC 49203, 27.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.5214709" httpUri="https://zenodo.org/record/5214709/files/figure.png" pageId="12" pageNumber="1417">12</figureCitation>
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)
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</paragraph>
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<paragraph id="8BD1204937116D1283F003DDFEA372E2" blockId="12.[138,344,1418,1442]" box="[138,344,1418,1442]" pageId="12" pageNumber="1417">
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<emphasis id="B91AFC5B37116D1283F003DDFEA372E2" box="[138,344,1418,1442]" italics="true" pageId="12" pageNumber="1417">Material examined</emphasis>
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</paragraph>
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<paragraph id="8BD1204937116D1283F003ECFEB771D5" blockId="12.[138,1185,1467,1685]" pageId="12" pageNumber="1417">
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<materialsCitation id="3B062A1437116D1283F003ECFCE57153" accessionNumber="ZRC 49202" collectingDate="1998-10-05" collectorName="M. Kottelat" country="Vietnam" location="Cam Pha District" pageId="12" pageNumber="1417" specimenCount="1" stateProvince="Quang Ninh" typeStatus="holotype">
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<typeStatus id="54D59EEB37116D1283F003ECFF017293" box="[138,250,1467,1491]" pageId="12" pageNumber="1417" type="holotype">Holotype</typeStatus>
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: ZRC 49202,
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<quantity id="4C968DAC37116D1282DE03ECFDF17293" box="[420,522,1467,1491]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" pageId="12" pageNumber="1417" unit="mm" value="26.8">26.8 mm</quantity>
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SL,
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<collectingCountry id="F37960D937116D12813F03ECFD577293" box="[581,684,1467,1491]" name="Vietnam" pageId="12" pageNumber="1417">Vietnam</collectingCountry>
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:
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<collectingRegion id="49AAEEAB37116D1281C203ECFC467293" box="[696,957,1467,1491]" country="Vietnam" name="Quang Ninh" pageId="12" pageNumber="1417">Quang Ninh Province</collectingRegion>
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,
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<location id="8EB1769237116D1280B003ECFB667293" LSID="urn:lsid:plazi:treatment:03C7915F37116D0F82C60345FD1E7576:8EB1769237116D1280B003ECFB667293" box="[970,1181,1467,1491]" country="Vietnam" name="Cam Pha District" pageId="12" pageNumber="1417" stateProvince="Quang Ninh">Cam Pha District</location>
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, small coastal stream, at about km 10 on road from Mong Duong to Tien Yen, 21
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<emphasis id="B91AFC5B37116D128756038CFBCD72B3" bold="true" box="[1068,1078,1499,1523]" italics="true" pageId="12" pageNumber="1417">°</emphasis>
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059380N, 107
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<emphasis id="B91AFC5B37116D1283CC03ACFF3B7153" bold="true" box="[182,192,1531,1555]" italics="true" pageId="12" pageNumber="1417">°</emphasis>
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21920E, coll.
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<collectorName id="269B459F37116D12821F03ACFE0F7153" box="[357,500,1531,1555]" pageId="12" pageNumber="1417">M. Kottelat</collectorName>
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||
et al.,
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<date id="FFD0068937116D12812C03ACFCE57153" box="[598,798,1531,1555]" pageId="12" pageNumber="1417" value="1998-10-05">
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<collectingDate id="EF94FF6137116D12812C03ACFCE57153" box="[598,798,1531,1555]" pageId="12" pageNumber="1417" value="1998-10-05">5 October 1998</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation id="3B062A1437116D12805503ACFEFF7174" accessionNumber="CMK 14982" country="Vietnam" pageId="12" pageNumber="1417" specimenCount="1" stateProvince="Quang Ninh" typeStatus="paratype">
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<typeStatus id="54D59EEB37116D12805503ACFC587153" box="[815,931,1531,1555]" pageId="12" pageNumber="1417" type="paratype">Paratypes</typeStatus>
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: CMK 14982,
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<specimenCount id="9D68EBC037116D12871C03ACFEFF7174" pageId="12" pageNumber="1417" type="generic">three specimens</specimenCount>
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</materialsCitation>
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,
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<materialsCitation id="3B062A1437116D12826B004BFBB47174" accessionNumber="ZRC 49203" box="[273,1103,1564,1588]" collectingDate="1998-10-05" collectorName="M. Kottelat" country="Vietnam" location="Cam Pha District" pageId="12" pageNumber="1417" specimenCount="1" stateProvince="Quang Ninh" typeStatus="paratype">
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ZRC 49203,
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<specimenCount id="9D68EBC037116D1282D4004BFDA07174" box="[430,603,1564,1588]" pageId="12" pageNumber="1417" type="generic">two specimens</specimenCount>
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,
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<quantity id="4C968DAC37116D128112004BFCF47174" box="[616,783,1564,1588]" metricMagnitude="-2" metricUnit="m" metricValue="2.405" metricValueMax="2.76" metricValueMin="2.05" pageId="12" pageNumber="1417" unit="mm" value="24.05" valueMax="27.6" valueMin="20.5">20.5–27.6 mm</quantity>
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SL, same data as holotype
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</materialsCitation>
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.
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<materialsCitation id="3B062A1437116D128726004BFEB371D5" accessionNumber="CMK 14951" collectingDate="1998-10-04" collectorName="M. Kottelat" country="Vietnam" location="Ba Che River" municipality="Ba Che District" pageId="12" pageNumber="1417" specimenCount="1" stateProvince="Quang Ninh" typeStatus="paratype">
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CMK 14951,
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<specimenCount id="9D68EBC037116D12839E006BFE7C7114" box="[228,391,1596,1620]" pageId="12" pageNumber="1417" type="generic">one specimen</specimenCount>
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,
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<quantity id="4C968DAC37116D1282EE006BFE017114" box="[404,506,1596,1620]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="12" pageNumber="1417" unit="mm" value="20.2">20.2 mm</quantity>
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SL,
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<collectingCountry id="F37960D937116D12814F006BFD607114" box="[565,667,1596,1620]" name="Vietnam" pageId="12" pageNumber="1417">Vietnam</collectingCountry>
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,
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<collectingRegion id="49AAEEAB37116D1281D2006BFC577114" box="[680,940,1596,1620]" country="Vietnam" name="Quang Ninh" pageId="12" pageNumber="1417">Quang Ninh Province</collectingRegion>
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,
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<collectingMunicipality id="6BB5BA3337116D1280C3006BFB8E7114" box="[953,1141,1596,1620]" pageId="12" pageNumber="1417">Ba Che District</collectingMunicipality>
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,
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<location id="8EB1769237116D1287F8006BFF047135" LSID="urn:lsid:plazi:treatment:03C7915F37116D0F82C60345FD1E7576:8EB1769237116D1287F8006BFF047135" country="Vietnam" municipality="Ba Che District" name="Ba Che River" pageId="12" pageNumber="1417" stateProvince="Quang Ninh">Ba Che River</location>
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<quantity id="4C968DAC37116D12827C000AFEC57135" box="[262,318,1629,1653]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="1417" unit="km" value="4.0">4 km</quantity>
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up-river of Ba Che city, 21
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<emphasis id="B91AFC5B37116D128103000BFD787134" bold="true" box="[633,643,1628,1652]" italics="true" pageId="12" pageNumber="1417">°</emphasis>
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169340N, 107
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<emphasis id="B91AFC5B37116D12805B000BFCD07134" bold="true" box="[801,811,1628,1652]" italics="true" pageId="12" pageNumber="1417">°</emphasis>
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149540E, coll.
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<collectorName id="269B459F37116D1280AB000AFBA17135" box="[977,1114,1629,1653]" pageId="12" pageNumber="1417">M. Kottelat</collectorName>
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||
et al.,
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<date id="FFD0068937116D1283F0002AFEB371D5" box="[138,328,1661,1685]" pageId="12" pageNumber="1417" value="1998-10-04">
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<collectingDate id="EF94FF6137116D1283F0002AFEB371D5" box="[138,328,1661,1685]" pageId="12" pageNumber="1417" value="1998-10-04">4 October 1998</collectingDate>
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</date>
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="8BD1204937106D13830706DEFF1D77E1" blockId="13.[125,230,137,161]" box="[125,230,137,161]" pageId="13" pageNumber="1418">
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<emphasis id="B91AFC5B37106D13830706DEFF1D77E1" box="[125,230,137,161]" italics="true" pageId="13" pageNumber="1418">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BD1204937106D13830706EEFE8076F1" blockId="13.[125,1172,185,433]" pageId="13" pageNumber="1418">
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<taxonomicName id="4C6E5BCA37106D13830706EEFE927791" box="[125,361,185,209]" class="Actinopterygii" family="Gobiidae" genus="Rhingobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="1418" phylum="Chordata" rank="species" species="variolatus">
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<emphasis id="B91AFC5B37106D13830706EEFE927791" box="[125,361,185,209]" italics="true" pageId="13" pageNumber="1418">Rhingobius variolatus</emphasis>
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</taxonomicName>
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is distinguished from all congeners by the unique combination of the following features: cheek and opercle with two rows of longitudinally arranged black spots in male, none in female; branchiostegal membrane without distinct mark in male; body with two major longitudinal rows of brownish black spots on lower half of lateral trunk in male; pectoral fin whitish, base with two conspicuous black spots in male; caudal fin base with a median black spot; second dorsal fin rays modally eight; anal fin rays modally seven; pectoral fin rays 15; longitudinal scale rows 26–27; predorsal median series 8–11; and vertebrae modally 27.
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</paragraph>
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<paragraph id="8BD1204937106D13830707BDFF0E7542" blockId="13.[125,245,490,514]" box="[125,245,490,514]" pageId="13" pageNumber="1418">
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<emphasis id="B91AFC5B37106D13830707BDFF0E7542" box="[125,245,490,514]" italics="true" pageId="13" pageNumber="1418">Description</emphasis>
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</paragraph>
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<paragraph id="8BD1204937106D138307044CFD9975B2" blockId="13.[125,1173,539,754]" pageId="13" pageNumber="1418">Body cylindrical anteriorly and compressed posteriorly. Body proportions in Table I. Head moderately large, slightly depressed anteriorly. Eye large, lips thick. Mouth oblique, rear edge reaching beyond vertical of anterior margin of pupil in male, not reaching vertical of anterior margin in female. Both jaws with three to four rows of conical teeth, and outer rows enlarged. Tongue margin rounded. Anterior nostril a short tube and posterior one a round hole. Gill opening extending to vertical through middle of opercle. Isthmus broad. 10+16217526227 vertebrae (mode 27).</paragraph>
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<paragraph id="8BD1204937106D138307054CFF1C7493" blockId="13.[125,1173,793,979]" pageId="13" pageNumber="1418">
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<emphasis id="B91AFC5B37106D138307054CFF4E7473" box="[125,181,795,819]" italics="true" pageId="13" pageNumber="1418">Fins.</emphasis>
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D1 VI; D2 I/7–8 (mode 8); A I/7–8 (mode 7); P 15–16 (mode 15); V I/5+I/5 (frequency distribution in Table II). D1 rays about equal, III, IV longest, rear tip not extending to origin of D
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||
<quantity id="4C968DAC37106D1382EA050CFE407433" box="[400,443,859,883]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="13" pageNumber="1418" unit="in" value="2.0">2 in</quantity>
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both sexes. A origin inserted below second branched ray of D2. P large, rear margin almost extending to vertical of anus in male; never reaching it in female. V disc rounded, spinous rays with pointed membranous lobe. C elliptical, rear edge rounded.
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</paragraph>
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<paragraph id="8BD1204937106D13830705ACFD6E7393" blockId="13.[125,1173,1019,1235]" pageId="13" pageNumber="1418">
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<emphasis id="B91AFC5B37106D13830705ACFF337353" box="[125,200,1019,1043]" italics="true" pageId="13" pageNumber="1418">Scales.</emphasis>
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Body with rather large ctenoid scales, anterior part of predorsal area naked; scales on posterior predorsal region and belly cycloid; scales in longitudinal series 26–27 (mode 26); transverse series seven to eight (mode seven); predorsal median series 8–11; series between first dorsal and upper pectoral fin origin seven to eight (mode seven) (frequency distribution in Table III). Head including opercle, preopercle, and prepelvic areas naked. Predorsal squamation with slightly trifurcate anterior edge, anterior extension of median series reaching beyond the vertical of pores
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<emphasis id="B91AFC5B37106D13810502ECFD777393" box="[639,652,1211,1235]" italics="true" pageId="13" pageNumber="1418">H</emphasis>
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.
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</paragraph>
|
||
<paragraph id="8BD1204937106D13830702ABFC9D72F4" blockId="13.[125,1173,1276,1556]" pageId="13" pageNumber="1418">
|
||
<emphasis id="B91AFC5B37106D13830702ABFE617254" box="[125,410,1276,1300]" italics="true" pageId="13" pageNumber="1418">Head lateral-line system.</emphasis>
|
||
Canals: nasal extension of anterior oculoscapular canal with terminal pores
|
||
<emphasis id="B91AFC5B37106D138240034BFEB27274" box="[314,329,1308,1332]" italics="true" pageId="13" pageNumber="1418">S</emphasis>
|
||
slightly in front of posterior nostril. Anterior interorbital section of oculoscapular canal separated, with paired pores
|
||
<emphasis id="B91AFC5B37106D1381CF036BFD397214" box="[693,706,1340,1364]" italics="true" pageId="13" pageNumber="1418">l</emphasis>
|
||
. A single pore
|
||
<emphasis id="B91AFC5B37106D13800B036BFC7B7214" box="[881,896,1340,1364]" italics="true" pageId="13" pageNumber="1418">K</emphasis>
|
||
in posterior interorbital region. Pore
|
||
<emphasis id="B91AFC5B37106D138265030BFEC97234" box="[287,306,1372,1396]" italics="true" pageId="13" pageNumber="1418">v</emphasis>
|
||
present at posterior edge of eye. Gap between anterior and posterior oculoscapular canals larger than the length of posterior oculoscapular canal. Preopercular canal usually with three pores
|
||
<emphasis id="B91AFC5B37106D13829903CBFE1472F4" box="[483,495,1436,1460]" italics="true" pageId="13" pageNumber="1418">c</emphasis>
|
||
,
|
||
<emphasis id="B91AFC5B37106D13828503CBFDF772F4" box="[511,524,1436,1460]" italics="true" pageId="13" pageNumber="1418">d</emphasis>
|
||
, E but few males lack pore
|
||
<emphasis id="B91AFC5B37106D13802903CBFCA472F4" box="[851,863,1436,1460]" italics="true" pageId="13" pageNumber="1418">c</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD1204937106D1383E203EBFDAF7154" blockId="13.[125,1173,1276,1556]" pageId="13" pageNumber="1418">
|
||
Sensory papillae: row
|
||
<emphasis id="B91AFC5B37106D1382EF03EAFE587294" box="[405,419,1469,1492]" italics="true" pageId="13" pageNumber="1418">a</emphasis>
|
||
extending to midline of orbit. Length of row
|
||
<emphasis id="B91AFC5B37106D1380D103EBFC4C7294" box="[939,951,1468,1492]" italics="true" pageId="13" pageNumber="1418">b</emphasis>
|
||
much smaller than orbit. Row
|
||
<emphasis id="B91AFC5B37106D138272038AFEE972B4" box="[264,274,1501,1524]" italics="true" pageId="13" pageNumber="1418">c</emphasis>
|
||
,
|
||
<emphasis id="B91AFC5B37106D13825F038BFEC972B4" box="[293,306,1500,1524]" italics="true" pageId="13" pageNumber="1418">d</emphasis>
|
||
long, not reaching vertical of pore
|
||
<emphasis id="B91AFC5B37106D138198038AFD1472B5" box="[738,751,1501,1525]" italics="true" pageId="13" pageNumber="1418">a</emphasis>
|
||
. A single
|
||
<emphasis id="B91AFC5B37106D138014038AFC7E72B4" box="[878,901,1501,1524]" italics="true" pageId="13" pageNumber="1418">cp</emphasis>
|
||
papilla. Row
|
||
<emphasis id="B91AFC5B37106D13874B038BFBC372B4" box="[1073,1080,1500,1524]" italics="true" pageId="13" pageNumber="1418">f</emphasis>
|
||
paired. Opercular rows
|
||
<emphasis id="B91AFC5B37106D13824003AAFEB57154" box="[314,334,1533,1556]" italics="true" pageId="13" pageNumber="1418">ot</emphasis>
|
||
and
|
||
<emphasis id="B91AFC5B37106D1382F103ABFE657154" box="[395,414,1532,1556]" italics="true" pageId="13" pageNumber="1418">oi</emphasis>
|
||
well separated.
|
||
</paragraph>
|
||
<paragraph id="8BD1204937106D118307006AFEE37662" blockId="13.[125,1173,1597,1685]" lastBlockId="15.[125,1173,137,485]" lastPageId="15" lastPageNumber="1420" pageId="13" pageNumber="1418">
|
||
<emphasis id="B91AFC5B37106D138307006AFE827115" box="[125,377,1597,1621]" italics="true" pageId="13" pageNumber="1418">Coloration in alcohol.</emphasis>
|
||
Head and body light yellowish brown. Body without distinct dark blotches. Two major longitudinal rows of brownish black spots on lower half of lateral trunk in male, but indistinct merely with darker brown margins of scale pockets. Dorsal side of snout with a pair of brown lines united at tip of snout. A longitudinal brown stripe behind orbit. Cheek and opercle with two rows of longitudinally arranged black spots in male, none in female. A somewhat indistinct longitudinal thin brown line on middle of cheek. Branchiostegal membrane without distinct marks, uniformly light yellowish in both sexes.
|
||
</paragraph>
|
||
<caption id="DF1170C137136D1083F00028FB5971D2" ID-DOI="http://doi.org/10.5281/zenodo.5214694" ID-Zenodo-Dep="5214694" box="[138,1186,1663,1682]" httpUri="https://zenodo.org/record/5214694/files/figure.png" pageId="14" pageNumber="1419" startId="14.[138,198,1663,1682]" targetBox="[235,1092,195,1625]" targetPageId="14">
|
||
<paragraph id="8BD1204937136D1083F00028FB5971D2" blockId="14.[138,1186,1663,1682]" box="[138,1186,1663,1682]" pageId="14" pageNumber="1419">
|
||
Figure 5. Head lateral-line system of
|
||
<taxonomicName id="4C6E5BCA37136D10829C0028FD5F71D2" authorityName="Chen & Kottelat" authorityYear="2005" box="[486,676,1663,1682]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="1419" phylum="Chordata" rank="species" species="virgigena">
|
||
<emphasis id="B91AFC5B37136D10829C0028FD5F71D2" box="[486,676,1663,1682]" italics="true" pageId="14" pageNumber="1419">Rhinogobius virgigena</emphasis>
|
||
</taxonomicName>
|
||
, holotype, ZRC 49204, 28.0 mm SL. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD1204937126D1183E2077CFD5276A5" blockId="15.[125,1173,137,485]" pageId="15" pageNumber="1420">First dorsal fin with light margin and a dark spot in front of third spinous ray in male, none in female. Second dorsal fin with two basal rows of blackish brown spots and greyish distal region in male, whitish with three rows of longitudinal brown spots in female. Pectoral fin whitish, base with two conspicuous black spots in male, a single brown spot in female. Caudal fin with three to five vertical rows of greyish brown lines and spots, base with a median black spot. Pelvic fin pale white.</paragraph>
|
||
<caption id="DF1170C137126D11830700D7FB6E71D3" ID-DOI="http://doi.org/10.5281/zenodo.5214696" ID-Zenodo-Dep="5214696" box="[125,1173,1664,1683]" httpUri="https://zenodo.org/record/5214696/files/figure.png" pageId="15" pageNumber="1420" startId="15.[125,185,1664,1683]" targetBox="[185,1120,585,1622]" targetPageId="15">
|
||
<paragraph id="8BD1204937126D11830700D7FB6E71D3" blockId="15.[125,1173,1664,1683]" box="[125,1173,1664,1683]" pageId="15" pageNumber="1420">
|
||
Figure 6.
|
||
<taxonomicName id="4C6E5BCA37126D1183A600D7FE9171D3" authorityName="Chen & Kottelat" authorityYear="2005" box="[220,362,1664,1683]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="1420" phylum="Chordata" rank="species" species="boa">
|
||
<emphasis id="B91AFC5B37126D1183A600D7FE9171D3" box="[220,362,1664,1683]" italics="true" pageId="15" pageNumber="1420">Rhinogobius boa</emphasis>
|
||
</taxonomicName>
|
||
, holotype, ZRC 49206, 27.7 mm SL, cephalic colour pattern of male. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD12049370D6D0E83F006DEFEF077E1" blockId="16.[138,267,137,161]" box="[138,267,137,161]" pageId="16" pageNumber="1421">
|
||
<emphasis id="B91AFC5B370D6D0E83F006DEFEF077E1" box="[138,267,137,161]" italics="true" pageId="16" pageNumber="1421">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD12049370D6D0E83F006EEFD547673" blockId="16.[138,1185,185,307]" pageId="16" pageNumber="1421">
|
||
<taxonomicName id="4C6E5BCA370D6D0E83F006EEFE877791" authorityName="Chen & Kottelat" authorityYear="2005" box="[138,380,185,209]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="1421" phylum="Chordata" rank="species" species="variolatus">
|
||
<emphasis id="B91AFC5B370D6D0E83F006EEFE877791" box="[138,380,185,209]" italics="true" pageId="16" pageNumber="1421">Rhinogobius variolatus</emphasis>
|
||
</taxonomicName>
|
||
is presently known only from the Ba Che drainage and an unnamed creek north of Cam Pha (
|
||
<figureCitation id="13553CCC370D6D0E82D0068DFDF177B2" box="[426,522,218,242]" captionStart="Figure 1" captionStartId="3.[125,185,1638,1657]" captionTargetBox="[173,1124,1007,1600]" captionTargetId="figure-254@3.[170,1127,1004,1601]" captionTargetPageId="3" captionText="Figure 1. Distribution of Rhinogobius boa (white triangles), R. sulcatus (diamond), R. variolatus (squares), and R. virgigena (black triangle) in north-eastern Vietnam. A, Mong Cai; B, Tien Yen; C, Ba Che; D, Mong Duong." figureDoi="http://doi.org/10.5281/zenodo.5214686" httpUri="https://zenodo.org/record/5214686/files/figure.png" pageId="16" pageNumber="1421">Figure 1</figureCitation>
|
||
). It was collected in a small, shallow coastal stream with a gravel to boulder substratum, and from the Ba Che river, which in the rainy season was flowing swift, turbid water over large boulders.
|
||
</paragraph>
|
||
<caption id="DF1170C1370D6D0E83F00032FF3E71D2" ID-DOI="http://doi.org/10.5281/zenodo.5214698" ID-Zenodo-Dep="5214698" httpUri="https://zenodo.org/record/5214698/files/figure.png" pageId="16" pageNumber="1421" startId="16.[138,198,1637,1656]" targetBox="[187,1136,443,1594]" targetPageId="16">
|
||
<paragraph id="8BD12049370D6D0E83F00032FF3E71D2" blockId="16.[138,1186,1637,1682]" pageId="16" pageNumber="1421">
|
||
Figure 7.
|
||
<emphasis id="B91AFC5B370D6D0E83910032FE5C7138" box="[235,423,1637,1656]" italics="true" pageId="16" pageNumber="1421">
|
||
<taxonomicName id="4C6E5BCA370D6D0E83910032FE587138" authorityName="Chen & Kottelat" authorityYear="2005" box="[235,419,1637,1656]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="1421" phylum="Chordata" rank="species" species="sulcatus">Rhinogobius sulcatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype, ZRC 49208, 27.0 mm SL, cephalic colour pattern of male. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD12049370C6D0F830706DEFF1577E1" blockId="17.[125,238,137,161]" box="[125,238,137,161]" pageId="17" pageNumber="1422">
|
||
<emphasis id="B91AFC5B370C6D0F830706DEFF1577E1" box="[125,238,137,161]" italics="true" pageId="17" pageNumber="1422">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD12049370C6D0F830706EEFCAA77B2" blockId="17.[125,1172,185,242]" pageId="17" pageNumber="1422">
|
||
From the latin
|
||
<emphasis id="B91AFC5B370C6D0F825106EEFE8C7791" box="[299,375,185,209]" italics="true" pageId="17" pageNumber="1422">variola</emphasis>
|
||
(‘‘spotted disease’’, smallpox), allusion to the spots on the cheek and opercle. An adjective (feminine:
|
||
<emphasis id="B91AFC5B370C6D0F8281068DFDA677B2" box="[507,605,218,242]" italics="true" pageId="17" pageNumber="1422">variolata</emphasis>
|
||
; neuter:
|
||
<emphasis id="B91AFC5B370C6D0F81B3068DFCBB77B2" box="[713,832,218,242]" italics="true" pageId="17" pageNumber="1422">variolatum</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD12049370C6D0F8307077CFF277603" blockId="17.[125,220,299,323]" box="[125,220,299,323]" pageId="17" pageNumber="1422">
|
||
<emphasis id="B91AFC5B370C6D0F8307077CFF277603" box="[125,220,299,323]" italics="true" pageId="17" pageNumber="1422">Remarks</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37473C2370C6D0F8307070CFD1E7576" pageId="17" pageNumber="1422" type="reference_group">
|
||
<paragraph id="8BD12049370C6D0F8307070CFD1E7576" blockId="17.[125,1173,347,566]" pageId="17" pageNumber="1422">
|
||
<taxonomicName id="4C6E5BCA370C6D0F8307070CFE8F7633" ID-CoL="4S936" authorityName="Chen & Kottelat" authorityYear="2005" box="[125,372,347,371]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="1422" phylum="Chordata" rank="species" species="variolatus">
|
||
<emphasis id="B91AFC5B370C6D0F8307070CFE8F7633" box="[125,372,347,371]" italics="true" pageId="17" pageNumber="1422">Rhinogobius variolatus</emphasis>
|
||
</taxonomicName>
|
||
has more similarities with
|
||
<taxonomicName id="4C6E5BCA370C6D0F81D3070BFBE57633" authority="Chen et al., 2002" authorityName="Chen" authorityYear="2002" box="[681,1054,347,371]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="1422" phylum="Chordata" rank="species" species="linshuienesis">
|
||
<emphasis id="B91AFC5B370C6D0F81D3070BFCB57633" box="[681,846,347,371]" italics="true" pageId="17" pageNumber="1422">R. linshuienesis</emphasis>
|
||
<bibRefCitation id="EFFF5DB8370C6D0F802F070CFBE57633" author="Chen I-S & Miller PJ & Wu HL & Fang LS" box="[853,1054,347,371]" pageId="17" pageNumber="1422" pagination="259 - 273" refId="ref8564" refString="Chen I-S, Miller PJ, Wu HL, Fang LS. 2002. Taxonomy and mitochondrial sequence evolution in non-diadromous species of Rhinogobius (Teleostei: Gobiidae) of Hainan Island, southern China. Marine and Freshwater Research 53: 259 - 273." type="journal article" year="2002">Chen et al., 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
(a species endemic to
|
||
<collectingRegion id="49AAEEAB370C6D0F827D072BFEA676D4" box="[263,349,380,404]" country="China" name="Hainan" pageId="17" pageNumber="1422">Hainan</collectingRegion>
|
||
island) than with any other congener in southern
|
||
<collectingCountry id="F37960D9370C6D0F80DD072BFC1576D4" box="[935,1006,380,404]" name="China" pageId="17" pageNumber="1422">China</collectingCountry>
|
||
and
|
||
<collectingCountry id="F37960D9370C6D0F8750072BFB6A76D4" box="[1066,1169,380,404]" name="Vietnam" pageId="17" pageNumber="1422">Vietnam</collectingCountry>
|
||
. They can be distinguished by the following features: (1) scale counts:
|
||
<taxonomicName id="4C6E5BCA370C6D0F80AB07CAFB9976F4" ID-CoL="4S936" authorityName="Chen & Kottelat" authorityYear="2005" box="[977,1122,412,436]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="1422" phylum="Chordata" rank="species" species="variolatus">
|
||
<emphasis id="B91AFC5B370C6D0F80AB07CAFB9976F4" box="[977,1122,412,436]" italics="true" pageId="17" pageNumber="1422">R. variolatus</emphasis>
|
||
</taxonomicName>
|
||
has fewer longitudinal scales than
|
||
<taxonomicName id="4C6E5BCA370C6D0F829107EAFD707695" ID-CoL="4S8ZR" authorityName="Chen, Miller, Wu & Fang" authorityYear="2002" box="[491,651,445,469]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="1422" phylum="Chordata" rank="species" species="linshuiensis">
|
||
<emphasis id="B91AFC5B370C6D0F829107EAFD707695" box="[491,651,445,469]" italics="true" pageId="17" pageNumber="1422">R. linshuiensis</emphasis>
|
||
</taxonomicName>
|
||
(26–27 versus 29–30); (2) male coloration pattern: cheek with two rows of black spots in
|
||
<taxonomicName id="4C6E5BCA370C6D0F81E20789FCDE76B5" ID-CoL="4S936" authorityName="Chen & Kottelat" authorityYear="2005" box="[664,805,477,501]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="1422" phylum="Chordata" rank="species" species="variolatus">
|
||
<emphasis id="B91AFC5B370C6D0F81E20789FCDE76B5" box="[664,805,477,501]" italics="true" pageId="17" pageNumber="1422">R. variolatus</emphasis>
|
||
</taxonomicName>
|
||
versus a single row of red spots in
|
||
<taxonomicName id="4C6E5BCA370C6D0F83E107A9FECE7556" ID-CoL="4S8ZR" authorityName="Chen, Miller, Wu & Fang" authorityYear="2002" box="[155,309,510,534]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="1422" phylum="Chordata" rank="species" species="linshuiensis">
|
||
<emphasis id="B91AFC5B370C6D0F83E107A9FECE7556" box="[155,309,510,534]" italics="true" pageId="17" pageNumber="1422">R. linshuiensis</emphasis>
|
||
</taxonomicName>
|
||
; caudal fin base with a single median black spot in
|
||
<taxonomicName id="4C6E5BCA370C6D0F80FA07A9FBF67556" ID-CoL="4S936" authorityName="Chen & Kottelat" authorityYear="2005" box="[896,1037,510,534]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="1422" phylum="Chordata" rank="species" species="variolatus">
|
||
<emphasis id="B91AFC5B370C6D0F80FA07A9FBF67556" box="[896,1037,510,534]" italics="true" pageId="17" pageNumber="1422">R. variolatus</emphasis>
|
||
</taxonomicName>
|
||
versus with two separated, greyish brown spots in
|
||
<taxonomicName id="4C6E5BCA370C6D0F813A0448FD267576" ID-CoL="4S8ZR" authorityName="Chen, Miller, Wu & Fang" authorityYear="2002" box="[576,733,542,566]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="1422" phylum="Chordata" rank="species" species="linshuiensis">
|
||
<emphasis id="B91AFC5B370C6D0F813A0448FD267576" box="[576,733,542,566]" italics="true" pageId="17" pageNumber="1422">R. linshuiensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |