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402 lines
85 KiB
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<document id="CDE5F4451979E0E3E6C62596D808FB27" ID-DOI="10.11646/zootaxa.5318.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8162469" ID-ZooBank="CF8E0B91-9AF7-4075-963D-9BC977B41852" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1689748652228" checkinUser="plazi" docAuthor="Gibson, Gary A. P. & Fusu, Lucian" docDate="2023" docId="03C7737CA60EFFB9FF39FDEEFEFFFB05" docLanguage="en" docName="zootaxa.5318.2.2.pdf" docOrigin="Zootaxa 5318 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5318.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptoomidae Gibson 2023, fam. nov." docType="treatment" docVersion="3" lastPageNumber="206" masterDocId="FFFE0B04A607FFB2FFAEFFCCFFE5FFA5" masterDocTitle="Leptoomidae, a new family of Eocene fossil Chalcidoidea (Hymenoptera), and family classification of Eocene fossil genera originally described in Neanastatinae (Eupelmidae)" masterLastPageNumber="216" masterPageNumber="195" pageNumber="204" updateTime="1689856743257" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="166C6E8547A845769EA24934E2067F97">Leptoomidae, a new family of Eocene fossil Chalcidoidea (Hymenoptera), and family classification of Eocene fossil genera originally described in Neanastatinae (Eupelmidae)</mods:title>
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<mods:name id="36B3D8DC2EDE811715B4A4127226E6CD" type="personal">
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<mods:namePart id="080314E12A6B6B7532818FD54ADF50A2">Gibson, Gary A. P.</mods:namePart>
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<mods:affiliation id="8753DD4C523DBA1D3B916CAB70BD60C7">Honorary Research Associate, Agriculture and Agri-Food Canada, Canadian National Collection of Insects, Arachnids and Nematodes, K. W. Neatby Bldg., 960 Carling Avenue, Ottawa, Ontario, CANADA, K 1 A 0 C 6.</mods:affiliation>
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<mods:namePart id="9238F178A23663678604270F5861DFB8">Fusu, Lucian</mods:namePart>
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<mods:affiliation id="ECDA86F1E3C2023D7D8B69D34C59ED07">Al. I. Cuza University, Faculty of Biology, Bd. Carol I, Nr 11, 700506 Iasi, ROMANIA.</mods:affiliation>
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<mods:typeOfResource id="73630D3387BB3874902E0D4EBBE8537D">text</mods:typeOfResource>
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<mods:title id="F2D5D5B9133E3BE825510C965CDFD5A2">Zootaxa</mods:title>
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<mods:part id="B13709806C16C4873E6A81FF60B0E70F">
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<mods:date id="A92B5422B90578DB0D260779F943DDEE">2023</mods:date>
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<mods:number id="E22B43BD0B1359BE701215E7C02A2AB8">2023-07-19</mods:number>
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<mods:number id="D7E6B9D357F6076CF937579156519B66">5318</mods:number>
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<mods:url id="AAB72A721B35E020D49F5166DD1A44C1">http://dx.doi.org/10.11646/zootaxa.5318.2.2</mods:url>
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<mods:classification id="6FA6D2BA89BDFEFCA0FA53EE97607F9D">journal article</mods:classification>
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<mods:identifier id="B56525C216A98A1A9439C888011BBD60" type="DOI">10.11646/zootaxa.5318.2.2</mods:identifier>
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<mods:identifier id="358E8A97321054E81ADAC64AB7AE1024" type="ISSN">1175-5326</mods:identifier>
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<paragraph id="8BD1C26AA60EFFBBFF39FDEEFDF2FD98" blockId="9.[151,1163,546,609]" box="[151,535,546,573]" pageId="9" pageNumber="204">
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<taxonomicName id="4C6EB9E9A60EFFBBFF39FDEEFE78FD99" authority="Gibson" authorityName="Gibson" authorityYear="2023" box="[151,413,546,572]" family="Leptoomidae" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="204" rank="family" status="fam. nov.">Leptoomidae Gibson</taxonomicName>
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<taxonomicNameLabel id="A229A303A60EFFBBFE0AFDEEFDF2FD98" box="[420,535,546,573]" pageId="9" pageNumber="204" rank="family">fam. nov.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="8BD1C26AA60EFFBBFF39FD8AFB6EFDC4" blockId="9.[151,1163,546,609]" box="[151,1163,582,609]" pageId="9" pageNumber="204">
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="C37491E1A60EFFBBFF39FD43FC7FFD68" pageId="9" pageNumber="204" type="type_taxon">
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<paragraph id="8BD1C26AA60EFFBBFF39FD43FDA0FD0C" blockId="9.[151,1437,655,2013]" box="[151,581,655,681]" pageId="9" pageNumber="204">
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<emphasis id="B91A1E78A60EFFBBFF39FD43FEF8FD0C" bold="true" box="[151,285,655,681]" pageId="9" pageNumber="204">
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<typeStatus id="54D57CC8A60EFFBBFF39FD43FF37FD0C" box="[151,210,655,681]" pageId="9" pageNumber="204">Type</typeStatus>
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genus
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</emphasis>
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.
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<taxonomicName id="4C6EB9E9A60EFFBBFE85FD43FDA4FD0C" authority="Gibson 2008" authorityName="Gibson" authorityYear="2008" box="[299,577,655,681]" class="Insecta" family="Tanaostigmatidae" genus="Leptoomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="204" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A1E78A60EFFBBFE85FD43FE43FD0D" box="[299,422,655,680]" italics="true" pageId="9" pageNumber="204">Leptoomus</emphasis>
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<bibRefCitation id="EFFFBF9BA60EFFBBFE03FD43FDA4FD0C" author="Gibson, G. A. P." box="[429,577,655,681]" pageId="9" pageNumber="204" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BD1C26AA60EFFBBFF69FD7FFC7FFD68" blockId="9.[151,1437,655,2013]" box="[199,922,691,717]" pageId="9" pageNumber="204">
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<emphasis id="B91A1E78A60EFFBBFF69FD7FFE6CFD68" bold="true" box="[199,393,691,717]" pageId="9" pageNumber="204">Included genera</emphasis>
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.
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<taxonomicName id="4C6EB9E9A60EFFBBFE39FD7FFDF7FD69" authorityName="Gibson" authorityYear="2008" box="[407,530,691,716]" class="Insecta" family="Tanaostigmatidae" genus="Leptoomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="204" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A1E78A60EFFBBFE39FD7FFDF7FD69" box="[407,530,691,716]" italics="true" pageId="9" pageNumber="204">Leptoomus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6EB9E9A60EFFBBFDE6FD7FFC72FD68" authority="Gibson, 2009" authorityName="Gibson" authorityYear="2009" box="[584,919,691,717]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="204" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A1E78A60EFFBBFDE6FD7FFD10FD68" box="[584,757,691,717]" italics="true" pageId="9" pageNumber="204">Neanaperiallus</emphasis>
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<bibRefCitation id="EFFFBF9BA60EFFBBFD52FD7FFC72FD68" author="Gibson, G. A. P." box="[764,919,691,717]" pageId="9" pageNumber="204" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson, 2009</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<subSubSection id="C37491E1A60EFFBBFF69FD1AFD0DFBB4" pageId="9" pageNumber="204" type="diagnosis">
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<paragraph id="8BD1C26AA60EFFBBFF69FD1AFD0DFBB4" blockId="9.[151,1437,655,2013]" pageId="9" pageNumber="204">
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<emphasis id="B91A1E78A60EFFBBFF69FD1AFEDFFD55" bold="true" box="[199,314,726,752]" pageId="9" pageNumber="204">Diagnosis</emphasis>
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. Antenna 12- or 13-segmented, with basal flagellomere not distinctly anelliform and clava 3- segmented. Mesoscutum with notaular furrows extending along length. Prepectus anteriorly rounded to angulate, extending to or slightly over posterolateral margin of pronotum; dorsal margin of prepectus intersecting base of the tegula distinctly anterior to and forming an almost right-angle with posterior margin of prepectus; posterior prepectal margin truncate along anterior margin of acropleuron.Acropleuron greatly enlarged, extending posteriorly to metapleuron and ventrally at least to anterodorsal margin of mesocoxa. In lateral view mesocoxa with anterior margin distinctly posterior to midline of acropleuron. Mesopectus with posterior margin abutting bases of mesocoxae. Mesotibia with 1 or 2 rows of apical pegs and mesotarsus with spine- to peg-like setae along both ventral margins. Fore wing with oblique bare band beyond basal fold.
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</paragraph>
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<subSubSection id="C37491E1A60EFFB9FF69FBD6FD17FE74" lastPageId="11" lastPageNumber="206" pageId="9" pageNumber="204" type="description">
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<paragraph id="8BD1C26AA60EFFB9FF69FBD6FD17FE74" blockId="9.[151,1437,655,2013]" lastBlockId="11.[151,1437,150,1184]" lastPageId="11" lastPageNumber="206" pageId="9" pageNumber="204">
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<emphasis id="B91A1E78A60EFFBBFF69FBD6FEB5FB91" bold="true" box="[199,336,1050,1076]" pageId="9" pageNumber="204">Description</emphasis>
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. Head with malar sulcus (
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<figureCitation id="1355DEEFA60EFFBBFD2DFBD7FD07FB90" box="[643,738,1051,1077]" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G & H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="9" pageNumber="204">Figs 2G</figureCitation>
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,
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<figureCitation id="1355DEEFA60EFFBBFD5FFBD7FCF6FB90" box="[753,787,1051,1077]" captionStart="FIGURE 3" captionStartId="13.[152,255,1776,1800]" captionTargetBox="[151,1436,181,1752]" captionTargetId="figure-19@13.[151,1436,181,1752]" captionTargetPageId="13" captionText="FIGURE 3. A‒C, Leptoomus janzeni, specimen LF2002: A, lateral habitus (mirrored image, arrow points to posterior margin of mesocoxa); B, head in frontal view; C, ventral habitus (mirrored image). D & E, Aspidopleura baltica, holotype: D, head and mesosoma, dorsal; E, mesosoma, dorsolateral. F & G, Metapelma sp.: F, mesosoma, dorsal; G, mesosoma, lateral. H, Calosota aestivalis, mesosoma, dorsolateral. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162476" httpUri="https://zenodo.org/record/8162476/files/figure.png" pageId="9" pageNumber="204">3B</figureCitation>
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,
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<figureCitation id="1355DEEFA60EFFBBFC8DFBD6FCA0FB90" box="[803,837,1050,1077]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="9" pageNumber="204">5E</figureCitation>
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); clypeus with apical margin straight transverse and not delimited by a subapical transverse groove (
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<figureCitation id="1355DEEFA60EFFBBFD1DFBF3FCF4FBFC" box="[691,785,1087,1113]" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G & H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="9" pageNumber="204">Figs 2G</figureCitation>
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,
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<figureCitation id="1355DEEFA60EFFBBFCB0FBF2FC85FBFC" box="[798,864,1086,1113]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="9" pageNumber="204">5A, E</figureCitation>
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inset). Antenna 12- or 13-segmented with 7- (
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<figureCitation id="1355DEEFA60EFFBBFAC5FBF3FF59FBD8" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G & H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="9" pageNumber="204">Figs 2G</figureCitation>
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,
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<figureCitation id="1355DEEFA60EFFBBFF66FBAFFF0CFBD8" box="[200,233,1123,1149]" captionStart="FIGURE 3" captionStartId="13.[152,255,1776,1800]" captionTargetBox="[151,1436,181,1752]" captionTargetId="figure-19@13.[151,1436,181,1752]" captionTargetPageId="13" captionText="FIGURE 3. A‒C, Leptoomus janzeni, specimen LF2002: A, lateral habitus (mirrored image, arrow points to posterior margin of mesocoxa); B, head in frontal view; C, ventral habitus (mirrored image). D & E, Aspidopleura baltica, holotype: D, head and mesosoma, dorsal; E, mesosoma, dorsolateral. F & G, Metapelma sp.: F, mesosoma, dorsal; G, mesosoma, lateral. H, Calosota aestivalis, mesosoma, dorsolateral. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162476" httpUri="https://zenodo.org/record/8162476/files/figure.png" pageId="9" pageNumber="204">3B</figureCitation>
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) or 8- (
|
||
<figureCitation id="1355DEEFA60EFFBBFE90FBAFFE5CFBD8" box="[318,441,1123,1149]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C & E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D & H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="9" pageNumber="204">Figs 4B, C</figureCitation>
|
||
) segmented funicle and 3-segmented clava; basal flagellomere lacking multiporous plate sensilla and the shortest funicular, subquadrate but not distinctly anelliform. Pronotum with strongly inclined neck abruptly merging into very short, transverse collar (
|
||
<figureCitation id="1355DEEFA60EFFBBFD63FB67FCCDFB60" box="[717,808,1195,1221]" captionStart="FIGURE 1" captionStartId="10.[152,255,1835,1859]" captionTargetBox="[152,1436,180,1809]" captionTargetId="figure-25@10.[151,1436,179,1809]" captionTargetPageId="10" captionText="FIGURE 1. Lateral view of mesosoma: A, Neanaperiallus masneri; B, Leptoomus janzeni; C, Tanaostigmodes sp.; D, Cynipencyrtus ishii; E, Eopelma gibsoni; F, Savzdargia sp.; G, Neanastatus sp.; H, Lambdobregma sp.; I, Brevivulva electroma. Insets in H and I are dorsal views of apex of mesoscutellum to propodeum; green arrows on images indicate the intersection of base of tegula with dorsal margin of prepectus, white arrows indicate posterior margin of prepectus; see ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162471" httpUri="https://zenodo.org/record/8162471/files/figure.png" pageId="9" pageNumber="204">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="1355DEEFA60EFFBBFC9AFB67FC93FB60" box="[820,886,1195,1221]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C & E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D & H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="9" pageNumber="204">4E, G</figureCitation>
|
||
,
|
||
<figureCitation id="1355DEEFA60EFFBBFC2FFB66FC44FB60" box="[897,929,1194,1221]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="9" pageNumber="204">5F</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFC02FB67FBD8FB60" author="Gibson, G. A. P." box="[940,1085,1195,1221]" pageId="9" pageNumber="204" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, figs 6‒8;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFB01FB67FADAFB60" author="Gibson, G. A. P." box="[1199,1343,1195,1221]" pageId="9" pageNumber="204" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
|
||
, figs 59, 60;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFF6CFB03FE4DFB4C" author="Simutnik, S. A. & Perkovsky, E. E. & Vasilenko, D. V." box="[194,424,1231,1257]" pageId="9" pageNumber="204" pagination="137 - 145" refId="ref17870" refString="Simutnik, S. A., Perkovsky, E. E. & Vasilenko, D. V. (2020) First record of Leptoomus janzeni Gibson (Hymenoptera, Chalcidoidea) from Rovno amber. Journal of Hymenoptera Research, 80, 137 - 145. https: // doi. org / 10.3897 / jhr. 80.58882" type="journal article" year="2020">
|
||
Simutnik
|
||
<emphasis id="B91A1E78A60EFFBBFE81FB03FE82FB4C" box="[303,359,1231,1257]" italics="true" pageId="9" pageNumber="204">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
, figs 1C, D) and without a median line (
|
||
<figureCitation id="1355DEEFA60EFFBBFCC6FB03FC01FB4C" box="[872,996,1231,1257]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C & E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D & H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="9" pageNumber="204">Figs 4E, G</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFC5EFB03FB64FB4C" author="Gibson, G. A. P." box="[1008,1153,1231,1257]" pageId="9" pageNumber="204" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, fig. 4). Mesonotum with transscutal articulation straight, articulated along its entire width (
|
||
<figureCitation id="1355DEEFA60EFFBBFCC8FB3FFC1AFAA8" box="[870,1023,1267,1293]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C & E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D & H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="9" pageNumber="204">Figs 4E, G, H</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFBA4FB3FFB7CFAA8" author="Gibson, G. A. P." box="[1034,1177,1267,1293]" pageId="9" pageNumber="204" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, figs 7, 8;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFAA6FB3FFA7DFAA8" author="Gibson, G. A. P." box="[1288,1432,1267,1293]" pageId="9" pageNumber="204" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
|
||
, fig. 59;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFF5DFADBFE05FA94" author="Simutnik, S. A. & Perkovsky, E. E. & Vasilenko, D. V." box="[243,480,1303,1329]" pageId="9" pageNumber="204" pagination="137 - 145" refId="ref17870" refString="Simutnik, S. A., Perkovsky, E. E. & Vasilenko, D. V. (2020) First record of Leptoomus janzeni Gibson (Hymenoptera, Chalcidoidea) from Rovno amber. Journal of Hymenoptera Research, 80, 137 - 145. https: // doi. org / 10.3897 / jhr. 80.58882" type="journal article" year="2020">
|
||
Simutnik
|
||
<emphasis id="B91A1E78A60EFFBBFECDFADBFE78FA94" box="[355,413,1303,1329]" italics="true" pageId="9" pageNumber="204">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
, fig. 1D); mesoscutum without parapsidal lines or sulcate notauli, but with notaular furrow extending from mesothoracic spiracle posterior of pronotum to transscutal articulation at anterolateral margin of axilla, the furrows delineating median and lateral mesoscutal lobes, with median lobe uniformly convex, not differentiated into anteromedian convex and posteromedian depressed region (
|
||
<figureCitation id="1355DEEFA60EFFBBFC40FA4FFBACFA38" box="[1006,1097,1411,1437]" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G & H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="9" pageNumber="204">Figs 2H</figureCitation>
|
||
,
|
||
<figureCitation id="1355DEEFA60EFFBBFBFCFA4FFB52FA38" box="[1106,1207,1411,1437]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C & E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D & H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="9" pageNumber="204">4A, E‒G</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFB6EFA4FFAAAFA38" author="Gibson, G. A. P." box="[1216,1359,1411,1437]" pageId="9" pageNumber="204" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, figs 6, 7;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFF1CFA6BFE76FA64" author="Simutnik, S. A. & Perkovsky, E. E. & Vasilenko, D. V." box="[178,403,1447,1473]" pageId="9" pageNumber="204" pagination="137 - 145" refId="ref17870" refString="Simutnik, S. A., Perkovsky, E. E. & Vasilenko, D. V. (2020) First record of Leptoomus janzeni Gibson (Hymenoptera, Chalcidoidea) from Rovno amber. Journal of Hymenoptera Research, 80, 137 - 145. https: // doi. org / 10.3897 / jhr. 80.58882" type="journal article" year="2020">
|
||
Simutnik
|
||
<emphasis id="B91A1E78A60EFFBBFEB3FA6BFEB1FA64" box="[285,340,1447,1473]" italics="true" pageId="9" pageNumber="204">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
, fig. 1D); mesoscutellar-axillar complex with axillae triangular, their inner angles separated by narrowly truncate base of mesoscutellum when mesonotum not flexed; mesoscutellum without differentiated frenum or axillular carina. Mesosoma in lateral view with convex acropleuron extending posteriorly to metapleuron and ventrally at least to anterodorsal margin of mesocoxa, with variably large mesepisternal region ventral to acropleuron (
|
||
<figureCitation id="1355DEEFA60EFFBBFF31F9FBFEFCF9F4" box="[159,281,1591,1617]" captionStart="FIGURE 1" captionStartId="10.[152,255,1835,1859]" captionTargetBox="[152,1436,180,1809]" captionTargetId="figure-25@10.[151,1436,179,1809]" captionTargetPageId="10" captionText="FIGURE 1. Lateral view of mesosoma: A, Neanaperiallus masneri; B, Leptoomus janzeni; C, Tanaostigmodes sp.; D, Cynipencyrtus ishii; E, Eopelma gibsoni; F, Savzdargia sp.; G, Neanastatus sp.; H, Lambdobregma sp.; I, Brevivulva electroma. Insets in H and I are dorsal views of apex of mesoscutellum to propodeum; green arrows on images indicate the intersection of base of tegula with dorsal margin of prepectus, white arrows indicate posterior margin of prepectus; see ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162471" httpUri="https://zenodo.org/record/8162471/files/figure.png" pageId="9" pageNumber="204">Figs 1A, B</figureCitation>
|
||
,
|
||
<figureCitation id="1355DEEFA60EFFBBFE8AF9FAFEAFF9F4" box="[292,330,1590,1617]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="9" pageNumber="204">5A</figureCitation>
|
||
). Mesopectus in ventral view with sulcate discrimen but apparently without transepisternal line (
|
||
<figureCitation id="1355DEEFA60EFFBBFAC5F9FBFF5CF9D0" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G & H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="9" pageNumber="204">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="1355DEEFA60EFFBBFF66F996FF09F9D0" box="[200,236,1626,1653]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="9" pageNumber="204">5A</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFF54F997FE03F9D0" author="Simutnik, S. A. & Perkovsky, E. E. & Vasilenko, D. V." box="[250,486,1627,1653]" pageId="9" pageNumber="204" pagination="137 - 145" refId="ref17870" refString="Simutnik, S. A., Perkovsky, E. E. & Vasilenko, D. V. (2020) First record of Leptoomus janzeni Gibson (Hymenoptera, Chalcidoidea) from Rovno amber. Journal of Hymenoptera Research, 80, 137 - 145. https: // doi. org / 10.3897 / jhr. 80.58882" type="journal article" year="2020">
|
||
Simutnik
|
||
<emphasis id="B91A1E78A60EFFBBFEC7F997FE46F9D0" box="[361,419,1627,1653]" italics="true" pageId="9" pageNumber="204">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
, fig. 2A); mesopectus with posterior margin abutting bases of mesocoxae (
|
||
<figureCitation id="1355DEEFA60EFFBBFA95F997FF49F93C" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G & H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="9" pageNumber="204">Figs 2B, C</figureCitation>
|
||
,
|
||
<figureCitation id="1355DEEFA60EFFBBFF16F9B2FF38F93C" box="[184,221,1662,1689]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="9" pageNumber="204">5A</figureCitation>
|
||
), without external membranous region between mesopectus and each mesocoxa so that mesocoxae unable to rotate anteriorly entirely out of their combined fossa. Prepectus bare; in lateral view anteriorly rounded to angulate and protuberant, extending to or slightly overlapping posterolateral margin of pronotum, with dorsal margin virtually straight and extending posteriorly distinctly beyond base of tegula (
|
||
<figureCitation id="1355DEEFA60EFFBBFC3EF927FBF4F8A0" box="[912,1041,1771,1797]" captionStart="FIGURE 1" captionStartId="10.[152,255,1835,1859]" captionTargetBox="[152,1436,180,1809]" captionTargetId="figure-25@10.[151,1436,179,1809]" captionTargetPageId="10" captionText="FIGURE 1. Lateral view of mesosoma: A, Neanaperiallus masneri; B, Leptoomus janzeni; C, Tanaostigmodes sp.; D, Cynipencyrtus ishii; E, Eopelma gibsoni; F, Savzdargia sp.; G, Neanastatus sp.; H, Lambdobregma sp.; I, Brevivulva electroma. Insets in H and I are dorsal views of apex of mesoscutellum to propodeum; green arrows on images indicate the intersection of base of tegula with dorsal margin of prepectus, white arrows indicate posterior margin of prepectus; see ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162471" httpUri="https://zenodo.org/record/8162471/files/figure.png" pageId="9" pageNumber="204">Figs 1A, B</figureCitation>
|
||
: arrows, 5A) and forming almost a right-angle relative to straight or only slightly curved posterior, truncate margin along anterior margin of acropleuron (
|
||
<figureCitation id="1355DEEFA60EFFBBFF31F8FFFEFFF8E8" box="[159,282,1843,1869]" captionStart="FIGURE 1" captionStartId="10.[152,255,1835,1859]" captionTargetBox="[152,1436,180,1809]" captionTargetId="figure-25@10.[151,1436,179,1809]" captionTargetPageId="10" captionText="FIGURE 1. Lateral view of mesosoma: A, Neanaperiallus masneri; B, Leptoomus janzeni; C, Tanaostigmodes sp.; D, Cynipencyrtus ishii; E, Eopelma gibsoni; F, Savzdargia sp.; G, Neanastatus sp.; H, Lambdobregma sp.; I, Brevivulva electroma. Insets in H and I are dorsal views of apex of mesoscutellum to propodeum; green arrows on images indicate the intersection of base of tegula with dorsal margin of prepectus, white arrows indicate posterior margin of prepectus; see ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162471" httpUri="https://zenodo.org/record/8162471/files/figure.png" pageId="9" pageNumber="204">Figs 1A, B</figureCitation>
|
||
,
|
||
<figureCitation id="1355DEEFA60EFFBBFE88F8FEFEA9F8E8" box="[294,332,1842,1869]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="9" pageNumber="204">5A</figureCitation>
|
||
); in dorsal view prepectus thin (
|
||
<figureCitation id="1355DEEFA60EFFBBFD03F8FFFCE4F8E8" box="[685,769,1843,1869]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C & E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D & H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="9" pageNumber="204">Fig. 4F</figureCitation>
|
||
: pre) to slightly convex (
|
||
<figureCitation id="1355DEEFA60EFFBBFBBDF8FFFB88F8E8" box="[1043,1133,1843,1869]" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G & H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="9" pageNumber="204">Fig. 2H</figureCitation>
|
||
: pre) but not conspicuously bulbous. Fore wing with typical venation consisting of submarginal, marginal, postmarginal and stigmal veins, and stigmal vein with distinct uncus (
|
||
<figureCitation id="1355DEEFA60EFFBBFDA6F8B7FD62F830" box="[520,647,1915,1941]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C & E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D & H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="9" pageNumber="204">Figs 4A, D</figureCitation>
|
||
,
|
||
<figureCitation id="1355DEEFA60EFFBBFD3DF8B6FD5DF830" box="[659,696,1914,1941]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="9" pageNumber="204">5G</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFD6AF8B7FCB0F830" author="Gibson, G. A. P." box="[708,853,1915,1941]" pageId="9" pageNumber="204" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, fig. 16); disc with broad, oblique bare band beyond basal fold either contiguous with (
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFDA2F853FD7BF81C" author="Gibson, G. A. P." box="[524,670,1951,1977]" pageId="9" pageNumber="204" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, fig. 16;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFD53F853FC39F81C" author="Simutnik, S. A. & Perkovsky, E. E. & Vasilenko, D. V." box="[765,988,1951,1977]" pageId="9" pageNumber="204" pagination="137 - 145" refId="ref17870" refString="Simutnik, S. A., Perkovsky, E. E. & Vasilenko, D. V. (2020) First record of Leptoomus janzeni Gibson (Hymenoptera, Chalcidoidea) from Rovno amber. Journal of Hymenoptera Research, 80, 137 - 145. https: // doi. org / 10.3897 / jhr. 80.58882" type="journal article" year="2020">
|
||
Simutnik
|
||
<emphasis id="B91A1E78A60EFFBBFCC6F853FC78F81C" box="[872,925,1951,1977]" italics="true" pageId="9" pageNumber="204">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
, fig. 1H) or separated from parastigma by setae (
|
||
<figureCitation id="1355DEEFA60EFFBBFF72F80FFEBCF878" box="[220,345,1986,2013]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="9" pageNumber="204">Figs 5G, H</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFFBF9BA60EFFBBFECAF80FFE17F878" author="Gibson, G. A. P." box="[356,498,1987,2013]" pageId="9" pageNumber="204" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
|
||
, fig. 62). Legs with mesocoxa inserted near metacoxa, distant from procoxa, in lateral view mesocoxa with anterior margin distinctly posterior to midline of acropleuron (
|
||
<figureCitation id="1355DEEFA60CFFB9FBDCFF5BFB1FFF14" box="[1138,1274,151,177]" captionStart="FIGURE 1" captionStartId="10.[152,255,1835,1859]" captionTargetBox="[152,1436,180,1809]" captionTargetId="figure-25@10.[151,1436,179,1809]" captionTargetPageId="10" captionText="FIGURE 1. Lateral view of mesosoma: A, Neanaperiallus masneri; B, Leptoomus janzeni; C, Tanaostigmodes sp.; D, Cynipencyrtus ishii; E, Eopelma gibsoni; F, Savzdargia sp.; G, Neanastatus sp.; H, Lambdobregma sp.; I, Brevivulva electroma. Insets in H and I are dorsal views of apex of mesoscutellum to propodeum; green arrows on images indicate the intersection of base of tegula with dorsal margin of prepectus, white arrows indicate posterior margin of prepectus; see ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162471" httpUri="https://zenodo.org/record/8162471/files/figure.png" pageId="11" pageNumber="206">Figs 1A, B</figureCitation>
|
||
,
|
||
<figureCitation id="1355DEEFA60CFFB9FAA4FF5BFACAFF14" box="[1290,1327,151,177]" captionStart="FIGURE 3" captionStartId="13.[152,255,1776,1800]" captionTargetBox="[151,1436,181,1752]" captionTargetId="figure-19@13.[151,1436,181,1752]" captionTargetPageId="13" captionText="FIGURE 3. A‒C, Leptoomus janzeni, specimen LF2002: A, lateral habitus (mirrored image, arrow points to posterior margin of mesocoxa); B, head in frontal view; C, ventral habitus (mirrored image). D & E, Aspidopleura baltica, holotype: D, head and mesosoma, dorsal; E, mesosoma, dorsolateral. F & G, Metapelma sp.: F, mesosoma, dorsal; G, mesosoma, lateral. H, Calosota aestivalis, mesosoma, dorsolateral. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162476" httpUri="https://zenodo.org/record/8162476/files/figure.png" pageId="11" pageNumber="206">3A</figureCitation>
|
||
); tarsi 5- segmented; protibia with bifurcate, curved tibial spur (
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FD52FF77FC74FF70" author="Gibson, G. A. P." box="[764,913,187,213]" pageId="11" pageNumber="206" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, fig. 21;
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FC5BFF77FB3BFF70" author="Simutnik, S. A. & Perkovsky, E. E. & Vasilenko, D. V." box="[1013,1246,187,213]" pageId="11" pageNumber="206" pagination="137 - 145" refId="ref17870" refString="Simutnik, S. A., Perkovsky, E. E. & Vasilenko, D. V. (2020) First record of Leptoomus janzeni Gibson (Hymenoptera, Chalcidoidea) from Rovno amber. Journal of Hymenoptera Research, 80, 137 - 145. https: // doi. org / 10.3897 / jhr. 80.58882" type="journal article" year="2020">
|
||
Simutnik
|
||
<emphasis id="B91A1E78A60CFFB9FBCAFF77FB79FF70" box="[1124,1180,187,213]" italics="true" pageId="11" pageNumber="206">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
, fig. 1I), without dorsal spicules but with dorsoapical spicule (
|
||
<figureCitation id="1355DEEFA60CFFB9FD27FF13FD3BFF5C" box="[649,734,222,249]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="11" pageNumber="206">Fig. 5A</figureCitation>
|
||
insert: arrow;
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FCD1FF13FBF4FF5C" author="Gibson, G. A. P." box="[895,1041,223,249]" pageId="11" pageNumber="206" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, fig. 21: pas); mesotibia with apical pegs arranged in 1 or 2 rows (
|
||
<figureCitation id="1355DEEFA60CFFB9FE49FECFFDD9FEB8" box="[487,572,258,285]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="11" pageNumber="206">Fig. 5C</figureCitation>
|
||
map;
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FD2CFECFFCF3FEB8" author="Gibson, G. A. P." box="[642,790,259,285]" pageId="11" pageNumber="206" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, figs 18, 19); mesotarsus ventrally with spine- to peg-like setae along both margins (
|
||
<figureCitation id="1355DEEFA60CFFB9FE14FEEBFDD3FEE4" box="[442,566,294,321]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="11" pageNumber="206">Figs 5B, C</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FDEFFEEBFD36FEE4" author="Gibson, G. A. P." box="[577,723,295,321]" pageId="11" pageNumber="206" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, figs 18, 19;
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FCCFFEEBFC16FEE4" author="Gibson, G. A. P." box="[865,1011,295,321]" pageId="11" pageNumber="206" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
|
||
, figs 64, 65); metatibia with two spurs (
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FF31FE87FE6DFEC0" author="Simutnik, S. A. & Perkovsky, E. E. & Vasilenko, D. V." box="[159,392,331,357]" pageId="11" pageNumber="206" pagination="137 - 145" refId="ref17870" refString="Simutnik, S. A., Perkovsky, E. E. & Vasilenko, D. V. (2020) First record of Leptoomus janzeni Gibson (Hymenoptera, Chalcidoidea) from Rovno amber. Journal of Hymenoptera Research, 80, 137 - 145. https: // doi. org / 10.3897 / jhr. 80.58882" type="journal article" year="2020">
|
||
Simutnik
|
||
<emphasis id="B91A1E78A60CFFB9FEA0FE87FEA2FEC0" box="[270,327,331,357]" italics="true" pageId="11" pageNumber="206">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
, fig. 2C). Gaster sessile; Gt
|
||
<subScript id="17EAC02FA60CFFB9FD12FE91FD21FECE" attach="left" box="[700,708,349,363]" fontSize="6" pageId="11" pageNumber="206">7</subScript>
|
||
and Gt
|
||
<subScript id="17EAC02FA60CFFB9FCB9FE91FCFAFECE" attach="left" box="[791,799,349,363]" fontSize="6" pageId="11" pageNumber="206">8</subScript>
|
||
fused into syntergum, though at least sometimes with an oblique groove posterior to cercus distinguishing presumptive Gt
|
||
<subScript id="17EAC02FA60CFFB9FCD9FE4DFC9AFE2A" attach="left" box="[887,895,385,399]" fontSize="6" pageId="11" pageNumber="206">7</subScript>
|
||
and Gt
|
||
<subScript id="17EAC02FA60CFFB9FC7DFE4DFC3EFE2A" attach="left" box="[979,987,385,399]" fontSize="6" pageId="11" pageNumber="206">8</subScript>
|
||
(
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FC46FEA3FB9BFE2C" author="Gibson, G. A. P." box="[1000,1150,367,393]" pageId="11" pageNumber="206" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, fig. 17); syntergum with cercus not advanced and cercal setae not kinked; ovipositor sheaths exerted for short distance beyond apex of gaster (
|
||
<figureCitation id="1355DEEFA60CFFB9FF31FE7BFF1CFE74" box="[159,249,438,465]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="11" pageNumber="206">Fig. 5D</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FEABFE7BFE7DFE74" author="Gibson, G. A. P." box="[261,408,439,465]" pageId="11" pageNumber="206" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, fig. 17;
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FE55FE7BFD6BFE74" author="Gibson, G. A. P." box="[507,654,439,465]" pageId="11" pageNumber="206" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
|
||
, fig. 63).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1192E2A60DFFB8FF36F8E7FEAEF876" ID-DOI="http://doi.org/10.5281/zenodo.8162471" ID-Zenodo-Dep="8162471" httpUri="https://zenodo.org/record/8162471/files/figure.png" pageId="10" pageNumber="205" startId="10.[152,255,1835,1859]" targetBox="[152,1436,180,1809]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8BD1C26AA60DFFB8FF36F8E7FEAEF876" blockId="10.[151,1436,1835,2003]" pageId="10" pageNumber="205">
|
||
<emphasis id="B91A1E78A60DFFB8FF36F8E7FEFDF8E6" bold="true" box="[152,280,1835,1859]" pageId="10" pageNumber="205">FIGURE 1</emphasis>
|
||
. Lateral view of mesosoma:
|
||
<emphasis id="B91A1E78A60DFFB8FDF9F8E7FD8CF8E6" bold="true" box="[599,617,1835,1859]" pageId="10" pageNumber="205">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C6EB9E9A60DFFB8FDD5F8E7FC90F8E6" authorityName="Gibson" authorityYear="2009" box="[635,885,1835,1859]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="masneri">
|
||
<emphasis id="B91A1E78A60DFFB8FDD5F8E7FC90F8E6" box="[635,885,1835,1859]" italics="true" pageId="10" pageNumber="205">Neanaperiallus masneri</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B91A1E78A60DFFB8FC29F8E7FC7DF8E6" bold="true" box="[903,920,1835,1859]" pageId="10" pageNumber="205">B</emphasis>
|
||
,
|
||
<taxonomicName id="4C6EB9E9A60DFFB8FC04F8E0FB8BF8E6" authorityName="Gibson" authorityYear="2008" box="[938,1134,1836,1859]" class="Insecta" family="Tanaostigmatidae" genus="Leptoomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="janzeni">
|
||
<emphasis id="B91A1E78A60DFFB8FC04F8E0FB8BF8E6" box="[938,1134,1836,1859]" italics="true" pageId="10" pageNumber="205">Leptoomus janzeni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B91A1E78A60DFFB8FB2FF8E7FB76F8E6" bold="true" box="[1153,1171,1835,1859]" pageId="10" pageNumber="205">C</emphasis>
|
||
,
|
||
<taxonomicName id="4C6EB9E9A60DFFB8FB0BF8E7FA91F8E6" box="[1189,1396,1835,1859]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigmodes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B91A1E78A60DFFB8FB0BF8E7FAACF8E6" box="[1189,1353,1835,1859]" italics="true" pageId="10" pageNumber="205">Tanaostigmodes</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B91A1E78A60DFFB8FA2AF8E7FA73F8E6" bold="true" box="[1412,1430,1835,1859]" pageId="10" pageNumber="205">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C6EB9E9A60DFFB8FF39F883FEBEF8C2" authorityName="Ishii" authorityYear="1928" box="[151,347,1871,1895]" class="Insecta" family="Tanaostigmatidae" genus="Cynipencyrtus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="ishii">
|
||
<emphasis id="B91A1E78A60DFFB8FF39F883FEBEF8C2" box="[151,347,1871,1895]" italics="true" pageId="10" pageNumber="205">Cynipencyrtus ishii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B91A1E78A60DFFB8FEC8F883FE92F8C2" bold="true" box="[358,375,1871,1895]" pageId="10" pageNumber="205">E</emphasis>
|
||
,
|
||
<taxonomicName id="4C6EB9E9A60DFFB8FE2DF883FDC8F8C2" box="[387,557,1871,1895]" class="Insecta" family="Eupelmidae" genus="Eopelma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="gibsoni">
|
||
<emphasis id="B91A1E78A60DFFB8FE2DF883FDC8F8C2" box="[387,557,1871,1895]" italics="true" pageId="10" pageNumber="205">Eopelma gibsoni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B91A1E78A60DFFB8FD97F883FDADF8C2" bold="true" box="[569,584,1871,1895]" pageId="10" pageNumber="205">F</emphasis>
|
||
,
|
||
<taxonomicName id="4C6EB9E9A60DFFB8FDFAF883FD0CF8C2" box="[596,745,1871,1895]" class="Insecta" family="Encyrtidae" genus="Savzdargia" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B91A1E78A60DFFB8FDFAF883FD20F8C2" box="[596,709,1871,1895]" italics="true" pageId="10" pageNumber="205">Savzdargia</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B91A1E78A60DFFB8FD5DF883FCE2F8C2" bold="true" box="[755,775,1871,1895]" pageId="10" pageNumber="205">G</emphasis>
|
||
,
|
||
<taxonomicName id="4C6EB9E9A60DFFB8FCBCF89CFC51F8C2" box="[786,948,1872,1895]" class="Insecta" family="Eupelmidae" genus="Neanastatus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B91A1E78A60DFFB8FCBCF89CFC6AF8C2" box="[786,911,1872,1895]" italics="true" pageId="10" pageNumber="205">Neanastatus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B91A1E78A60DFFB8FC13F883FC34F8C2" bold="true" box="[957,977,1871,1895]" pageId="10" pageNumber="205">H</emphasis>
|
||
,
|
||
<taxonomicName id="4C6EB9E9A60DFFB8FC73F883FB44F8C2" box="[989,1185,1871,1895]" class="Insecta" family="Eupelmidae" genus="Lambdobregma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B91A1E78A60DFFB8FC73F883FB99F8C2" box="[989,1148,1871,1895]" italics="true" pageId="10" pageNumber="205">Lambdobregma</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B91A1E78A60DFFB8FB04F883FB51F8C2" bold="true" box="[1194,1204,1871,1895]" pageId="10" pageNumber="205">I</emphasis>
|
||
,
|
||
<taxonomicName id="4C6EB9E9A60DFFB8FB6EF883FA73F8C2" authorityName="Gibson" authorityYear="2009" box="[1216,1430,1871,1895]" class="Insecta" family="Eupelmidae" genus="Brevivulva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="electroma">
|
||
<emphasis id="B91A1E78A60DFFB8FB6EF883FA73F8C2" box="[1216,1430,1871,1895]" italics="true" pageId="10" pageNumber="205">Brevivulva electroma</emphasis>
|
||
</taxonomicName>
|
||
. Insets in H and I are dorsal views of apex of mesoscutellum to propodeum; green arrows on images indicate the intersection of base of tegula with dorsal margin of prepectus, white arrows indicate posterior margin of prepectus; see ‘Material and methods’ for abbreviations.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37491E1A60CFFB9FF69FE17FEFFFB05" pageId="11" pageNumber="206" type="discussion">
|
||
<paragraph id="8BD1C26AA60CFFB9FF69FE17FB1AFC24" blockId="11.[151,1437,150,1184]" pageId="11" pageNumber="206">
|
||
<emphasis id="B91A1E78A60CFFB9FF69FE17FED6FE50" bold="true" box="[199,307,475,501]" pageId="11" pageNumber="206">Remarks</emphasis>
|
||
.
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FEEEFE17FE04FE50" author="Gibson, G. A. P." box="[320,481,475,501]" pageId="11" pageNumber="206" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson (2009)</bibRefCitation>
|
||
described the mesoscutum of
|
||
<taxonomicName id="4C6EB9E9A60CFFB9FC83FE17FC43FE51" authorityName="Gibson" authorityYear="2009" box="[813,934,475,500]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="masneri">
|
||
<emphasis id="B91A1E78A60CFFB9FC83FE17FC43FE51" box="[813,934,475,500]" italics="true" pageId="11" pageNumber="206">N. masneri</emphasis>
|
||
</taxonomicName>
|
||
as uniformly convex, without distinct notauli except for a very short furrow anteriorly at the level of the lateral margin of the pronotum mesal to the spiracle visible in lateral view (
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FEE7FDEFFE3BFD98" author="Gibson, G. A. P." box="[329,478,547,573]" pageId="11" pageNumber="206" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
|
||
, fig. 60). He also stated that minute air bubbles reduced clarity of observation of the inclusion and that the mostly slightly lanceolate- or spatulate-appearing setae, as seen in dorsal view, likely resulted from the setae being surrounded by a thin layer of air because in lateral view the pronotal and mesoscutal setae appear more normal, hair-like. With the discovery of the new species of
|
||
<taxonomicName id="4C6EB9E9A60CFFB9FC2AFD43FBCAFD0C" authorityName="Gibson" authorityYear="2009" box="[900,1071,655,681]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="206" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91A1E78A60CFFB9FC2AFD43FBCAFD0C" box="[900,1071,655,681]" italics="true" pageId="11" pageNumber="206">Neanaperiallus</emphasis>
|
||
</taxonomicName>
|
||
, which clearly shows complete, furrow-like notauli (
|
||
<figureCitation id="1355DEEFA60CFFB9FED9FD7FFDF6FD68" box="[375,531,691,717]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C & E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D & H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="11" pageNumber="206">Figs 4A, E‒G</figureCitation>
|
||
) similar to those of
|
||
<taxonomicName id="4C6EB9E9A60CFFB9FD5EFD7FFCB8FD69" authorityName="Gibson" authorityYear="2008" box="[752,861,691,716]" class="Insecta" family="Tanaostigmatidae" genus="Leptoomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="janzeni">
|
||
<emphasis id="B91A1E78A60CFFB9FD5EFD7FFCB8FD69" box="[752,861,691,716]" italics="true" pageId="11" pageNumber="206">L. janzeni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1355DEEFA60CFFB9FCC2FD7FFC21FD68" box="[876,964,691,717]" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G & H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="11" pageNumber="206">Fig. 2H</figureCitation>
|
||
), the apparent absence of complete notauli for
|
||
<taxonomicName id="4C6EB9E9A60CFFB9FF6EFD1BFEDFFD55" authorityName="Gibson" authorityYear="2009" box="[192,314,727,752]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="masneri">
|
||
<emphasis id="B91A1E78A60CFFB9FF6EFD1BFEDFFD55" box="[192,314,727,752]" italics="true" pageId="11" pageNumber="206">N. masneri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1355DEEFA60CFFB9FEE5FD1BFE40FD54" box="[331,421,727,753]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C & E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D & H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="11" pageNumber="206">Fig. 4H</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FE1CFD1BFDA1FD54" author="Gibson, G. A. P." box="[434,580,727,753]" pageId="11" pageNumber="206" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
|
||
, fig. 59) likely reflects an artefact of the furrows not being visible except in lateral view immediately behind the pronotum. Different views and states of preservation of different individuals of
|
||
<taxonomicName id="4C6EB9E9A60CFFB9FF39FCD3FEE1FC9D" authorityName="Gibson" authorityYear="2008" box="[151,260,799,824]" class="Insecta" family="Tanaostigmatidae" genus="Leptoomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="janzeni">
|
||
<emphasis id="B91A1E78A60CFFB9FF39FCD3FEE1FC9D" box="[151,260,799,824]" italics="true" pageId="11" pageNumber="206">L. janzeni</emphasis>
|
||
</taxonomicName>
|
||
imaged by
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FE2BFCD3FDC3FC9C" author="Gibson, G. A. P." box="[389,550,799,825]" pageId="11" pageNumber="206" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson (2008)</bibRefCitation>
|
||
also show different forms and relative visibility of the notaular furrows (
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FAE4FCD3FF34FCF8" author="Gibson, G. A. P." pageId="11" pageNumber="206" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
,
|
||
<emphasis id="B91A1E78A60CFFB9FF71FC8FFF16FCF8" box="[223,243,835,861]" italics="true" pageId="11" pageNumber="206">cf</emphasis>
|
||
. figs 4, 6‒8), the furrows being least visible in direct dorsal view (
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FC5FFC8FFB63FCF8" author="Gibson, G. A. P." box="[1009,1158,835,861]" pageId="11" pageNumber="206" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
, figs 4, 8). We therefore interpret complete notaular furrows as a feature of
|
||
<taxonomicName id="4C6EB9E9A60CFFB9FD65FCABFCA1FC25" authorityName="Gibson" authorityYear="2009" box="[715,836,871,896]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="masneri">
|
||
<emphasis id="B91A1E78A60CFFB9FD65FCABFCA1FC25" box="[715,836,871,896]" italics="true" pageId="11" pageNumber="206">N. masneri</emphasis>
|
||
</taxonomicName>
|
||
and describe this feature for the family.
|
||
</paragraph>
|
||
<paragraph id="8BD1C26AA60CFFB9FF69FC47FEFFFB05" blockId="11.[151,1437,150,1184]" pageId="11" pageNumber="206">
|
||
Both available inclusions of
|
||
<taxonomicName id="4C6EB9E9A60CFFB9FDA1FC47FD59FC00" authorityName="Gibson" authorityYear="2009" box="[527,700,907,933]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="206" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91A1E78A60CFFB9FDA1FC47FD59FC00" box="[527,700,907,933]" italics="true" pageId="11" pageNumber="206">Neanaperiallus</emphasis>
|
||
</taxonomicName>
|
||
have an unflexed mesonotum in which the axillae are separated medially (
|
||
<figureCitation id="1355DEEFA60CFFB9FEA4FC63FE6FFC6C" box="[266,394,943,969]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C & E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D & H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="11" pageNumber="206">Figs 4E‒H</figureCitation>
|
||
). Because of this, it is unknown whether the basally truncate mesoscutellum separating the inner angles of the axillae results from the posterior margin of the mesoscutum overlapping the axillae medially, as for
|
||
<taxonomicName id="4C6EB9E9A60CFFB9FF6EFC3BFED5FBB5" authorityName="Gibson" authorityYear="2008" box="[192,304,1015,1040]" class="Insecta" family="Tanaostigmatidae" genus="Leptoomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="janzeni">
|
||
<emphasis id="B91A1E78A60CFFB9FF6EFC3BFED5FBB5" box="[192,304,1015,1040]" italics="true" pageId="11" pageNumber="206">L. janzeni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFFFBF9BA60CFFB9FE91FC3BFE30FBB4" author="Gibson, G. A. P." box="[319,469,1015,1041]" pageId="11" pageNumber="206" pagination="1 - 26" refId="ref17197" refString="Gibson, G. A. P. (2008) Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae. Zootaxa, 1730 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1730.1.1" type="journal article" year="2008">Gibson 2008</bibRefCitation>
|
||
,
|
||
<emphasis id="B91A1E78A60CFFB9FE4FFC3BFE10FBB4" box="[481,501,1015,1041]" italics="true" pageId="11" pageNumber="206">cf</emphasis>
|
||
. figs 7, 8), or whether the mesonotum articulates, hinge-like, along the transscutal articulation and the inner angles of the axillae are separated by an anteriorly truncate mesoscutellum. Consequently, our family description includes description of relative mesoscutellar-axillar structure when the mesonotum is not flexed, but it remains uncertain whether the mesonotal articulatory structure in
|
||
<taxonomicName id="4C6EB9E9A60CFFB9FB88FBAFFB36FBD8" authorityName="Gibson" authorityYear="2009" box="[1062,1235,1123,1149]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="206" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91A1E78A60CFFB9FB88FBAFFB36FBD8" box="[1062,1235,1123,1149]" italics="true" pageId="11" pageNumber="206">Neanaperiallus</emphasis>
|
||
</taxonomicName>
|
||
is the same as in
|
||
<emphasis id="B91A1E78A60CFFB9FF39FB4BFEFFFB05" box="[151,282,1159,1184]" italics="true" pageId="11" pageNumber="206">
|
||
<taxonomicName id="4C6EB9E9A60CFFB9FF39FB4BFEF3FB05" authorityName="Gibson" authorityYear="2008" box="[151,278,1159,1184]" class="Insecta" family="Tanaostigmatidae" genus="Leptoomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="206" phylum="Arthropoda" rank="genus">Leptoomus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |