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<document id="7970A9B0E1063B080FA6900496624D0B" ID-DOI="10.11646/zootaxa.5318.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8162469" ID-ZooBank="CF8E0B91-9AF7-4075-963D-9BC977B41852" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1689748652228" checkinUser="plazi" docAuthor="Gibson, Gary A. P. &amp; Fusu, Lucian" docDate="2023" docId="03C7737CA609FFBDFF39FC73FC31FEC0" docLanguage="en" docName="zootaxa.5318.2.2.pdf" docOrigin="Zootaxa 5318 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5318.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neanaperiallus Gibson 2009" docType="treatment" docVersion="1" lastPageNumber="209" masterDocId="FFFE0B04A607FFB2FFAEFFCCFFE5FFA5" masterDocTitle="Leptoomidae, a new family of Eocene fossil Chalcidoidea (Hymenoptera), and family classification of Eocene fossil genera originally described in Neanastatinae (Eupelmidae)" masterLastPageNumber="216" masterPageNumber="195" pageNumber="209" updateTime="1689856743257" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="A5E03CA839CCDED1AD5B604D898838D6">Leptoomidae, a new family of Eocene fossil Chalcidoidea (Hymenoptera), and family classification of Eocene fossil genera originally described in Neanastatinae (Eupelmidae)</mods:title>
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<mods:name id="27AA7CDD4699434C3888A7F9EF74D7B1" type="personal">
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<mods:namePart id="1AA32E2802B8C4F3363F88123033CE33">Gibson, Gary A. P.</mods:namePart>
<mods:affiliation id="C991D96AABB9CAC47079340874C31838">Honorary Research Associate, Agriculture and Agri-Food Canada, Canadian National Collection of Insects, Arachnids and Nematodes, K. W. Neatby Bldg., 960 Carling Avenue, Ottawa, Ontario, CANADA, K 1 A 0 C 6.</mods:affiliation>
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</mods:role>
<mods:namePart id="2B62EEFB730565B8DC51CA90FE2A33B4">Fusu, Lucian</mods:namePart>
<mods:affiliation id="30986B003A6892757D4007D01CBB2F9E">Al. I. Cuza University, Faculty of Biology, Bd. Carol I, Nr 11, 700506 Iasi, ROMANIA.</mods:affiliation>
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<mods:title id="458067ADB45F3CA001588B0A2542752B">Zootaxa</mods:title>
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<mods:date id="EB5ABB946ADFB442FA8EF1686229C316">2023</mods:date>
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<mods:number id="6F539C973D5E9FA87FA5CD7B6E8E6AE6">2023-07-19</mods:number>
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<mods:number id="5EE446B9CDFA08433C67FB92DE5A7470">5318</mods:number>
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<mods:url id="33DB88F19DA373098FB8982C7F035ADB">http://dx.doi.org/10.11646/zootaxa.5318.2.2</mods:url>
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<mods:classification id="669F47B7663835672AB87CB6F8A29240">journal article</mods:classification>
<mods:identifier id="BA9D9CFCA1C8FCEBE21CB2FF52591406" type="DOI">10.11646/zootaxa.5318.2.2</mods:identifier>
<mods:identifier id="D85464BD0D3D7023D46FEFB464397988" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="C37491E1A609FFBCFF39FC73FE29FC79" box="[151,460,959,988]" pageId="14" pageNumber="209" type="nomenclature">
<paragraph id="8BD1C26AA609FFBCFF39FC73FE29FC79" blockId="14.[151,460,959,1024]" box="[151,460,959,988]" pageId="14" pageNumber="209">
<taxonomicName id="4C6EB9E9A609FFBCFF39FC73FE29FC79" authority="Gibson" authorityName="Gibson" authorityYear="2009" box="[151,460,959,988]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B91A1E78A609FFBCFF39FC73FE29FC79" bold="true" box="[151,460,959,988]" pageId="14" pageNumber="209">
<emphasis id="B91A1E78A609FFBCFF39FC73FE87FC79" bold="true" box="[151,354,959,988]" italics="true" pageId="14" pageNumber="209">Neanaperiallus</emphasis>
Gibson
</emphasis>
</taxonomicName>
</paragraph>
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<subSubSection id="C37491E1A609FFBCFF39FC29FE8FFBA5" box="[151,362,997,1024]" pageId="14" pageNumber="209" type="description">
<paragraph id="8BD1C26AA609FFBCFF39FC29FE8FFBA5" blockId="14.[151,460,959,1024]" box="[151,362,997,1024]" pageId="14" pageNumber="209">
(
<figureCitation id="1355DEEFA609FFBCFF31FC29FF19FC5A" box="[159,252,997,1023]" captionStart="FIGURE 1" captionStartId="10.[152,255,1835,1859]" captionTargetBox="[152,1436,180,1809]" captionTargetId="figure-25@10.[151,1436,179,1809]" captionTargetPageId="10" captionText="FIGURE 1. Lateral view of mesosoma: A, Neanaperiallus masneri; B, Leptoomus janzeni; C, Tanaostigmodes sp.; D, Cynipencyrtus ishii; E, Eopelma gibsoni; F, Savzdargia sp.; G, Neanastatus sp.; H, Lambdobregma sp.; I, Brevivulva electroma. Insets in H and I are dorsal views of apex of mesoscutellum to propodeum; green arrows on images indicate the intersection of base of tegula with dorsal margin of prepectus, white arrows indicate posterior margin of prepectus; see ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162471" httpUri="https://zenodo.org/record/8162471/files/figure.png" pageId="14" pageNumber="209">Figs 1A</figureCitation>
,
<figureCitation id="1355DEEFA609FFBCFEA6FC29FECEFC5A" box="[264,299,997,1023]" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G &amp; H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="14" pageNumber="209">2B</figureCitation>
,
<figureCitation id="1355DEEFA609FFBCFE99FC29FEA2FC5A" box="[311,327,997,1023]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C &amp; E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D &amp; H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="14" pageNumber="209">4</figureCitation>
,
<figureCitation id="1355DEEFA609FFBCFEFDFC29FE84FBA5" box="[339,353,997,1024]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37491E1A609FFBCFF39FBE8FDD5FB98" box="[151,560,1060,1085]" pageId="14" pageNumber="209" type="reference_group">
<paragraph id="8BD1C26AA609FFBCFF39FBE8FDD5FB98" blockId="14.[151,560,1060,1085]" box="[151,560,1060,1085]" pageId="14" pageNumber="209">
<treatmentCitationGroup id="AB7EE544A609FFBCFF39FBE8FDD5FB98" box="[151,560,1060,1085]" pageId="14" pageNumber="209">
<treatmentCitation id="0ACFE47BA609FFBCFF39FBE8FDCEFB98" ID-DOI="http://doi.org/10.5281/zenodo.3791301" ID-Zenodo-Dep="3791301" LSID="urn:lsid:zoobank.org:act:6654E412-1382-46E9-BDE1-3BDA2469B95C" author="Gibson, G. A. P." box="[151,555,1060,1085]" httpUri="http://treatment.plazi.org/id/A02887B6FFB9FFE39BDB14C3FD87FEA3" page="203" pageId="14" pageNumber="209" year="2009">
<taxonomicName id="4C6EB9E9A609FFBCFF39FBE8FDCEFB98" authority="Gibson, 2009: 200 - 203" authorityName="Gibson" authorityPageNumber="200 - 203" authorityYear="2009" box="[151,555,1060,1085]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B91A1E78A609FFBCFF39FBE8FED6FB99" box="[151,307,1060,1084]" italics="true" pageId="14" pageNumber="209">Neanaperiallus</emphasis>
<bibRefCitation id="EFFFBF9BA609FFBCFE94FBE8FDCEFB98" author="Gibson, G. A. P." box="[314,555,1060,1085]" pageId="14" pageNumber="209" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson, 2009: 200‒203</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
</subSubSection>
<subSubSection id="C37491E1A609FFBCFF39FBA9FB5DFBDA" box="[151,1208,1125,1152]" pageId="14" pageNumber="209" type="multiple">
<paragraph id="8BD1C26AA609FFBCFF39FBA9FB5DFBDA" blockId="14.[151,1437,1125,2016]" box="[151,1208,1125,1152]" pageId="14" pageNumber="209">
<emphasis id="B91A1E78A609FFBCFF39FBA9FEBFFBDA" bold="true" box="[151,346,1125,1151]" pageId="14" pageNumber="209">Included species</emphasis>
.
<taxonomicName id="4C6EB9E9A609FFBCFEC6FBA9FD20FBDA" authority="Gibson" authorityName="Gibson" box="[360,709,1125,1151]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="janzeni">
<emphasis id="B91A1E78A609FFBCFEC6FBA9FD88FBDA" box="[360,621,1125,1151]" italics="true" pageId="14" pageNumber="209">Neanaperiallus janzeni</emphasis>
Gibson
</taxonomicName>
and
<taxonomicName id="4C6EB9E9A609FFBCFD55FBA9FBB2FB25" authority="Fusu" authorityName="Gibson &amp; Fusu" authorityYear="2023" box="[763,1111,1125,1152]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="defunctus" status="sp. nov.">
<emphasis id="B91A1E78A609FFBCFD55FBA9FBFFFBDA" box="[763,1050,1125,1151]" italics="true" pageId="14" pageNumber="209">Neanaperiallus defunctus</emphasis>
Fusu
</taxonomicName>
<emphasis id="B91A1E78A609FFBCFBF1FBA9FB5DFBDA" bold="true" box="[1119,1208,1125,1151]" pageId="14" pageNumber="209">
<taxonomicNameLabel id="A229A303A609FFBCFBF1FBA9FB5DFBDA" box="[1119,1208,1125,1151]" pageId="14" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37491E1A609FFBCFF69FB45FCDBF819" pageId="14" pageNumber="209" type="diagnosis">
<paragraph id="8BD1C26AA609FFBCFF69FB45FCDBF819" blockId="14.[151,1437,1125,2016]" pageId="14" pageNumber="209">
<emphasis id="B91A1E78A609FFBCFF69FB45FEDFFB06" bold="true" box="[199,314,1161,1187]" pageId="14" pageNumber="209">Diagnosis</emphasis>
. Head and mesosoma non-metallic, brown. Head with inner margins of eyes apparently not divergent ventrally or at least not conspicuously divergent (not visible in
<taxonomicName id="4C6EB9E9A609FFBCFCC8FB62FC3CFB62" box="[870,985,1198,1223]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="janzeni">
<emphasis id="B91A1E78A609FFBCFCC8FB62FC3CFB62" box="[870,985,1198,1223]" italics="true" pageId="14" pageNumber="209">N. janzeni</emphasis>
</taxonomicName>
and direct frontal view not possible for
<taxonomicName id="4C6EB9E9A609FFBCFF39FB1EFEC1FB4E" authorityName="Gibson &amp; Fusu" authorityYear="2023" box="[151,292,1233,1259]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="defunctus">
<emphasis id="B91A1E78A609FFBCFF39FB1EFEC1FB4E" box="[151,292,1233,1259]" italics="true" pageId="14" pageNumber="209">N. defunctus</emphasis>
</taxonomicName>
,
<figureCitation id="1355DEEFA609FFBCFE9DFB1DFE6CFB49" box="[307,393,1233,1260]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">Fig. 5E</figureCitation>
); clypeal area concave and labrum perpendicular to clypeus (
<emphasis id="B91A1E78A609FFBCFB93FB1EFB82FB4E" box="[1085,1127,1234,1259]" italics="true" pageId="14" pageNumber="209">i. e.</emphasis>
not visible in direct frontal view of clypeus,
<emphasis id="B91A1E78A609FFBCFEF7FB39FE88FAAA" box="[345,365,1269,1295]" italics="true" pageId="14" pageNumber="209">cf</emphasis>
.
<figureCitation id="1355DEEFA609FFBCFED3FB39FE35FAB5" box="[381,464,1269,1296]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">Fig. 5E</figureCitation>
and inset in 5E); occipital carina
<emphasis id="B91A1E78A609FFBCFCFEFB39FC81FAAA" bold="true" box="[848,868,1269,1295]" pageId="14" pageNumber="209">∩</emphasis>
-shaped (
<figureCitation id="1355DEEFA609FFBCFC62FB39FBC4FAB5" box="[972,1057,1269,1296]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">Fig. 5F</figureCitation>
;
<bibRefCitation id="EFFFBF9BA609FFBCFB81FB39FB27FAAA" author="Gibson, G. A. P." box="[1071,1218,1269,1295]" pageId="14" pageNumber="209" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
, fig. 59). Mandible not visible in
<taxonomicName id="4C6EB9E9A609FFBCFE81FAD6FE45FA96" box="[303,416,1306,1331]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="janzeni">
<emphasis id="B91A1E78A609FFBCFE81FAD6FE45FA96" box="[303,416,1306,1331]" italics="true" pageId="14" pageNumber="209">N. janzeni</emphasis>
</taxonomicName>
but apparently tridentate in
<taxonomicName id="4C6EB9E9A609FFBCFD74FAD6FC83FA96" authorityName="Gibson &amp; Fusu" authorityYear="2023" box="[730,870,1305,1331]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="defunctus">
<emphasis id="B91A1E78A609FFBCFD74FAD6FC83FA96" box="[730,870,1305,1331]" italics="true" pageId="14" pageNumber="209">N. defunctus</emphasis>
</taxonomicName>
. Antenna with lower margin of torulus in line with or slightly ventral to lower margin of eyes; 13-segmented (1:1:8:3) with 8 slightly transverse to slightly elongate funiculars and with mps on fu
<subScript id="17EAC02FA609FFBCFE49FAB8FE0AFA27" attach="right" box="[487,495,1396,1410]" fontSize="6" pageId="14" pageNumber="209">3</subScript>
‒fu
<subScript id="17EAC02FA609FFBCFDBAFAB8FDF9FA27" attach="left" box="[532,540,1396,1410]" fontSize="6" pageId="14" pageNumber="209">8</subScript>
(
<figureCitation id="1355DEEFA609FFBCFD85FAADFD23FAD9" box="[555,710,1377,1404]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C &amp; E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D &amp; H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="14" pageNumber="209">Figs 4B, C, F</figureCitation>
;
<bibRefCitation id="EFFFBF9BA609FFBCFD7CFAADFC80FADE" author="Gibson, G. A. P." box="[722,869,1377,1403]" pageId="14" pageNumber="209" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
, fig. 58); clava elongate-ovoid, more than twice as long as wide, and apical clavomere with minute micropilose sensory region apically, appearing as rounded terminal button (
<figureCitation id="1355DEEFA609FFBCFF41FA65FEAFFA66" box="[239,330,1449,1475]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C &amp; E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D &amp; H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="14" pageNumber="209">Fig. 4C</figureCitation>
: tb). Mesoscutellar-axillar complex with axillae equilateral to somewhat elongate-triangular, with their inner angles slightly separated and mesoscutellum truncate basally when mesonotum not flexed (
<figureCitation id="1355DEEFA609FFBCFA99FA01FF49F9AE" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C &amp; E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D &amp; H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="14" pageNumber="209">Figs 4G, H</figureCitation>
) (flexed condition unknown); mesoscutellum with sculpture along apical margin aligned, differentiating a thin carina (
<figureCitation id="1355DEEFA609FFBCFF41F9D9FE61F98A" box="[239,388,1557,1583]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C &amp; E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D &amp; H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="14" pageNumber="209">Figs 4E, 4H</figureCitation>
,
<figureCitation id="1355DEEFA609FFBCFE3AF9D9FE5FF995" box="[404,442,1557,1584]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">5H</figureCitation>
: smc) separating lunate, down-sloped region apically. Metanotum with dorsellum an oblique band projecting over apex of mesoscutellum (
<figureCitation id="1355DEEFA609FFBCFCA6F9F5FC86F9F1" box="[776,867,1593,1620]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">Fig. 5H</figureCitation>
) but the two sclerites separated by a narrow gap (
<figureCitation id="1355DEEFA609FFBCFF31F991FF18F9DD" box="[159,253,1629,1656]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">Fig. 5H</figureCitation>
: double arrow in inset). Prepectus flat (
<figureCitation id="1355DEEFA609FFBCFD67F991FCC7F9D2" box="[713,802,1629,1655]" captionStart="FIGURE 4" captionStartId="17.[152,255,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-19@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 4. A‒C &amp; E‒G, Neanaperiallus defunctus sp. nov., holotype ♀: A, habitus with amber piece containing the fossil as an inset; B, head and outer view of antenna; C, antenna in dorsal view; E‒G, head and mesosoma with arrow in G indicating the posteriorly dilated part of the propodeal carina. D &amp; H, Neanaperiallus masneri: D, fore and hind wings; H, mesosoma in dorsal view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162478" httpUri="https://zenodo.org/record/8162478/files/figure.png" pageId="14" pageNumber="209">Fig. 4F</figureCitation>
: pre). Acropleuron enlarged posteriorly to metapleuron and posteroventrally to dorsolateral level of mesocoxa, with crenulate acropleural sulcus directed obliquely toward posterodorsal angle of prepectus over most of length (
<figureCitation id="1355DEEFA609FFBCFD57F969FCB4F91A" box="[761,849,1701,1727]" captionStart="FIGURE 1" captionStartId="10.[152,255,1835,1859]" captionTargetBox="[152,1436,180,1809]" captionTargetId="figure-25@10.[151,1436,179,1809]" captionTargetPageId="10" captionText="FIGURE 1. Lateral view of mesosoma: A, Neanaperiallus masneri; B, Leptoomus janzeni; C, Tanaostigmodes sp.; D, Cynipencyrtus ishii; E, Eopelma gibsoni; F, Savzdargia sp.; G, Neanastatus sp.; H, Lambdobregma sp.; I, Brevivulva electroma. Insets in H and I are dorsal views of apex of mesoscutellum to propodeum; green arrows on images indicate the intersection of base of tegula with dorsal margin of prepectus, white arrows indicate posterior margin of prepectus; see ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162471" httpUri="https://zenodo.org/record/8162471/files/figure.png" pageId="14" pageNumber="209">Fig. 1A</figureCitation>
) or only over about anterior one-third (
<figureCitation id="1355DEEFA609FFBCFABEF969FA8DF965" box="[1296,1384,1701,1728]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">Fig. 5A</figureCitation>
) but differentiating acropleuron from comparatively large mesepisternal region ventral to acropleural suture (
<figureCitation id="1355DEEFA609FFBCFA97F905FA72F946" box="[1337,1431,1737,1763]" captionStart="FIGURE 1" captionStartId="10.[152,255,1835,1859]" captionTargetBox="[152,1436,180,1809]" captionTargetId="figure-25@10.[151,1436,179,1809]" captionTargetPageId="10" captionText="FIGURE 1. Lateral view of mesosoma: A, Neanaperiallus masneri; B, Leptoomus janzeni; C, Tanaostigmodes sp.; D, Cynipencyrtus ishii; E, Eopelma gibsoni; F, Savzdargia sp.; G, Neanastatus sp.; H, Lambdobregma sp.; I, Brevivulva electroma. Insets in H and I are dorsal views of apex of mesoscutellum to propodeum; green arrows on images indicate the intersection of base of tegula with dorsal margin of prepectus, white arrows indicate posterior margin of prepectus; see ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162471" httpUri="https://zenodo.org/record/8162471/files/figure.png" pageId="14" pageNumber="209">Figs 1A</figureCitation>
,
<figureCitation id="1355DEEFA609FFBCFF39F921FF58F8AD" box="[151,189,1773,1800]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">5A</figureCitation>
). Mesopectus in ventral view with posterior margin abutting base of mesocoxa (
<figureCitation id="1355DEEFA609FFBCFB95F921FB75F8A2" box="[1083,1168,1773,1799]" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G &amp; H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="14" pageNumber="209">Fig. 2B</figureCitation>
) though mesocoxa with triangular basolateral cavity opposite angle formed between acropleuron and mesopectus (
<figureCitation id="1355DEEFA609FFBCFB32F8DDFB13F88E" box="[1180,1270,1809,1835]" captionStart="FIGURE 2" captionStartId="12.[152,255,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-25@12.[151,1436,181,1879]" captionTargetPageId="12" captionText="FIGURE 2. A, Cynipencyrtus ishii, mesosoma, ventrolateral view. B, Neanaperiallus masneri, mesosoma, ventral view. C‒E, Leptoomus janzeni: C, mesosoma, ventrolateral view; D, posterior of mesopectus and mesocoxa, posterior view; E, mesopectus and mesocoxae, posteroventral view. F, Tanaostigmodes sp., mesosoma, ventral view. G &amp; H, Leptoomus janzeni: G, head and antenna, frontal view; H, head and mesosoma, dorsolateral view. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162474" httpUri="https://zenodo.org/record/8162474/files/figure.png" pageId="14" pageNumber="209">Fig. 2B</figureCitation>
;
<bibRefCitation id="EFFFBF9BA609FFBCFAAAF8DDFA7DF88E" author="Gibson, G. A. P." box="[1284,1432,1809,1835]" pageId="14" pageNumber="209" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
, fig. 61). Fore wing disc with bare band beyond basal fold separated from parastigma by region of setae (
<figureCitation id="1355DEEFA609FFBCFA97F8F9FF4BF8D1" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">Figs 5G, H</figureCitation>
;
<bibRefCitation id="EFFFBF9BA609FFBCFF14F895FEA8F8D6" author="Gibson, G. A. P." box="[186,333,1881,1907]" pageId="14" pageNumber="209" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson 2009</bibRefCitation>
, fig. 62), parastigma not separated from marginal vein by hyaline break though apparently slightly less pigmented distally (
<figureCitation id="1355DEEFA609FFBCFE0BF8B1FDC0F832" box="[421,549,1917,1944]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">Figs 5G, H</figureCitation>
). Protibia apparently without dorsal spicules but with dorsoapical spicules (
<figureCitation id="1355DEEFA609FFBCFAC1F8B1FF59F819" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">Fig. 5A</figureCitation>
, inset); mesotibia with spine-like pegs apically (
<figureCitation id="1355DEEFA609FFBCFD77F86DFCD7F819" box="[729,818,1953,1980]" captionStart="FIGURE 5" captionStartId="18.[152,255,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-25@18.[151,1436,181,1558]" captionTargetPageId="18" captionText="FIGURE 5. Neanaperiallus defunctus sp. nov., holotype ♀: A, head and mesosoma in ventral view, protibia as an inset with arrow indicating dorsoapical pegs; B, apex of mesotibia and mesotarsus in dorsal (posterior) view; C, apex of mesotibia and mesotarsus in ventral (anterior) view; D, distal part of gaster in ventral view; E, head in ventral view with the inset showing the clypeus, labrum and mandibles and arrows indicating the mandibular teeth; F, head in posterior view; G, fore wing; H, fore wing base and, as inset, mesoscutellar-axillar complex, apex of mesoscutellum and dorsellum with double-headed arrow within gap between the two sclerites and single arrow indicating the subvertical mesoscutellar apex. See ‘Material and methods’ for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.8162480" httpUri="https://zenodo.org/record/8162480/files/figure.png" pageId="14" pageNumber="209">Fig. 5C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37491E1A609FFBDFF69F809FC31FEC0" lastPageId="15" lastPageNumber="210" pageId="14" pageNumber="209" type="discussion">
<paragraph id="8BD1C26AA609FFBDFF69F809FC31FEC0" blockId="14.[151,1437,1125,2016]" lastBlockId="15.[151,1436,151,357]" lastPageId="15" lastPageNumber="210" pageId="14" pageNumber="209">
<emphasis id="B91A1E78A609FFBCFF69F809FED1F87A" bold="true" box="[199,308,1989,2015]" pageId="14" pageNumber="209">Remarks</emphasis>
. The extended generic diagnosis given above for
<taxonomicName id="4C6EB9E9A609FFBCFCD9F809FBC1F87A" authorityName="Gibson" authorityYear="2009" box="[887,1060,1989,2015]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B91A1E78A609FFBCFCD9F809FBC1F87A" box="[887,1060,1989,2015]" italics="true" pageId="14" pageNumber="209">Neanaperiallus</emphasis>
</taxonomicName>
is to supplement the description in
<bibRefCitation id="EFFFBF9BA608FFBDFF1BFF5BFEBFFF14" author="Gibson, G. A. P." box="[181,346,151,177]" pageId="15" pageNumber="210" pagination="175 - 214" refId="ref17269" refString="Gibson, G. A. P. (2009) Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa. ZooKeys, 20, 175 - 214. https: // doi. org / 10.3897 / zookeys. 20.161" type="journal article" year="2009">Gibson (2009)</bibRefCitation>
after the discovery of a second fossil that represents a second species in the genus, and to have a comparative description with
<taxonomicName id="4C6EB9E9A608FFBDFE53FF77FD9DFF71" authorityName="Gibson" authorityYear="2008" box="[509,632,187,212]" class="Insecta" family="Tanaostigmatidae" genus="Leptoomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B91A1E78A608FFBDFE53FF77FD9DFF71" box="[509,632,187,212]" italics="true" pageId="15" pageNumber="210">Leptoomus</emphasis>
</taxonomicName>
. As noted above, the state of preservation and the inability to examine the two available inclusions equally from all angles prevents directly comparable, accurate description of all parts of the two species so the generic diagnosis and/or species descriptions may be inaccurate in some respects for one or both species. Additional inclusions are necessary to further clarify apparent similarities and differences between
<taxonomicName id="4C6EB9E9A608FFBDFF39FE87FEA1FEC0" authorityName="Gibson" authorityYear="2009" box="[151,324,331,357]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B91A1E78A608FFBDFF39FE87FEA1FEC0" box="[151,324,331,357]" italics="true" pageId="15" pageNumber="210">Neanaperiallus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6EB9E9A608FFBDFED4FE87FE10FEC1" authorityName="Gibson" authorityYear="2008" box="[378,501,331,356]" class="Insecta" family="Tanaostigmatidae" genus="Leptoomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B91A1E78A608FFBDFED4FE87FE10FEC1" box="[378,501,331,356]" italics="true" pageId="15" pageNumber="210">Leptoomus</emphasis>
</taxonomicName>
and between
<taxonomicName id="4C6EB9E9A608FFBDFD3EFE87FCF9FEC0" authorityName="Gibson &amp; Fusu" authorityYear="2023" box="[656,796,331,357]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="species" species="defunctus">
<emphasis id="B91A1E78A608FFBDFD3EFE87FCF9FEC0" box="[656,796,331,357]" italics="true" pageId="15" pageNumber="210">N. defunctus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6EB9E9A608FFBDFCFCFE87FC28FEC1" authorityName="Gibson" authorityYear="2009" box="[850,973,331,356]" class="Insecta" family="Eupelmidae" genus="Neanaperiallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="species" species="masneri">
<emphasis id="B91A1E78A608FFBDFCFCFE87FC28FEC1" box="[850,973,331,356]" italics="true" pageId="15" pageNumber="210">N. masneri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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