709 lines
112 KiB
XML
709 lines
112 KiB
XML
<document id="076455B76D3141BFDB49A3622095865E" ID-CLB-Dataset="6499" ID-DOI="10.11646/zootaxa.4965.3.8" ID-GBIF-Dataset="01b94d4b-9006-43fb-bc7c-d953eb189ce8" ID-ISSN="1175-5326" ID-Zenodo-Dep="4736172" ID-ZooBank="A42D0F3D-3145-4540-82A4-E936ADBDDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620111888585" checkinUser="plazi" docAuthor="Tiunova, Tatiana M., Semenchenko, Alexander A. & Tong, Xiaoli" docDate="2021" docId="03B91F39FFA8FF8CE1D1A243FD85BC91" docLanguage="en" docName="zootaxa.4965.3.8.pdf" docOrigin="Zootaxa 4965 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Baetis majus Tiunova & Semenchenko & Tong 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="552" masterDocId="FF806741FFAAFF87E146A32CFF99BD40" masterDocTitle="Baetis majus sp. nov., new species of mayfly (Ephemeroptera: Baetidae) from Far East of Russia" masterLastPageNumber="557" masterPageNumber="541" pageNumber="543" updateTime="1698938243106" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="2FCFFE4439744B72C2AB51993B64A2CA">
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<mods:title id="C5405FBB4CEA791300AB3038B893BEDB">Baetis majus sp. nov., new species of mayfly (Ephemeroptera: Baetidae) from Far East of Russia</mods:title>
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<mods:name id="8C8B931979CA1A92AD4CF3B152BB48CC" type="personal">
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<mods:roleTerm id="729CF690CBA8E632E6FACDCDAAB0F500">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F9E224BBABFA7E186458FD0BC2159B6B">Tiunova, Tatiana M.</mods:namePart>
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<mods:affiliation id="AFD05C22F307D600F2ABE2E005535E6E">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok, 690022, Russia.</mods:affiliation>
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</mods:name>
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<mods:name id="E18ECE75C9C8D40F29C67F4A5DF6E83F" type="personal">
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<mods:roleTerm id="7A4AEF362E51B46EC120DA039943F619">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="930852C566BACF14E9F96A810A237D1A">Semenchenko, Alexander A.</mods:namePart>
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<mods:affiliation id="C0ED75C0DE2E8ACEFB1212E487393ED5">Laboratory of Ecology and Evolutionary Biology of Aquatic Organisms, School of Natural Sciences, Far Eastern Federal University, Suhanova St. 8, 690950 Vladivostok, Russia.</mods:affiliation>
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<mods:roleTerm id="9310AA32781AE35A2F7B8067DDCDD791">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A79A8E72A9056E95F30D83A0AABAA7ED">Tong, Xiaoli</mods:namePart>
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<mods:nameIdentifier id="699C23717EA48D14C5620D6B145D8B42" type="ORCID">0000-0003-1731-229X</mods:nameIdentifier>
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<mods:affiliation id="6F9D38D0313AEFFDFCC5C16F490FCABF">Department of Entomology, College of Plant Protection, South China Agricultural University, Guangzhou 510642, Guangdong, China. xtong @ scau. edu. cn; https: // orcid. org / 0000 - 0003 - 1731 - 229 X</mods:affiliation>
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<mods:nameIdentifier id="ED9A39DBC3A352ACB58817C7AF4DE872" type="email">xtong@scau.edu.cn</mods:nameIdentifier>
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<mods:title id="13F53974B7EBF00B39FA5B5CAC719E2A">Zootaxa</mods:title>
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<mods:part id="B5478A9A51D192916B8ABCD53139B001">
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<mods:date id="3C0EF3E9610DB6C4B6BA917D7C112706">2021</mods:date>
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<mods:number id="2AD825F5E381236AB896A086135267E9">2021-04-29</mods:number>
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<mods:number id="226D80F68431AC793E1AF2AD2DD34580">4965</mods:number>
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<treatment id="03B91F39FFA8FF8CE1D1A243FD85BC91" ID-DOI="http://doi.org/10.5281/zenodo.4790950" ID-GBIF-Taxon="182387813" ID-Zenodo-Dep="4790950" LSID="urn:lsid:plazi:treatment:03B91F39FFA8FF8CE1D1A243FD85BC91" httpUri="http://treatment.plazi.org/id/03B91F39FFA8FF8CE1D1A243FD85BC91" lastPageId="11" lastPageNumber="552" pageId="2" pageNumber="543">
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<subSubSection id="C30AFDA4FFA8FF85E1D1A243FD9FBCC9" box="[151,518,366,393]" pageId="2" pageNumber="543" type="nomenclature">
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<paragraph id="8BAFAE2FFFA8FF85E1D1A243FD9FBCC9" blockId="2.[151,993,366,465]" box="[151,518,366,393]" pageId="2" pageNumber="543">
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<heading id="D0E71943FFA8FF85E1D1A243FD9FBCC9" bold="true" box="[151,518,366,393]" fontSize="11" level="1" pageId="2" pageNumber="543" reason="1">
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<emphasis id="B964723DFFA8FF85E1D1A243FD9FBCC9" bold="true" box="[151,518,366,393]" pageId="2" pageNumber="543">
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<taxonomicName id="4C10D5ACFFA8FF85E1D1A243FE3BBCC8" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[151,418,366,393]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
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<emphasis id="B964723DFFA8FF85E1D1A243FEABBCC9" bold="true" box="[151,306,367,393]" italics="true" pageId="2" pageNumber="543">Baetis majus</emphasis>
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Tiunova
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</taxonomicName>
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<taxonomicNameLabel id="A257CF46FFA8FF85E0EFA243FD9FBCC9" box="[425,518,367,393]" pageId="2" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C30AFDA4FFA8FF85E1D1A2BEFE80BC91" pageId="2" pageNumber="543" type="description">
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<paragraph id="8BAFAE2FFFA8FF85E1D1A2BEFC78BCED" blockId="2.[151,993,366,465]" box="[151,993,402,429]" pageId="2" pageNumber="543">
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<uri id="FF81A22DFFA8FF85E1D1A2BEFC78BCED" box="[151,993,402,429]" pageId="2" pageNumber="543">
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urn:lsid:zoobank.org:pub:
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</uri>
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</paragraph>
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<paragraph id="8BAFAE2FFFA8FF85E1D1A29BFE80BC91" blockId="2.[151,993,366,465]" box="[151,281,439,465]" pageId="2" pageNumber="543">
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(
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<figureCitation id="132BB2AAFFA8FF85E1D9A29BFE89BC91" box="[159,272,439,465]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–4" captionStart-2="FIGURES 5–7" captionStart-3="FIGURES 8–13" captionStart-4="FIGURES 14–20" captionStart-5="FIGURES 21–26" captionStart-6="FIGURES 27–32" captionStart-7="FIGURES 33–38" captionStart-8="FIGURES 39–41" captionStartId-0="3.[151,264,1943,1967]" captionStartId-1="4.[151,264,1968,1992]" captionStartId-2="5.[151,264,1550,1574]" captionStartId-3="6.[151,264,1930,1954]" captionStartId-4="7.[151,264,1931,1955]" captionStartId-5="8.[151,264,1761,1785]" captionStartId-6="9.[151,264,1966,1990]" captionStartId-7="10.[151,264,1218,1242]" captionStartId-8="11.[151,264,1992,2016]" captionTargetBox-0="[252,1335,182,1919]" captionTargetBox-1="[171,1416,189,1933]" captionTargetBox-2="[170,1418,309,1527]" captionTargetBox-3="[151,1436,669,1907]" captionTargetBox-4="[156,1421,190,1907]" captionTargetBox-5="[160,1422,198,1719]" captionTargetBox-6="[151,1436,490,1943]" captionTargetBox-7="[162,1420,194,1185]" captionTargetBox-8="[152,1436,489,1969]" captionTargetId-0="figure-17@3.[252,1335,181,1919]" captionTargetId-1="figure-27@4.[160,1427,181,1944]" captionTargetId-2="figure-92@5.[170,1418,309,1527]" captionTargetId-3="figure-277@6.[151,1436,669,1907]" captionTargetId-4="figure-17@7.[151,1436,181,1907]" captionTargetId-5="figure-27@8.[151,1436,181,1737]" captionTargetId-6="figure-161@9.[151,1436,489,1943]" captionTargetId-7="figure-27@10.[151,1436,181,1193]" captionTargetId-8="figure-163@11.[151,1436,489,1969]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionText-0="FIGURES 1–2. Baetis majus Tiunova sp. nov., 1, male imago, holotype, lateral view; 2, female imago, dorsal view." captionText-1="FIGURES 3–4. Baetis majus Tiunova sp. nov., male imagoes, holotype: 3, hind wing; 4, unistyliger and forceps." captionText-2="FIGURES 5–7. Color pattern of Baetis majus Tiunova sp. nov., larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view." captionText-3="FIGURES 8–13. Baetis majus Tiunova sp. nov., larvae, dorsal view, paratypes: 8, scape; 9, surface of the head; 10, spatulas on head; 11, canines of left and right mandible; 12, glossae and paraglossae; 13, setae on paraglossa. Scale bar: 8, 11–12, 0.01 mm; 9, 0.02 mm; 10, 13, 0.002 mm." captionText-4="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." captionText-5="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." captionText-6="FIGURES 27–32. Baetis majus Tiunova sp. nov., larvae, paratypes: 27, 29, inner margin of fore tibia; 28, inner margin of fore tarsus; 30, tarsal claw; 31, paraproct; 32, spatulas on paraproct. Scale bar: 27–31, 0.01 mm; 32, 0.001 mm." captionText-7="FIGURES 33–38. Baetis majus Tiunova sp. nov., larvae, gills shape, dorsal view, roman numbers belong to the respective gill pairs." captionText-8="FIGURES 39–41. Baetis majus Tiunova sp. nov., egg: 39, general view; 40, structure of chorion; 41, micropyle." figureDoi-0="http://doi.org/10.5281/zenodo.4736174" figureDoi-1="http://doi.org/10.5281/zenodo.4736176" figureDoi-2="http://doi.org/10.5281/zenodo.4736178" figureDoi-3="http://doi.org/10.5281/zenodo.4736180" figureDoi-4="http://doi.org/10.5281/zenodo.4736182" figureDoi-5="http://doi.org/10.5281/zenodo.4736186" figureDoi-6="http://doi.org/10.5281/zenodo.4736188" figureDoi-7="http://doi.org/10.5281/zenodo.4736190" figureDoi-8="http://doi.org/10.5281/zenodo.4736192" httpUri-0="https://zenodo.org/record/4736174/files/figure.png" httpUri-1="https://zenodo.org/record/4736176/files/figure.png" httpUri-2="https://zenodo.org/record/4736178/files/figure.png" httpUri-3="https://zenodo.org/record/4736180/files/figure.png" httpUri-4="https://zenodo.org/record/4736182/files/figure.png" httpUri-5="https://zenodo.org/record/4736186/files/figure.png" httpUri-6="https://zenodo.org/record/4736188/files/figure.png" httpUri-7="https://zenodo.org/record/4736190/files/figure.png" httpUri-8="https://zenodo.org/record/4736192/files/figure.png" pageId="2" pageNumber="543">Figs 1–41</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30AFDA4FFA8FF85E1D1A2D2FB66BFC5" pageId="2" pageNumber="543" type="materials_examined">
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<emphasis id="B964723DFFA8FF85E1D1A2D2FE9FBF58" bold="true" box="[151,262,510,536]" pageId="2" pageNumber="543">Material.</emphasis>
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<materialsCitation id="3B78A472FFA8FF85E04BA2D3FE02BF21" ID-GBIF-Occurrence="3118550303" collectingDate="2014-05-22" collectionCode="T" country="Russia" county="Shkotovskiy district" latitude="43.078007" location="Shkotovo village" longLatPrecision="1" longitude="132.35881" municipality="Shkotovka River" pageId="2" pageNumber="543" specimenCount="1" stateProvince="Primorskiy" typeStatus="holotype">
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<typeStatus id="54AB108DFFA8FF85E04BA2D3FEE3BF59" box="[269,378,511,537]" pageId="2" pageNumber="543" type="holotype">Holotype</typeStatus>
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</emphasis>
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male imago (reared from larva, with larval skin),
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<emphasis id="B964723DFFA8FF85E2E1A2D2FB73BF58" bold="true" box="[935,1258,510,536]" pageId="2" pageNumber="543">
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<collectingCountry id="F307EEBFFFA8FF85E2E1A2D2FB7CBF58" box="[935,1253,510,536]" name="Russia" pageId="2" pageNumber="543">RUSSIAN FEDERATION</collectingCountry>
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:
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</emphasis>
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<collectingRegion id="49D460CDFFA8FF85E5B7A2D3FF7ABF7D" country="Russia" name="Primorskiy" pageId="2" pageNumber="543">PRIMORSKIY KRAY</collectingRegion>
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,
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<collectingCounty id="62CED6A3FFA8FF85E1B0A10FFE7BBF7D" box="[246,482,547,573]" pageId="2" pageNumber="543">Shkotovskiy district</collectingCounty>
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,
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<collectingMunicipality id="6BCB3455FFA8FF85E0B3A10FFD22BF7D" box="[501,699,547,573]" pageId="2" pageNumber="543">Shkotovka River</collectingMunicipality>
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, bridge,
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<location id="8ECFF8F4FFA8FF85E26AA10FFC6CBF7D" LSID="urn:lsid:plazi:treatment:03B91F39FFA8FF8CE1D1A243FD85BC91:8ECFF8F4FFA8FF85E26AA10FFC6CBF7D" box="[812,1013,547,573]" country="Russia" county="Shkotovskiy district" latitude="43.078007" longLatPrecision="1" longitude="132.35881" municipality="Shkotovka River" name="Shkotovo village" pageId="2" pageNumber="543" stateProvince="Primorskiy">Shkotovo village</location>
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,
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<geoCoordinate id="EE24C8E8FFA8FF85E54EA10FFB50BF7D" box="[1032,1225,547,573]" degrees="43" direction="north" minutes="04" orientation="latitude" pageId="2" pageNumber="543" precision="1" seconds="40.83" value="43.078007">43°04´40.83´´ N</geoCoordinate>
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<geoCoordinate id="EE24C8E8FFA8FF85E59EA10FFA01BF7D" box="[1240,1432,547,573]" degrees="132" direction="east" minutes="21" orientation="longitude" pageId="2" pageNumber="543" precision="1" seconds="31.72" value="132.35881">132°21´31.72´´E</geoCoordinate>
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,
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<date id="FFAE88EFFFA8FF85E1D1A16BFE95BF21" box="[151,268,583,609]" pageId="2" pageNumber="543" value="2014-05-22">
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<collectingDate id="EFEA7107FFA8FF85E1D1A16BFE95BF21" box="[151,268,583,609]" pageId="2" pageNumber="543" value="2014-05-22">22.V.2014</collectingDate>
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</date>
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,
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<collectionCode id="ED0136EAFFA8FF85E05DA16BFEB4BF21" box="[283,301,583,609]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="543">T</collectionCode>
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. Tiunova
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||
</materialsCitation>
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.
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<emphasis id="B964723DFFA8FF85E0EFA16BFDB8BF21" bold="true" box="[425,545,583,609]" pageId="2" pageNumber="543">
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<typeStatus id="54AB108DFFA8FF85E0EFA16BFDB8BF21" box="[425,545,583,609]" pageId="2" pageNumber="543" type="paratype">Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount id="9D1665A6FFA8FF85E372A16BFD1CBF21" box="[564,645,583,609]" pageId="2" pageNumber="543" type="male">2 male</specimenCount>
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,
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<specimenCount id="9D1665A6FFA8FF85E3D2A16BFD60BF21" box="[660,761,583,609]" pageId="2" pageNumber="543" type="female">1 female</specimenCount>
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imagines,
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<specimenCount id="9D1665A6FFA8FF85E23DA16BFC42BF21" box="[891,987,583,609]" pageId="2" pageNumber="543" type="larva">2 larvae</specimenCount>
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, and
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<materialsCitation id="3B78A472FFA8FF85E55BA16BFE36BFC5" ID-GBIF-Occurrence="3118550301" collectingDate="2014-05-22" collectionCode="BM" country="Russia" county="Shkotovskiy district" latitude="43.078007" location="Shkotovo village" longLatPrecision="1" longitude="132.35881" municipality="Shkotovka River" pageId="2" pageNumber="543" specimenCode="BM175" specimenCount="1" specimenCount-larva="1" stateProvince="Primorskiy" typeStatus="paratype">
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<specimenCount id="9D1665A6FFA8FF85E55BA16BFB11BF21" box="[1053,1160,583,609]" pageId="2" pageNumber="543" type="larva">one larva</specimenCount>
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(
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<specimenCode id="DBB60654FFA8FF85E5DAA16AFB6CBF20" box="[1180,1269,582,608]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="2" pageNumber="543">BM175</specimenCode>
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)
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</emphasis>
|
||
same data as holotype; same place,
|
||
<date id="FFAE88EFFFA8FF85E0D4A146FE36BFC5" box="[402,431,618,645]" pageId="2" pageNumber="543" value="2015-05-11">11.</date>
|
||
</materialsCitation>
|
||
<collectionCode id="ED0136EAFFA8FF85E0E8A146FE5DBFC5" box="[430,452,618,645]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="543" type="Museum">V</collectionCode>
|
||
.2015,
|
||
<materialsCitation id="3B78A472FFA8FF85E357A146FC31BFC5" ID-GBIF-Occurrence="3118550302" box="[529,936,618,645]" collectionCode="T" country="Russia" pageId="2" pageNumber="543" specimenCount="5" specimenCount-larva="5" stateProvince="Primorskiy" typeStatus="paratype">
|
||
<specimenCount id="9D1665A6FFA8FF85E357A146FDF4BFC5" box="[529,621,618,645]" pageId="2" pageNumber="543" type="larva">5 larvae</specimenCount>
|
||
,
|
||
<collectionCode id="ED0136EAFFA8FF85E33EA147FD13BFC5" box="[632,650,619,645]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="543">T</collectionCode>
|
||
. Tiunova; same place,
|
||
<date id="FFAE88EFFFA8FF85E2CFA146FC31BFC5" box="[905,936,618,645]" pageId="2" pageNumber="543" value="2018-05-15">15.</date>
|
||
</materialsCitation>
|
||
<collectionCode id="ED0136EAFFA8FF85E2E1A146FC24BFC5" box="[935,957,618,645]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="543" type="Museum">V</collectionCode>
|
||
.2018,
|
||
<materialsCitation id="3B78A472FFA8FF85E54CA147FB62BFC5" ID-GBIF-Occurrence="3118550304" box="[1034,1275,619,645]" collectionCode="T" country="Russia" pageId="2" pageNumber="543" specimenCount="27" specimenCount-larva="27" stateProvince="Primorskiy" typeStatus="paratype">
|
||
<specimenCount id="9D1665A6FFA8FF85E54CA147FBEDBFC5" box="[1034,1140,619,645]" pageId="2" pageNumber="543" type="larva">27 larvae</specimenCount>
|
||
,
|
||
<collectionCode id="ED0136EAFFA8FF85E539A147FB08BFC5" box="[1151,1169,619,645]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="543">T</collectionCode>
|
||
. Tiunova
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30AFDA4FFA8FF8FE181A1A2FCDCBA92" lastPageId="8" lastPageNumber="549" pageId="2" pageNumber="543" type="description">
|
||
<paragraph id="8BAFAE2FFFA8FF85E181A1A2FB94BF8D" blockId="2.[151,1437,510,2013]" pageId="2" pageNumber="543">
|
||
<emphasis id="B964723DFFA8FF85E181A1A2FE7EBFE9" bold="true" box="[199,487,654,681]" pageId="2" pageNumber="543">
|
||
Description.
|
||
<emphasis id="B964723DFFA8FF85E018A1A2FE7EBFE9" bold="true" box="[350,487,654,681]" italics="true" pageId="2" pageNumber="543">Male imago</emphasis>
|
||
</emphasis>
|
||
(in alcohol) (
|
||
<figureCitation id="132BB2AAFFA8FF85E33DA1A3FD22BFE9" box="[635,699,655,681]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[252,1335,182,1919]" captionTargetId="figure-17@3.[252,1335,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–2. Baetis majus Tiunova sp. nov., 1, male imago, holotype, lateral view; 2, female imago, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736174" httpUri="https://zenodo.org/record/4736174/files/figure.png" pageId="2" pageNumber="543">Fig. 1</figureCitation>
|
||
). Length (mm): body 4.7–5.2; forewings 5.5–5.6; cerci 12.0–13.0; femora 1.1; tibia 1.7–1.8.
|
||
<emphasis id="B964723DFFA8FF85E0FCA19FFD9ABF8D" bold="true" box="[442,515,691,717]" pageId="2" pageNumber="543">Head:</emphasis>
|
||
light brown or brownish. Antennae yellowish.
|
||
</paragraph>
|
||
<paragraph id="8BAFAE2FFFA8FF85E181A1FBFC62BE55" blockId="2.[151,1437,510,2013]" pageId="2" pageNumber="543">
|
||
Turbinate eyes moderately high (
|
||
<figureCitation id="132BB2AAFFA8FF85E375A1FBFDEABFB1" box="[563,627,727,753]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[252,1335,182,1919]" captionTargetId="figure-17@3.[252,1335,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–2. Baetis majus Tiunova sp. nov., 1, male imago, holotype, lateral view; 2, female imago, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736174" httpUri="https://zenodo.org/record/4736174/files/figure.png" pageId="2" pageNumber="543">Fig. 1</figureCitation>
|
||
); faceted surface oval in dorsal view, 1.2 times longer than wide; facetted surface of turbinate eyes light orange; the shaft grayish with a yellowish ring.
|
||
</paragraph>
|
||
<paragraph id="8BAFAE2FFFA8FF85E181A033FBF7BEAD" blockId="2.[151,1437,510,2013]" pageId="2" pageNumber="543">
|
||
<emphasis id="B964723DFFA8FF85E181A033FEB9BE79" bold="true" box="[199,288,799,825]" pageId="2" pageNumber="543">Thorax</emphasis>
|
||
: Medioscutum and submedioscutum light brown; posterior scutal protuberance dark brown with light lateral sides; scutellum grayish; scuto-scutellar impression white; median longitudinal suture dark brown; sublateroscutum dark brown; anterior phragma dark brown.All legs white (
|
||
<figureCitation id="132BB2AAFFA8FF85E288A04BFB94BEC1" box="[974,1037,871,897]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[252,1335,182,1919]" captionTargetId="figure-17@3.[252,1335,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–2. Baetis majus Tiunova sp. nov., 1, male imago, holotype, lateral view; 2, female imago, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736174" httpUri="https://zenodo.org/record/4736174/files/figure.png" pageId="2" pageNumber="543">Fig. 1</figureCitation>
|
||
). Lengths ratio of individual foreleg segments: 1.1:1.8:0.8:0.5:0.35:0.2. Forewing transparent, all veins pale; pterostigma milky on a dark background, with 5–6 cross veins. Hind wing hyaline, transparent, and approximately 2 times longer than wide, rounded apex and two simple longitudinal veins; cross veins absent; costal projection present (
|
||
<figureCitation id="132BB2AAFFA8FF85E55AA0FFFBF9BEAD" box="[1052,1120,979,1005]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1968,1992]" captionTargetBox="[171,1416,189,1933]" captionTargetId="figure-27@4.[160,1427,181,1944]" captionTargetPageId="4" captionText="FIGURES 3–4. Baetis majus Tiunova sp. nov., male imagoes, holotype: 3, hind wing; 4, unistyliger and forceps." figureDoi="http://doi.org/10.5281/zenodo.4736176" httpUri="https://zenodo.org/record/4736176/files/figure.png" pageId="2" pageNumber="543">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAFAE2FFFA8FF85E181A0DAFD23B9A9" blockId="2.[151,1437,510,2013]" pageId="2" pageNumber="543">
|
||
<emphasis id="B964723DFFA8FF85E181A0DAFEA5B950" bold="true" box="[199,316,1014,1040]" pageId="2" pageNumber="543">Abdomen</emphasis>
|
||
: Tergum I brownish; terga II–VI yellowish or whitish, translucent (
|
||
<figureCitation id="132BB2AAFFA8FF85E512A0DBFB00B951" box="[1108,1177,1015,1041]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[252,1335,182,1919]" captionTargetId="figure-17@3.[252,1335,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–2. Baetis majus Tiunova sp. nov., 1, male imago, holotype, lateral view; 2, female imago, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736174" httpUri="https://zenodo.org/record/4736174/files/figure.png" pageId="2" pageNumber="543">Fig. 1</figureCitation>
|
||
); terga VIII–X brown with darker lateral margins. Sterna I–VI white, translucent; sterna VII–IX brown, opaque. Styliger whitish (
|
||
<figureCitation id="132BB2AAFFA8FF85E40CA737FA16B975" box="[1354,1423,1051,1077]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1968,1992]" captionTargetBox="[171,1416,189,1933]" captionTargetId="figure-27@4.[160,1427,181,1944]" captionTargetPageId="4" captionText="FIGURES 3–4. Baetis majus Tiunova sp. nov., male imagoes, holotype: 3, hind wing; 4, unistyliger and forceps." figureDoi="http://doi.org/10.5281/zenodo.4736176" httpUri="https://zenodo.org/record/4736176/files/figure.png" pageId="2" pageNumber="543">Fig. 4</figureCitation>
|
||
). Unistiliger and segment I of forceps brownish; segments II and III whitish; unistiliger with parallel margins, slightly elongated, almost as long as wide, with a distinct small projection at the apex of the inner edge; segment I of forceps conical and elongate; segment II relatively narrow, slightly widening towards the apex and with beveled inner apical margin; segment III small, well defined, strongly expanded to the apex, its length only slightly exceeds its maximum width (
|
||
<figureCitation id="132BB2AAFFA8FF85E1A0A7E3FEB5B9A9" box="[230,300,1231,1257]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1968,1992]" captionTargetBox="[171,1416,189,1933]" captionTargetId="figure-27@4.[160,1427,181,1944]" captionTargetPageId="4" captionText="FIGURES 3–4. Baetis majus Tiunova sp. nov., male imagoes, holotype: 3, hind wing; 4, unistyliger and forceps." figureDoi="http://doi.org/10.5281/zenodo.4736176" httpUri="https://zenodo.org/record/4736176/files/figure.png" pageId="2" pageNumber="543">Fig. 4</figureCitation>
|
||
). Caudal filaments whitish (
|
||
<figureCitation id="132BB2AAFFA8FF85E321A7E3FD30B9A9" box="[615,681,1231,1257]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[252,1335,182,1919]" captionTargetId="figure-17@3.[252,1335,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–2. Baetis majus Tiunova sp. nov., 1, male imago, holotype, lateral view; 2, female imago, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736174" httpUri="https://zenodo.org/record/4736174/files/figure.png" pageId="2" pageNumber="543">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAFAE2FFFA8FF85E181A7DEFC28B839" blockId="2.[151,1437,510,2013]" pageId="2" pageNumber="543">
|
||
<emphasis id="B964723DFFA8FF85E181A7DEFEEDB84D" bold="true" box="[199,372,1266,1293]" italics="true" pageId="2" pageNumber="543">Female imago.</emphasis>
|
||
Length (mm): body 5.5; forewings 6.5; cerci lost; foreleg: femora 1.0; tibia 1.3. General color of body brown (
|
||
<figureCitation id="132BB2AAFFA8FF85E001A63BFE12B871" box="[327,395,1303,1329]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[252,1335,182,1919]" captionTargetId="figure-17@3.[252,1335,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–2. Baetis majus Tiunova sp. nov., 1, male imago, holotype, lateral view; 2, female imago, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736174" httpUri="https://zenodo.org/record/4736174/files/figure.png" pageId="2" pageNumber="543">Fig. 2</figureCitation>
|
||
). Head grayish, posterior margin brownish; antennae whitish. Thorax light brown, with brown median longitudinal suture. Wings transparent; all veins brownish. Legs yellowish or brownish. Abdomen terga I–VIII (with eggs) light brown without maculation; terga IX–X lighter.
|
||
</paragraph>
|
||
<paragraph id="8BAFAE2FFFA8FF85E181A6AFFE0CB881" blockId="2.[151,1437,510,2013]" pageId="2" pageNumber="543">
|
||
<emphasis id="B964723DFFA8FF85E181A6AFFEF7B8DD" bold="true" box="[199,366,1410,1437]" italics="true" pageId="2" pageNumber="543">Mature larvae</emphasis>
|
||
(in alcohol): Length (mm): body 4.7–5.3; cerci 2.5–3.1. General body color brown with light maculation (
|
||
<figureCitation id="132BB2AAFFA8FF85E064A68BFE1FB881" box="[290,390,1446,1473]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1550,1574]" captionTargetBox="[170,1418,309,1527]" captionTargetId="figure-92@5.[170,1418,309,1527]" captionTargetPageId="5" captionText="FIGURES 5–7. Color pattern of Baetis majus Tiunova sp. nov., larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736178" httpUri="https://zenodo.org/record/4736178/files/figure.png" pageId="2" pageNumber="543">Figs 5–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAFAE2FFFA8FF82E181A6E7FACCBC5D" blockId="2.[151,1437,510,2013]" lastBlockId="5.[151,1437,150,285]" lastPageId="5" lastPageNumber="546" pageId="2" pageNumber="543">
|
||
<emphasis id="B964723DFFA8FF85E181A6E7FE89B8A5" bold="true" box="[199,272,1483,1509]" pageId="2" pageNumber="543">Head:</emphasis>
|
||
antennae brownish; antennal pedicel and scape without spatulas, with rare fine hair-like setae only (
|
||
<figureCitation id="132BB2AAFFA8FF85E1E6A6C3FF7CBB49" box="[160,229,1519,1545]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1930,1954]" captionTargetBox="[151,1436,669,1907]" captionTargetId="figure-277@6.[151,1436,669,1907]" captionTargetPageId="6" captionText="FIGURES 8–13. Baetis majus Tiunova sp. nov., larvae, dorsal view, paratypes: 8, scape; 9, surface of the head; 10, spatulas on head; 11, canines of left and right mandible; 12, glossae and paraglossae; 13, setae on paraglossa. Scale bar: 8, 11–12, 0.01 mm; 9, 0.02 mm; 10, 13, 0.002 mm." figureDoi="http://doi.org/10.5281/zenodo.4736180" httpUri="https://zenodo.org/record/4736180/files/figure.png" pageId="2" pageNumber="543">Fig. 8</figureCitation>
|
||
); head brown with light maculation on vertex; surface of the head covered numerous strong spatulas and fine hair-like setae (
|
||
<figureCitation id="132BB2AAFFA8FF85E0C6A53FFE61BB6D" box="[384,504,1555,1581]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1930,1954]" captionTargetBox="[151,1436,669,1907]" captionTargetId="figure-277@6.[151,1436,669,1907]" captionTargetPageId="6" captionText="FIGURES 8–13. Baetis majus Tiunova sp. nov., larvae, dorsal view, paratypes: 8, scape; 9, surface of the head; 10, spatulas on head; 11, canines of left and right mandible; 12, glossae and paraglossae; 13, setae on paraglossa. Scale bar: 8, 11–12, 0.01 mm; 9, 0.02 mm; 10, 13, 0.002 mm." figureDoi="http://doi.org/10.5281/zenodo.4736180" httpUri="https://zenodo.org/record/4736180/files/figure.png" pageId="2" pageNumber="543">Figs 9–10</figureCitation>
|
||
); frons, clypeus and labrum white; fronto-clypeal suture black; gena with a pair of dark brown stripes. Eyes of female black (
|
||
<figureCitation id="132BB2AAFFA8FF85E3D9A51BFD7CBB11" box="[671,741,1591,1617]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1550,1574]" captionTargetBox="[170,1418,309,1527]" captionTargetId="figure-92@5.[170,1418,309,1527]" captionTargetPageId="5" captionText="FIGURES 5–7. Color pattern of Baetis majus Tiunova sp. nov., larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736178" httpUri="https://zenodo.org/record/4736178/files/figure.png" pageId="2" pageNumber="543">Fig. 7</figureCitation>
|
||
), larval turbinate eyes of male light orange (
|
||
<figureCitation id="132BB2AAFFA8FF85E5A0A51BFAB5BB11" box="[1254,1324,1590,1617]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1550,1574]" captionTargetBox="[170,1418,309,1527]" captionTargetId="figure-92@5.[170,1418,309,1527]" captionTargetPageId="5" captionText="FIGURES 5–7. Color pattern of Baetis majus Tiunova sp. nov., larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736178" httpUri="https://zenodo.org/record/4736178/files/figure.png" pageId="2" pageNumber="543">Fig. 5</figureCitation>
|
||
). Labrum more or less rounded (width/length ratio of 1.3); dorsal surface with 1 + 2–3 long submarginal setae, arranged in one regular row, and row of long pointed setae laterally on both margins; posterior area with rare thick and short setae (
|
||
<figureCitation id="132BB2AAFFA8FF85E1A7A58FFEAFBBFD" box="[225,310,1699,1725]" captionStart="FIGURES 14–20" captionStartId="7.[151,264,1931,1955]" captionTargetBox="[156,1421,190,1907]" captionTargetId="figure-17@7.[151,1436,181,1907]" captionTargetPageId="7" captionText="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="2" pageNumber="543">Fig. 14</figureCitation>
|
||
). Canines of right mandible with 7 teeth divided into two groups; incisor with three large teeth of relatively equal size; kinetodontium with four teeth; the first, second and third relatively equal size, fourth smallest (
|
||
<figureCitation id="132BB2AAFFA8FF85E1D9A5C7FF6FBA45" box="[159,246,1771,1797]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1930,1954]" captionTargetBox="[151,1436,669,1907]" captionTargetId="figure-277@6.[151,1436,669,1907]" captionTargetPageId="6" captionText="FIGURES 8–13. Baetis majus Tiunova sp. nov., larvae, dorsal view, paratypes: 8, scape; 9, surface of the head; 10, spatulas on head; 11, canines of left and right mandible; 12, glossae and paraglossae; 13, setae on paraglossa. Scale bar: 8, 11–12, 0.01 mm; 9, 0.02 mm; 10, 13, 0.002 mm." figureDoi="http://doi.org/10.5281/zenodo.4736180" httpUri="https://zenodo.org/record/4736180/files/figure.png" pageId="2" pageNumber="543">Figs 11</figureCitation>
|
||
,
|
||
<figureCitation id="132BB2AAFFA8FF85E045A5C6FEBBBA45" box="[259,290,1770,1797]" captionStart="FIGURES 14–20" captionStartId="7.[151,264,1931,1955]" captionTargetBox="[156,1421,190,1907]" captionTargetId="figure-17@7.[151,1436,181,1907]" captionTargetPageId="7" captionText="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="2" pageNumber="543">15</figureCitation>
|
||
); prostheca toothbrush-like. Left mandible canines with 7 teeth; incisor with three teeth, first two the largest; kinetodontium with four teeth, first and fourth the smallest and second the largest (
|
||
<figureCitation id="132BB2AAFFA8FF85E5DFA423FB76BA69" box="[1177,1263,1807,1833]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1930,1954]" captionTargetBox="[151,1436,669,1907]" captionTargetId="figure-277@6.[151,1436,669,1907]" captionTargetPageId="6" captionText="FIGURES 8–13. Baetis majus Tiunova sp. nov., larvae, dorsal view, paratypes: 8, scape; 9, surface of the head; 10, spatulas on head; 11, canines of left and right mandible; 12, glossae and paraglossae; 13, setae on paraglossa. Scale bar: 8, 11–12, 0.01 mm; 9, 0.02 mm; 10, 13, 0.002 mm." figureDoi="http://doi.org/10.5281/zenodo.4736180" httpUri="https://zenodo.org/record/4736180/files/figure.png" pageId="2" pageNumber="543">Figs 11</figureCitation>
|
||
,
|
||
<figureCitation id="132BB2AAFFA8FF85E5BAA423FA82BA6A" box="[1276,1307,1807,1834]" captionStart="FIGURES 14–20" captionStartId="7.[151,264,1931,1955]" captionTargetBox="[156,1421,190,1907]" captionTargetId="figure-17@7.[151,1436,181,1907]" captionTargetPageId="7" captionText="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="2" pageNumber="543">16</figureCitation>
|
||
); prostheca brush-like, with mostly short blunt appendages and two or three pointed ones on inner side. Maxillary palp twosegmented, surface covered with hair-like setae, more densely located on distal part of second segment; second segment longer than first segment (1.7 times) (
|
||
<figureCitation id="132BB2AAFFA8FF85E3DFA457FD72BAD5" box="[665,747,1915,1941]" captionStart="FIGURES 14–20" captionStartId="7.[151,264,1931,1955]" captionTargetBox="[156,1421,190,1907]" captionTargetId="figure-17@7.[151,1436,181,1907]" captionTargetPageId="7" captionText="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="2" pageNumber="543">Fig. 17</figureCitation>
|
||
). Labium with paraglossae concave in middle, 1.4 times wider than glossae; apical part of paraglossae with two regular rows of long setae; 6 long bristles located along outer margin in ventral surface (
|
||
<figureCitation id="132BB2AAFFA8FF85E02CA4EFFE27BA9D" box="[362,446,1987,2013]" captionStart="FIGURES 14–20" captionStartId="7.[151,264,1931,1955]" captionTargetBox="[156,1421,190,1907]" captionTargetId="figure-17@7.[151,1436,181,1907]" captionTargetPageId="7" captionText="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="2" pageNumber="543">Fig. 18</figureCitation>
|
||
); distal part of dorsal surface of paraglossa covered with small stout setae (
|
||
<figureCitation id="132BB2AAFFA8FF85E44BA4EFFA09BA9D" box="[1293,1424,1987,2013]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1930,1954]" captionTargetBox="[151,1436,669,1907]" captionTargetId="figure-277@6.[151,1436,669,1907]" captionTargetPageId="6" captionText="FIGURES 8–13. Baetis majus Tiunova sp. nov., larvae, dorsal view, paratypes: 8, scape; 9, surface of the head; 10, spatulas on head; 11, canines of left and right mandible; 12, glossae and paraglossae; 13, setae on paraglossa. Scale bar: 8, 11–12, 0.01 mm; 9, 0.02 mm; 10, 13, 0.002 mm." figureDoi="http://doi.org/10.5281/zenodo.4736180" httpUri="https://zenodo.org/record/4736180/files/figure.png" pageId="2" pageNumber="543">Figs 12–13</figureCitation>
|
||
). Glossae triangular with row of 9–10 stout setae on outer and 5–6 long setae on inner margins (
|
||
<figureCitation id="132BB2AAFFAFFF82E59EA3BBFAB4BDF1" box="[1240,1325,151,177]" captionStart="FIGURES 14–20" captionStartId="7.[151,264,1931,1955]" captionTargetBox="[156,1421,190,1907]" captionTargetId="figure-17@7.[151,1436,181,1907]" captionTargetPageId="7" captionText="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="5" pageNumber="546">Fig. 18</figureCitation>
|
||
). General shape of the second and third segments of the labial palp rather stocky (
|
||
<figureCitation id="132BB2AAFFAFFF82E283A397FBD0BD95" box="[965,1097,187,213]" captionStart="FIGURES 14–20" captionStartId="7.[151,264,1931,1955]" captionTargetBox="[156,1421,190,1907]" captionTargetId="figure-17@7.[151,1436,181,1907]" captionTargetPageId="7" captionText="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="5" pageNumber="546">Figs 19–20</figureCitation>
|
||
); third segment wide, its apex truncated; 1.3 times wider than the second segment; ventral surface covered with long stout setae (
|
||
<figureCitation id="132BB2AAFFAFFF82E5A0A3F3FAA0BDB9" box="[1254,1337,223,249]" captionStart="FIGURES 14–20" captionStartId="7.[151,264,1931,1955]" captionTargetBox="[156,1421,190,1907]" captionTargetId="figure-17@7.[151,1436,181,1907]" captionTargetPageId="7" captionText="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="5" pageNumber="546">Fig. 19</figureCitation>
|
||
); second segment with rounded apicomedial projection; dorsal surface with a row of 4–5 long pointed setae (
|
||
<figureCitation id="132BB2AAFFAFFF82E5B2A22FFAD1BC5D" box="[1268,1352,259,285]" captionStart="FIGURES 14–20" captionStartId="7.[151,264,1931,1955]" captionTargetBox="[156,1421,190,1907]" captionTargetId="figure-17@7.[151,1436,181,1907]" captionTargetPageId="7" captionText="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="5" pageNumber="546">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6FFEA7FFA9FF84E1D1A4BBFAAABAF0" ID-DOI="http://doi.org/10.5281/zenodo.4736174" ID-Zenodo-Dep="4736174" box="[151,1331,1943,1968]" httpUri="https://zenodo.org/record/4736174/files/figure.png" pageId="3" pageNumber="544" startId="3.[151,264,1943,1967]" targetBox="[252,1335,182,1919]" targetPageId="3">
|
||
<paragraph id="8BAFAE2FFFA9FF84E1D1A4BBFAAABAF0" blockId="3.[151,1331,1943,1968]" box="[151,1331,1943,1968]" pageId="3" pageNumber="544">
|
||
<emphasis id="B964723DFFA9FF84E1D1A4BBFEA3BAEF" bold="true" box="[151,314,1943,1967]" pageId="3" pageNumber="544">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="4C10D5ACFFA9FF84E007A4B4FD84BAF0" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[321,541,1944,1968]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA9FF84E007A4B4FE5ABAEF" box="[321,451,1944,1967]" italics="true" pageId="3" pageNumber="544">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFA9FF84E365A4B4FDEABAF0" bold="true" box="[547,627,1944,1968]" pageId="3" pageNumber="544">
|
||
<taxonomicNameLabel id="A257CF46FFA9FF84E365A4B4FDEABAF0" box="[547,627,1944,1968]" pageId="3" pageNumber="544" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, 1, male imago, holotype, lateral view; 2, female imago, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6FFEA7FFAEFF83E1D1A49CFA8CBA88" ID-DOI="http://doi.org/10.5281/zenodo.4736176" ID-Zenodo-Dep="4736176" box="[151,1301,1968,1992]" httpUri="https://zenodo.org/record/4736176/files/figure.png" pageId="4" pageNumber="545" startId="4.[151,264,1968,1992]" targetBox="[171,1416,189,1933]" targetPageId="4">
|
||
<paragraph id="8BAFAE2FFFAEFF83E1D1A49CFA8CBA88" blockId="4.[151,1301,1968,1992]" box="[151,1301,1968,1992]" pageId="4" pageNumber="545">
|
||
<emphasis id="B964723DFFAEFF83E1D1A49CFEA2BA88" bold="true" box="[151,315,1968,1992]" pageId="4" pageNumber="545">FIGURES 3–4.</emphasis>
|
||
<taxonomicName id="4C10D5ACFFAEFF83E007A49DFD84BA88" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[321,541,1968,1992]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="545" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFAEFF83E007A49DFE5ABA88" box="[321,451,1969,1992]" italics="true" pageId="4" pageNumber="545">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFAEFF83E365A49CFDEABA88" bold="true" box="[547,627,1968,1992]" pageId="4" pageNumber="545">
|
||
<taxonomicNameLabel id="A257CF46FFAEFF83E365A49CFDEABA88" box="[547,627,1968,1992]" pageId="4" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male imagoes, holotype: 3, hind wing; 4, unistyliger and forceps.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6FFEA7FFAFFF82E1D1A522FEFDBB0A" ID-DOI="http://doi.org/10.5281/zenodo.4736178" ID-Zenodo-Dep="4736178" httpUri="https://zenodo.org/record/4736178/files/figure.png" pageId="5" pageNumber="546" startId="5.[151,264,1550,1574]" targetBox="[170,1418,309,1527]" targetPageId="5">
|
||
<paragraph id="8BAFAE2FFFAFFF82E1D1A522FEFDBB0A" blockId="5.[151,1437,1550,1610]" pageId="5" pageNumber="546">
|
||
<emphasis id="B964723DFFAFFF82E1D1A522FEA4BB67" bold="true" box="[151,317,1550,1575]" pageId="5" pageNumber="546">FIGURES 5–7.</emphasis>
|
||
Color pattern of
|
||
<taxonomicName id="4C10D5ACFFAFFF82E0B2A523FD4DBB66" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[500,724,1550,1574]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="546" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFAFFF82E0B2A523FDE0BB66" box="[500,633,1551,1574]" italics="true" pageId="5" pageNumber="546">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFAFFF82E39AA523FCB6BB67" bold="true" box="[732,815,1551,1575]" pageId="5" pageNumber="546">
|
||
<taxonomicNameLabel id="A257CF46FFAFFF82E39AA523FCB6BB67" box="[732,815,1551,1575]" pageId="5" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAFAE2FFFAFFF82E181A555FAE4BABB" blockId="5.[151,1437,1656,2043]" pageId="5" pageNumber="546">
|
||
<emphasis id="B964723DFFAFFF82E181A555FEB0BBD3" bold="true" box="[199,297,1657,1683]" pageId="5" pageNumber="546">Thorax:</emphasis>
|
||
brown with contrasting whitish or brownish stripes on the mesonotum (
|
||
<figureCitation id="132BB2AAFFAFFF82E518A555FB5FBBD3" box="[1118,1222,1656,1683]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1550,1574]" captionTargetBox="[170,1418,309,1527]" captionTargetId="figure-92@5.[170,1418,309,1527]" captionTargetPageId="5" captionText="FIGURES 5–7. Color pattern of Baetis majus Tiunova sp. nov., larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736178" httpUri="https://zenodo.org/record/4736178/files/figure.png" pageId="5" pageNumber="546">Figs 5, 7</figureCitation>
|
||
). Mesonotum with a V-shaped white spot near anterior margin. Pronotum with a pair of light spots laterally. Legs white, joints of leg segments brown (
|
||
<figureCitation id="132BB2AAFFAFFF82E018A5EDFE3ABB9C" box="[350,419,1729,1756]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1550,1574]" captionTargetBox="[170,1418,309,1527]" captionTargetId="figure-92@5.[170,1418,309,1527]" captionTargetPageId="5" captionText="FIGURES 5–7. Color pattern of Baetis majus Tiunova sp. nov., larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736178" httpUri="https://zenodo.org/record/4736178/files/figure.png" pageId="5" pageNumber="546">Fig. 6</figureCitation>
|
||
). Femora with brown rectangular spot near inner margin (
|
||
<figureCitation id="132BB2AAFFAFFF82E574A5EDFB1CBB9B" box="[1074,1157,1729,1755]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="5" pageNumber="546">Fig. 21</figureCitation>
|
||
); outer edge with row of long, apically widened setae in distal part (
|
||
<figureCitation id="132BB2AAFFAFFF82E3C6A5C9FD4ABBBF" box="[640,723,1765,1791]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="5" pageNumber="546">Fig. 22</figureCitation>
|
||
) and rounded setae in basal part; inner margin with a irregular row of small pointed setae in basal part; femoral villopore present (
|
||
<figureCitation id="132BB2AAFFAFFF82E2DEA425FC77BA63" box="[920,1006,1801,1827]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="5" pageNumber="546">Fig. 23</figureCitation>
|
||
). Tibia and tarsi brown in apical part; stout setae arranged in an irregular rows along outer margin of tibia (
|
||
<figureCitation id="132BB2AAFFAFFF82E2D6A401FC7FBA07" box="[912,998,1837,1863]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="5" pageNumber="546">Figs 21</figureCitation>
|
||
,
|
||
<figureCitation id="132BB2AAFFAFFF82E2B7A401FBAEBA07" box="[1009,1079,1837,1863]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1966,1990]" captionTargetBox="[151,1436,490,1943]" captionTargetId="figure-161@9.[151,1436,489,1943]" captionTargetPageId="9" captionText="FIGURES 27–32. Baetis majus Tiunova sp. nov., larvae, paratypes: 27, 29, inner margin of fore tibia; 28, inner margin of fore tarsus; 30, tarsal claw; 31, paraproct; 32, spatulas on paraproct. Scale bar: 27–31, 0.01 mm; 32, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.4736188" httpUri="https://zenodo.org/record/4736188/files/figure.png" pageId="5" pageNumber="546">27, 29</figureCitation>
|
||
) and a regular row on tarsi (
|
||
<figureCitation id="132BB2AAFFAFFF82E429A401FF2CBA2B" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="5" pageNumber="546">Fig. 21</figureCitation>
|
||
,
|
||
<figureCitation id="132BB2AAFFAFFF82E187A47DFF78BA2B" box="[193,225,1873,1899]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1966,1990]" captionTargetBox="[151,1436,490,1943]" captionTargetId="figure-161@9.[151,1436,489,1943]" captionTargetPageId="9" captionText="FIGURES 27–32. Baetis majus Tiunova sp. nov., larvae, paratypes: 27, 29, inner margin of fore tibia; 28, inner margin of fore tarsus; 30, tarsal claw; 31, paraproct; 32, spatulas on paraproct. Scale bar: 27–31, 0.01 mm; 32, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.4736188" httpUri="https://zenodo.org/record/4736188/files/figure.png" pageId="5" pageNumber="546">28</figureCitation>
|
||
); smaller setae located along inner margin of tibia and tarsi; dorsal surface of femora, tibia and tarsi covered with small bluntly rounded spatulas (
|
||
<figureCitation id="132BB2AAFFAFFF82E378A459FD5DBACF" box="[574,708,1909,1935]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1966,1990]" captionTargetBox="[151,1436,490,1943]" captionTargetId="figure-161@9.[151,1436,489,1943]" captionTargetPageId="9" captionText="FIGURES 27–32. Baetis majus Tiunova sp. nov., larvae, paratypes: 27, 29, inner margin of fore tibia; 28, inner margin of fore tarsus; 30, tarsal claw; 31, paraproct; 32, spatulas on paraproct. Scale bar: 27–31, 0.01 mm; 32, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.4736188" httpUri="https://zenodo.org/record/4736188/files/figure.png" pageId="5" pageNumber="546">Figs 27–29</figureCitation>
|
||
). Patella-tibial suture present (
|
||
<figureCitation id="132BB2AAFFAFFF82E558A459FBEFBACF" box="[1054,1142,1909,1935]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="5" pageNumber="546">Figs 21</figureCitation>
|
||
,
|
||
<figureCitation id="132BB2AAFFAFFF82E5C2A459FB3DBACF" box="[1156,1188,1909,1935]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1966,1990]" captionTargetBox="[151,1436,490,1943]" captionTargetId="figure-161@9.[151,1436,489,1943]" captionTargetPageId="9" captionText="FIGURES 27–32. Baetis majus Tiunova sp. nov., larvae, paratypes: 27, 29, inner margin of fore tibia; 28, inner margin of fore tarsus; 30, tarsal claw; 31, paraproct; 32, spatulas on paraproct. Scale bar: 27–31, 0.01 mm; 32, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.4736188" httpUri="https://zenodo.org/record/4736188/files/figure.png" pageId="5" pageNumber="546">27</figureCitation>
|
||
). Tarsal claws brown, with row of 13–15 teeth increasing in length toward apex. Apex of tarsal claw without subapical setae (
|
||
<figureCitation id="132BB2AAFFAFFF82E449A4B5FAFCBAF3" box="[1295,1381,1945,1971]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="5" pageNumber="546">Figs 24</figureCitation>
|
||
,
|
||
<figureCitation id="132BB2AAFFAFFF82E436A4B5FA09BAF3" box="[1392,1424,1945,1971]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1966,1990]" captionTargetBox="[151,1436,490,1943]" captionTargetId="figure-161@9.[151,1436,489,1943]" captionTargetPageId="9" captionText="FIGURES 27–32. Baetis majus Tiunova sp. nov., larvae, paratypes: 27, 29, inner margin of fore tibia; 28, inner margin of fore tarsus; 30, tarsal claw; 31, paraproct; 32, spatulas on paraproct. Scale bar: 27–31, 0.01 mm; 32, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.4736188" httpUri="https://zenodo.org/record/4736188/files/figure.png" pageId="5" pageNumber="546">30</figureCitation>
|
||
). Lengths (mm) of the leg segments as follows: Foreleg: femur 0.9‒0.95; tibia 0.5‒0.6; and tarsus 0.5‒0.55. Middle leg: femur 0.9‒0.95; tibia 0.5‒0.6; and tarsus 0.5. Hind leg: femur 0.9‒1.05; tibia 0.55‒0.9; and tarsus 0.45‒0.5.
|
||
</paragraph>
|
||
<paragraph id="8BAFAE2FFFACFF81E181A3BAFC3CBFC5" blockId="6.[151,1437,150,645]" pageId="6" pageNumber="547">
|
||
<emphasis id="B964723DFFACFF81E181A3BAFEDCBDF0" bold="true" box="[199,325,150,176]" pageId="6" pageNumber="547">Abdomen:</emphasis>
|
||
Terga with contrasted white and black or dark brown maculations (
|
||
<figureCitation id="132BB2AAFFACFF81E504A3BBFB31BDF1" box="[1090,1192,150,177]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1550,1574]" captionTargetBox="[170,1418,309,1527]" captionTargetId="figure-92@5.[170,1418,309,1527]" captionTargetPageId="5" captionText="FIGURES 5–7. Color pattern of Baetis majus Tiunova sp. nov., larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736178" httpUri="https://zenodo.org/record/4736178/files/figure.png" pageId="6" pageNumber="547">Figs 5, 7</figureCitation>
|
||
). Tergum I brownish, posterior margin darker; terga II–VIII with a pair of antero-median light stripes and spots contoured with dark brown; terga II–III and VII darker than terga IV and VI with a pair of large white spots joining along the middle part at the posterior margin; antero-lateral corners and lateral margins white; postero-lateral corners black; terga IX and X completely white (
|
||
<figureCitation id="132BB2AAFFACFF81E0A0A20BFDC8BC01" box="[486,593,294,321]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1550,1574]" captionTargetBox="[170,1418,309,1527]" captionTargetId="figure-92@5.[170,1418,309,1527]" captionTargetPageId="5" captionText="FIGURES 5–7. Color pattern of Baetis majus Tiunova sp. nov., larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736178" httpUri="https://zenodo.org/record/4736178/files/figure.png" pageId="6" pageNumber="547">Figs 5, 7</figureCitation>
|
||
); posterior margins of tergum VI with regular row of quadrangular teeth of various sizes; surface of terga densely covered with numerous semilunar impressions, rare hair-like setae and conical spatulas (
|
||
<figureCitation id="132BB2AAFFACFF81E01EA243FE32BCCA" box="[344,427,367,394]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="6" pageNumber="547">Fig. 26</figureCitation>
|
||
). Sterna generally dirty white; sterna VII–VIII brownish (
|
||
<figureCitation id="132BB2AAFFACFF81E568A243FBEBBCCA" box="[1070,1138,367,394]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1550,1574]" captionTargetBox="[170,1418,309,1527]" captionTargetId="figure-92@5.[170,1418,309,1527]" captionTargetPageId="5" captionText="FIGURES 5–7. Color pattern of Baetis majus Tiunova sp. nov., larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736178" httpUri="https://zenodo.org/record/4736178/files/figure.png" pageId="6" pageNumber="547">Fig. 6</figureCitation>
|
||
). Six pairs of gills, present on abdominal segments II–VII; gills elongated, oval-shaped; all gills matt, with dark band surrounding margin and apparent tracheation; margins with numerous hair-like setae inserted at the base of small teeth (
|
||
<figureCitation id="132BB2AAFFACFF81E588A29BFACBBC91" box="[1230,1362,439,465]" captionStart="FIGURES 33–38" captionStartId="10.[151,264,1218,1242]" captionTargetBox="[162,1420,194,1185]" captionTargetId="figure-27@10.[151,1436,181,1193]" captionTargetPageId="10" captionText="FIGURES 33–38. Baetis majus Tiunova sp. nov., larvae, gills shape, dorsal view, roman numbers belong to the respective gill pairs." figureDoi="http://doi.org/10.5281/zenodo.4736190" httpUri="https://zenodo.org/record/4736190/files/figure.png" pageId="6" pageNumber="547">Figs 33–38</figureCitation>
|
||
). Gills II and VI almost equal in length and only slightly smaller than gill III; gills IV–V equal in length, 2 times longer than wide (
|
||
<figureCitation id="132BB2AAFFACFF81E054A2D3FE0CBF59" box="[274,405,510,537]" captionStart="FIGURES 33–38" captionStartId="10.[151,264,1218,1242]" captionTargetBox="[162,1420,194,1185]" captionTargetId="figure-27@10.[151,1436,181,1193]" captionTargetPageId="10" captionText="FIGURES 33–38. Baetis majus Tiunova sp. nov., larvae, gills shape, dorsal view, roman numbers belong to the respective gill pairs." figureDoi="http://doi.org/10.5281/zenodo.4736190" httpUri="https://zenodo.org/record/4736190/files/figure.png" pageId="6" pageNumber="547">Figs 35–36</figureCitation>
|
||
); gill VII the smallest,1.3 times shorter than gill II and 1.6 times shorter than gill V (
|
||
<figureCitation id="132BB2AAFFACFF81E47AA2D3FA09BF59" box="[1340,1424,511,537]" captionStart="FIGURES 33–38" captionStartId="10.[151,264,1218,1242]" captionTargetBox="[162,1420,194,1185]" captionTargetId="figure-27@10.[151,1436,181,1193]" captionTargetPageId="10" captionText="FIGURES 33–38. Baetis majus Tiunova sp. nov., larvae, gills shape, dorsal view, roman numbers belong to the respective gill pairs." figureDoi="http://doi.org/10.5281/zenodo.4736190" httpUri="https://zenodo.org/record/4736190/files/figure.png" pageId="6" pageNumber="547">Fig. 38</figureCitation>
|
||
). Paraproct with 9–10 large pointed teeth like spines of different size (
|
||
<figureCitation id="132BB2AAFFACFF81E2D1A10FFC72BF7D" box="[919,1003,546,573]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="6" pageNumber="547">Fig. 25</figureCitation>
|
||
); surface of paraproct without notched scales, with fine hair-like setae located disorderly and 3–5 rounded scales, scattered in middle area (
|
||
<figureCitation id="132BB2AAFFACFF81E5A3A16BFAA2BF21" box="[1253,1339,582,609]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="6" pageNumber="547">Figs 25</figureCitation>
|
||
,
|
||
<figureCitation id="132BB2AAFFACFF81E400A16BFA09BF21" box="[1350,1424,583,609]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1966,1990]" captionTargetBox="[151,1436,490,1943]" captionTargetId="figure-161@9.[151,1436,489,1943]" captionTargetPageId="9" captionText="FIGURES 27–32. Baetis majus Tiunova sp. nov., larvae, paratypes: 27, 29, inner margin of fore tibia; 28, inner margin of fore tarsus; 30, tarsal claw; 31, paraproct; 32, spatulas on paraproct. Scale bar: 27–31, 0.01 mm; 32, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.4736188" httpUri="https://zenodo.org/record/4736188/files/figure.png" pageId="6" pageNumber="547">31–32</figureCitation>
|
||
). Cerci brownish with dark band at the middle and dark tips (
|
||
<figureCitation id="132BB2AAFFACFF81E274A147FC01BFC5" box="[818,920,618,645]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1550,1574]" captionTargetBox="[170,1418,309,1527]" captionTargetId="figure-92@5.[170,1418,309,1527]" captionTargetPageId="5" captionText="FIGURES 5–7. Color pattern of Baetis majus Tiunova sp. nov., larvae, paratypes: 5, male, dorsal view; 6, ventral view; 7, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736178" httpUri="https://zenodo.org/record/4736178/files/figure.png" pageId="6" pageNumber="547">Figs 5–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6FFEA7FFACFF81E1D1A4A6FE4BBAAA" ID-DOI="http://doi.org/10.5281/zenodo.4736180" ID-Zenodo-Dep="4736180" httpUri="https://zenodo.org/record/4736180/files/figure.png" pageId="6" pageNumber="547" startId="6.[151,264,1930,1954]" targetBox="[151,1436,669,1907]" targetPageId="6">
|
||
<paragraph id="8BAFAE2FFFACFF81E1D1A4A6FE4BBAAA" blockId="6.[151,1436,1930,2027]" pageId="6" pageNumber="547">
|
||
<emphasis id="B964723DFFACFF81E1D1A4A6FEDEBAE2" bold="true" box="[151,327,1930,1954]" pageId="6" pageNumber="547">FIGURES 8–13.</emphasis>
|
||
<taxonomicName id="4C10D5ACFFACFF81E00BA4A7FDB1BAE2" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[333,552,1930,1954]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="547" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFACFF81E00BA4A7FE56BAE2" box="[333,463,1931,1954]" italics="true" pageId="6" pageNumber="547">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFACFF81E368A4A7FDE7BAE3" bold="true" box="[558,638,1931,1955]" pageId="6" pageNumber="547">
|
||
<taxonomicNameLabel id="A257CF46FFACFF81E368A4A7FDE7BAE3" box="[558,638,1931,1955]" pageId="6" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, larvae, dorsal view, paratypes: 8, scape; 9, surface of the head; 10, spatulas on head; 11, canines of left and right mandible; 12, glossae and paraglossae; 13, setae on paraglossa. Scale bar: 8, 11–12, 0.01 mm; 9, 0.02 mm; 10, 13, 0.002 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6FFEA7FFADFF80E1D1A4A7FB36BAAB" ID-DOI="http://doi.org/10.5281/zenodo.4736182" ID-Zenodo-Dep="4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="7" pageNumber="548" startId="7.[151,264,1931,1955]" targetBox="[156,1421,190,1907]" targetPageId="7">
|
||
<paragraph id="8BAFAE2FFFADFF80E1D1A4A7FB36BAAB" blockId="7.[151,1436,1931,2028]" pageId="7" pageNumber="548">
|
||
<emphasis id="B964723DFFADFF80E1D1A4A7FECDBAE3" bold="true" box="[151,340,1931,1955]" pageId="7" pageNumber="548">FIGURES 14–20.</emphasis>
|
||
<taxonomicName id="4C10D5ACFFADFF80E01FA4A0FDADBAE3" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[345,564,1931,1955]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="548" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFADFF80E01FA4A0FE42BAE3" box="[345,475,1932,1955]" italics="true" pageId="7" pageNumber="548">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFADFF80E37CA4A0FD13BAE4" bold="true" box="[570,650,1932,1956]" pageId="7" pageNumber="548">
|
||
<taxonomicNameLabel id="A257CF46FFADFF80E37CA4A0FD13BAE4" box="[570,650,1932,1956]" pageId="7" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6FFEA7FFA2FF8FE1D1A5CDFE9FBA01" ID-DOI="http://doi.org/10.5281/zenodo.4736186" ID-Zenodo-Dep="4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="8" pageNumber="549" startId="8.[151,264,1761,1785]" targetBox="[160,1422,198,1719]" targetPageId="8">
|
||
<paragraph id="8BAFAE2FFFA2FF8FE1D1A5CDFE9FBA01" blockId="8.[151,1436,1761,1858]" pageId="8" pageNumber="549">
|
||
<emphasis id="B964723DFFA2FF8FE1D1A5CDFECABBB9" bold="true" box="[151,339,1761,1785]" pageId="8" pageNumber="549">FIGURES 21–26.</emphasis>
|
||
<taxonomicName id="4C10D5ACFFA2FF8FE01FA5CEFDAABBB9" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[345,563,1761,1785]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="549" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA2FF8FE01FA5CEFE42BBB9" box="[345,475,1762,1785]" italics="true" pageId="8" pageNumber="549">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFA2FF8FE37EA5CEFD11BBBA" bold="true" box="[568,648,1762,1786]" pageId="8" pageNumber="549">
|
||
<taxonomicNameLabel id="A257CF46FFA2FF8FE37EA5CEFD11BBBA" box="[568,648,1762,1786]" pageId="8" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAFAE2FFFA2FF8FE181A45CFCDCBA92" blockId="8.[151,1437,1904,2038]" pageId="8" pageNumber="549">
|
||
<emphasis id="B964723DFFA2FF8FE181A45CFE98BACA" bold="true" box="[199,257,1904,1930]" pageId="8" pageNumber="549">Eggs</emphasis>
|
||
. General form oval 102–106 μm length and 62–68 μm width (
|
||
<figureCitation id="132BB2AAFFA2FF8FE28EA45CFB87BACA" box="[968,1054,1904,1930]" captionStart="FIGURES 39–41" captionStartId="11.[151,264,1992,2016]" captionTargetBox="[152,1436,489,1969]" captionTargetId="figure-163@11.[151,1436,489,1969]" captionTargetPageId="11" captionText="FIGURES 39–41. Baetis majus Tiunova sp. nov., egg: 39, general view; 40, structure of chorion; 41, micropyle." figureDoi="http://doi.org/10.5281/zenodo.4736192" httpUri="https://zenodo.org/record/4736192/files/figure.png" pageId="8" pageNumber="549">Fig. 39</figureCitation>
|
||
). Chorion corrugated and densely wrinkled, with numerous small granules and with small fossae (
|
||
<figureCitation id="132BB2AAFFA2FF8FE22BA4B8FC5ABAEE" box="[877,963,1940,1966]" captionStart="FIGURES 39–41" captionStartId="11.[151,264,1992,2016]" captionTargetBox="[152,1436,489,1969]" captionTargetId="figure-163@11.[151,1436,489,1969]" captionTargetPageId="11" captionText="FIGURES 39–41. Baetis majus Tiunova sp. nov., egg: 39, general view; 40, structure of chorion; 41, micropyle." figureDoi="http://doi.org/10.5281/zenodo.4736192" httpUri="https://zenodo.org/record/4736192/files/figure.png" pageId="8" pageNumber="549">Fig. 40</figureCitation>
|
||
). One or two small and round micropyles with a collar-like ring located in the equatorial area (
|
||
<figureCitation id="132BB2AAFFA2FF8FE3A2A494FCAEBA92" box="[740,823,1976,2002]" captionStart="FIGURES 39–41" captionStartId="11.[151,264,1992,2016]" captionTargetBox="[152,1436,489,1969]" captionTargetId="figure-163@11.[151,1436,489,1969]" captionTargetPageId="11" captionText="FIGURES 39–41. Baetis majus Tiunova sp. nov., egg: 39, general view; 40, structure of chorion; 41, micropyle." figureDoi="http://doi.org/10.5281/zenodo.4736192" httpUri="https://zenodo.org/record/4736192/files/figure.png" pageId="8" pageNumber="549">Fig. 41</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30AFDA4FFA2FF8EE181A4F7FF46BC91" lastPageId="9" lastPageNumber="550" pageId="8" pageNumber="549" type="diagnosis">
|
||
<paragraph id="8BAFAE2FFFA2FF8EE181A4F7FF46BC91" blockId="8.[151,1437,1904,2038]" lastBlockId="9.[151,1437,151,465]" lastPageId="9" lastPageNumber="550" pageId="8" pageNumber="549">
|
||
<emphasis id="B964723DFFA2FF8FE181A4F7FEDBBAB5" bold="true" box="[199,322,2011,2037]" pageId="8" pageNumber="549">Diagnosis.</emphasis>
|
||
Larva. Six pairs of gills, present on abdominal segments II–VII; dorsal surface of the head, femora, tibia and tarsi covered with numerous small bluntly rounded spatulas and fine hairs (
|
||
<figureCitation id="132BB2AAFFA3FF8EE524A3BBFB4EBDF1" box="[1122,1239,151,177]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1930,1954]" captionTargetBox="[151,1436,669,1907]" captionTargetId="figure-277@6.[151,1436,669,1907]" captionTargetPageId="6" captionText="FIGURES 8–13. Baetis majus Tiunova sp. nov., larvae, dorsal view, paratypes: 8, scape; 9, surface of the head; 10, spatulas on head; 11, canines of left and right mandible; 12, glossae and paraglossae; 13, setae on paraglossa. Scale bar: 8, 11–12, 0.01 mm; 9, 0.02 mm; 10, 13, 0.002 mm." figureDoi="http://doi.org/10.5281/zenodo.4736180" httpUri="https://zenodo.org/record/4736180/files/figure.png" pageId="9" pageNumber="550">Figs 9–10</figureCitation>
|
||
,
|
||
<figureCitation id="132BB2AAFFA3FF8EE5A2A3BBFAB7BDF1" box="[1252,1326,151,177]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1966,1990]" captionTargetBox="[151,1436,490,1943]" captionTargetId="figure-161@9.[151,1436,489,1943]" captionTargetPageId="9" captionText="FIGURES 27–32. Baetis majus Tiunova sp. nov., larvae, paratypes: 27, 29, inner margin of fore tibia; 28, inner margin of fore tarsus; 30, tarsal claw; 31, paraproct; 32, spatulas on paraproct. Scale bar: 27–31, 0.01 mm; 32, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.4736188" httpUri="https://zenodo.org/record/4736188/files/figure.png" pageId="9" pageNumber="550">27–29</figureCitation>
|
||
); the first tree teeth of kinetodontium and incisor of right mandible of relatively equal size (
|
||
<figureCitation id="132BB2AAFFA3FF8EE575A397FB1EBD95" box="[1075,1159,186,213]" captionStart="FIGURES 14–20" captionStartId="7.[151,264,1931,1955]" captionTargetBox="[156,1421,190,1907]" captionTargetId="figure-17@7.[151,1436,181,1907]" captionTargetPageId="7" captionText="FIGURES 14–20. Baetis majus Tiunova sp. nov., larvae, details of mouthparts, paratypes: 14, labrum, dorsal view; 15, canines and prostheca of right mandible, dorsal view; 16, canines and prostheca of left mandible, dorsal view; 17, maxillary palp, dorsal view; 18, glossa and paraglossa, ventral view; 19, labial palp, ventral view; 20, labial palp, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4736182" httpUri="https://zenodo.org/record/4736182/files/figure.png" pageId="9" pageNumber="550">Fig. 15</figureCitation>
|
||
); third segment of labial palp wide, its apex truncated; 1.3 times wider than the second segment; second segment of labial palp with rounded apicomedial projection (
|
||
<figureCitation id="132BB2AAFFA3FF8EE0EDA22FFD93BC5D" box="[427,522,258,285]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="9" pageNumber="550">Figs. 25</figureCitation>
|
||
,
|
||
<figureCitation id="132BB2AAFFA3FF8EE35EA22FFDFBBC5D" box="[536,610,259,285]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1966,1990]" captionTargetBox="[151,1436,490,1943]" captionTargetId="figure-161@9.[151,1436,489,1943]" captionTargetPageId="9" captionText="FIGURES 27–32. Baetis majus Tiunova sp. nov., larvae, paratypes: 27, 29, inner margin of fore tibia; 28, inner margin of fore tarsus; 30, tarsal claw; 31, paraproct; 32, spatulas on paraproct. Scale bar: 27–31, 0.01 mm; 32, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.4736188" httpUri="https://zenodo.org/record/4736188/files/figure.png" pageId="9" pageNumber="550">31–32</figureCitation>
|
||
); posterior margins of tergum VI with regular row of quadrangular teeth of various sizes (
|
||
<figureCitation id="132BB2AAFFA3FF8EE01FA20BFE37BC02" box="[345,430,295,322]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1761,1785]" captionTargetBox="[160,1422,198,1719]" captionTargetId="figure-27@8.[151,1436,181,1737]" captionTargetPageId="8" captionText="FIGURES 21–26. Baetis majus Tiunova sp. nov., larvae, paratypes, dorsal view: 21, foreleg, 22, shape of long bristles on distal part of femora; 23, shape of short setae on inner margin femora; 24, tarsus claw; 25, paraproct plate; 26, posterior margin of tergum VI." figureDoi="http://doi.org/10.5281/zenodo.4736186" httpUri="https://zenodo.org/record/4736186/files/figure.png" pageId="9" pageNumber="550">Fig. 26</figureCitation>
|
||
); paraproct without notched spatulas, with fine hairs located disorderly and 3–5 rounded spatulas, scattered in middle area. Imago: hind wings with two simple longitudinal veins (
|
||
<figureCitation id="132BB2AAFFA3FF8EE5D2A267FB41BC25" box="[1172,1240,331,357]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1968,1992]" captionTargetBox="[171,1416,189,1933]" captionTargetId="figure-27@4.[160,1427,181,1944]" captionTargetPageId="4" captionText="FIGURES 3–4. Baetis majus Tiunova sp. nov., male imagoes, holotype: 3, hind wing; 4, unistyliger and forceps." figureDoi="http://doi.org/10.5281/zenodo.4736176" httpUri="https://zenodo.org/record/4736176/files/figure.png" pageId="9" pageNumber="550">Fig. 3</figureCitation>
|
||
); unistiliger with parallel margins, slightly elongated, with a distinct small projection at the apex of the inner edge (
|
||
<figureCitation id="132BB2AAFFA3FF8EE5A2A243FAB0BCC9" box="[1252,1321,367,393]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1968,1992]" captionTargetBox="[171,1416,189,1933]" captionTargetId="figure-27@4.[160,1427,181,1944]" captionTargetPageId="4" captionText="FIGURES 3–4. Baetis majus Tiunova sp. nov., male imagoes, holotype: 3, hind wing; 4, unistyliger and forceps." figureDoi="http://doi.org/10.5281/zenodo.4736176" httpUri="https://zenodo.org/record/4736176/files/figure.png" pageId="9" pageNumber="550">Fig. 4</figureCitation>
|
||
); segment III of forceps small, well defined, strongly expanded to the apex, its length only slightly exceeds its maximum width.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6FFEA7FFA3FF8EE1D1A482FB56BAAA" ID-DOI="http://doi.org/10.5281/zenodo.4736188" ID-Zenodo-Dep="4736188" httpUri="https://zenodo.org/record/4736188/files/figure.png" pageId="9" pageNumber="550" startId="9.[151,264,1966,1990]" targetBox="[151,1436,490,1943]" targetPageId="9">
|
||
<paragraph id="8BAFAE2FFFA3FF8EE1D1A482FB56BAAA" blockId="9.[151,1437,1966,2026]" pageId="9" pageNumber="550">
|
||
<emphasis id="B964723DFFA3FF8EE1D1A482FED4BA86" bold="true" box="[151,333,1966,1990]" pageId="9" pageNumber="550">FIGURES 27–32</emphasis>
|
||
.
|
||
<taxonomicName id="4C10D5ACFFA3FF8EE01CA483FDAFBA86" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[346,566,1966,1990]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="550" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA3FF8EE01CA483FE44BA86" box="[346,477,1967,1990]" italics="true" pageId="9" pageNumber="550">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFA3FF8EE37AA482FD15BA86" bold="true" box="[572,652,1966,1990]" pageId="9" pageNumber="550">
|
||
<taxonomicNameLabel id="A257CF46FFA3FF8EE37AA482FD15BA86" box="[572,652,1966,1990]" pageId="9" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, larvae, paratypes: 27, 29, inner margin of fore tibia; 28, inner margin of fore tarsus; 30, tarsal claw; 31, paraproct; 32, spatulas on paraproct. Scale bar: 27–31, 0.01 mm; 32, 0.001 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6FFEA7FFA0FF8DE1D1A7EEFF49B9BE" ID-DOI="http://doi.org/10.5281/zenodo.4736190" ID-Zenodo-Dep="4736190" httpUri="https://zenodo.org/record/4736190/files/figure.png" pageId="10" pageNumber="551" startId="10.[151,264,1218,1242]" targetBox="[162,1420,194,1185]" targetPageId="10">
|
||
<paragraph id="8BAFAE2FFFA0FF8DE1D1A7EEFF49B9BE" blockId="10.[151,1436,1218,1278]" pageId="10" pageNumber="551">
|
||
<emphasis id="B964723DFFA0FF8DE1D1A7EEFECDB99A" bold="true" box="[151,340,1218,1242]" pageId="10" pageNumber="551">FIGURES 33–38.</emphasis>
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE01CA7EFFDAFB99A" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[346,566,1218,1242]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA0FF8DE01CA7EFFE44B99A" box="[346,477,1219,1242]" italics="true" pageId="10" pageNumber="551">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFA0FF8DE37AA7EEFD14B99A" bold="true" box="[572,653,1218,1242]" pageId="10" pageNumber="551">
|
||
<taxonomicNameLabel id="A257CF46FFA0FF8DE37AA7EEFD14B99A" box="[572,653,1218,1242]" pageId="10" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, larvae, gills shape, dorsal view, roman numbers belong to the respective gill pairs.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30AFDA4FFA0FF8DE181A600FCCBB8F3" pageId="10" pageNumber="551" type="distribution">
|
||
<paragraph id="8BAFAE2FFFA0FF8DE181A600FCCBB8F3" blockId="10.[151,1437,1324,2035]" pageId="10" pageNumber="551">
|
||
<emphasis id="B964723DFFA0FF8DE181A600FEF8B806" bold="true" box="[199,353,1324,1350]" pageId="10" pageNumber="551">Distribution.</emphasis>
|
||
Russian Far East.
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE373A601FCB3B806" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[565,810,1324,1350]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA0FF8DE373A601FD5FB806" box="[565,710,1325,1350]" italics="true" pageId="10" pageNumber="551">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257CF46FFA0FF8DE274A600FC17B807" box="[818,910,1324,1351]" pageId="10" pageNumber="551" rank="species">
|
||
<emphasis id="B964723DFFA0FF8DE274A600FC1EB806" bold="true" box="[818,903,1324,1350]" pageId="10" pageNumber="551">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is known from its
|
||
<typeStatus id="54AB108DFFA0FF8DE523A601FB0CB807" box="[1125,1173,1325,1351]" pageId="10" pageNumber="551">type</typeStatus>
|
||
locality only (
|
||
<figureCitation id="132BB2AAFFA0FF8DE47DA600FA09B806" box="[1339,1424,1324,1350]" captionStart="FIGURE 42" captionStartId="12.[151,250,1130,1154]" captionTargetBox="[151,1436,205,1107]" captionTargetId="figure-27@12.[151,1436,179,1107]" captionTargetPageId="12" captionText="FIGURE 42. Locality of Baetis majus Tiunova sp. nov., Shkotovka River (Primorskiy Kray)." figureDoi="http://doi.org/10.5281/zenodo.4736194" httpUri="https://zenodo.org/record/4736194/files/figure.png" pageId="10" pageNumber="551">Fig. 42</figureCitation>
|
||
). The larvae of the new species were found in the lower part of the river on swift riffles with cobble and gravel substrate. No larvae were found at other sites upstream of the river. Water temperature was 12–15°C, depth
|
||
<quantity id="4CE803CAFFA0FF8DE410A658FF24B8F3" metricMagnitude="-1" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="10" pageNumber="551" unit="cm" value="40.0" valueMax="60.0" valueMin="20.0">20–60 cm</quantity>
|
||
. According to our data, the adult emergence period is May.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30AFDA4FFA0FF8DE181A690FE77B8BB" pageId="10" pageNumber="551" type="etymology">
|
||
<paragraph id="8BAFAE2FFFA0FF8DE181A690FE77B8BB" blockId="10.[151,1437,1324,2035]" pageId="10" pageNumber="551">
|
||
<emphasis id="B964723DFFA0FF8DE181A690FED1B896" bold="true" box="[199,328,1468,1494]" pageId="10" pageNumber="551">Etymology</emphasis>
|
||
. The species name is derived from the Latin word
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE2E6A691FC7AB896" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[928,995,1469,1494]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="majus">
|
||
<emphasis id="B964723DFFA0FF8DE2E6A691FC7AB896" box="[928,995,1469,1494]" italics="true" pageId="10" pageNumber="551">majus</emphasis>
|
||
</taxonomicName>
|
||
, which means May. All the examined material was collected in May.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30AFDA4FFA0FF8CE181A528FD85BC91" lastPageId="11" lastPageNumber="552" pageId="10" pageNumber="551" type="description">
|
||
<paragraph id="8BAFAE2FFFA0FF8DE181A528FAB4BBEF" blockId="10.[151,1437,1324,2035]" pageId="10" pageNumber="551">
|
||
<emphasis id="B964723DFFA0FF8DE181A528FD9EBB5E" bold="true" box="[199,519,1540,1566]" pageId="10" pageNumber="551">Results of DNA barcoding.</emphasis>
|
||
We obtained partial COI sequences (657 bp) of
|
||
<specimenCount id="9D1665A6FFA0FF8DE561A529FB58BB5E" box="[1063,1217,1540,1567]" pageId="10" pageNumber="551" type="generic">one specimen</specimenCount>
|
||
(BM175) of
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE410A529FEA6BB02" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA0FF8DE410A529FF43BB02" italics="true" pageId="10" pageNumber="551">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFA0FF8DE00EA504FE32BB02" bold="true" box="[328,427,1576,1602]" pageId="10" pageNumber="551">
|
||
<taxonomicNameLabel id="A257CF46FFA0FF8DE00EA504FE3CBB02" box="[328,421,1576,1602]" pageId="10" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
<specimenCount id="9D1665A6FFA0FF8DE0F2A504FDE4BB02" box="[436,637,1576,1602]" pageId="10" pageNumber="551" type="generic">twelve specimens</specimenCount>
|
||
(BP30, BP34, BP35, BP178, BP179, BP190, BP358, BP387, BP388, BP672, BP674 and BP794) of
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE344A561FD7ABB26" authorityName="Kluge" authorityYear="1983" box="[514,739,1612,1638]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="pseudothermicus">
|
||
<emphasis id="B964723DFFA0FF8DE344A561FD7ABB26" box="[514,739,1612,1638]" italics="true" pageId="10" pageNumber="551">B. pseudothermicus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<specimenCount id="9D1665A6FFA0FF8DE3B3A560FC3ABB26" box="[757,931,1612,1638]" pageId="10" pageNumber="551" type="generic">four specimens</specimenCount>
|
||
(BU29, BU177, BU542 and BU552) of
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE4C2A561FF70BBCA" authorityName="Kazlauskas" authorityYear="1963" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="ursinus">
|
||
<emphasis id="B964723DFFA0FF8DE4C2A561FF70BBCA" italics="true" pageId="10" pageNumber="551">B. ursinus</emphasis>
|
||
</taxonomicName>
|
||
; and
|
||
<specimenCount id="9D1665A6FFA0FF8DE061A55DFE55BBCA" box="[295,460,1648,1675]" pageId="10" pageNumber="551" type="generic">two specimens</specimenCount>
|
||
(BU543 and BU556) of
|
||
<emphasis id="B964723DFFA0FF8DE3A6A55DFCF6BBCA" box="[736,879,1649,1674]" italics="true" pageId="10" pageNumber="551">
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE3A6A55DFCF2BBCA" authorityName="Kluge" authorityYear="1983" box="[736,875,1649,1674]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="ussuricus">B. ussuricus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The mean nucleotide base compositions of
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE410A55DFEA4BBEE" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA0FF8DE410A55DFF43BBEE" italics="true" pageId="10" pageNumber="551">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFA0FF8DE002A5B8FE04BBEE" bold="true" box="[324,413,1684,1710]" pageId="10" pageNumber="551">
|
||
<taxonomicNameLabel id="A257CF46FFA0FF8DE002A5B8FE04BBEE" box="[324,413,1684,1710]" pageId="10" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were A 23.6%, C 18.4%, G 21.8% and T 36.2% across all sites for all sequences.
|
||
</paragraph>
|
||
<paragraph id="8BAFAE2FFFA0FF8DE181A595FD9FBAEB" blockId="10.[151,1437,1324,2035]" pageId="10" pageNumber="551">
|
||
We used nucleotide BLAST algorithm in NCBI GenBank (nucleotide collection database) and animal identification in BOLD systems to find the nearest neighbours to
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE2E5A5F1FB06BBB6" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[931,1183,1756,1782]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA0FF8DE2E5A5F1FBA1BBB6" box="[931,1080,1757,1782]" italics="true" pageId="10" pageNumber="551">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257CF46FFA0FF8DE5EDA5F0FA92BBB7" box="[1195,1291,1756,1783]" pageId="10" pageNumber="551" rank="species">
|
||
<emphasis id="B964723DFFA0FF8DE5EDA5F0FA9DBBB6" bold="true" box="[1195,1284,1756,1782]" pageId="10" pageNumber="551">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
A search in GenBank showed that the nearest neighbours have a similarity less than 85% and relate to unidentified
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE410A42DFF71BA7E" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B964723DFFA0FF8DE410A42DFA05BA5A" box="[1366,1436,1793,1818]" italics="true" pageId="10" pageNumber="551">Baetis</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(
|
||
<accessionNumber id="944333CCFFA0FF8DE1BEA408FEECBA7E" box="[248,373,1828,1854]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KP970694" pageId="10" pageNumber="551">KP970694</accessionNumber>
|
||
,
|
||
<accessionNumber id="944333CCFFA0FF8DE0C6A408FE63BA7F" box="[384,506,1828,1855]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KP970695" pageId="10" pageNumber="551">KP970695</accessionNumber>
|
||
) and several species of genus
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE215A408FC5EBA7E" authorityName="Bengtsson" authorityYear="1912" box="[851,967,1828,1854]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964723DFFA0FF8DE215A408FC5EBA7E" box="[851,967,1828,1854]" italics="true" pageId="10" pageNumber="551">Acentrella</emphasis>
|
||
</taxonomicName>
|
||
. Highly similar (98.75–100%) sequences of unidentified
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE00CA465FE75BA22" box="[330,492,1864,1890]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B964723DFFA0FF8DE00CA465FE09BA22" box="[330,400,1865,1890]" italics="true" pageId="10" pageNumber="551">Baetis</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F307EEBFFFA0FF8DE370A464FDE0BA22" box="[566,633,1864,1890]" name="China" pageId="10" pageNumber="551">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49D460CDFFA0FF8DE3CDA464FD46BA22" box="[651,735,1864,1890]" country="China" name="Beijing" pageId="10" pageNumber="551">Beijing</collectingRegion>
|
||
and
|
||
<collectingRegion id="49D460CDFFA0FF8DE25AA464FCC6BA22" box="[796,863,1864,1890]" country="China" name="Hebei" pageId="10" pageNumber="551">Hebei</collectingRegion>
|
||
provinces) were found in the BOLD system, but these data was unavailable and set to private status. Probably,
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE20EA441FBA3BAC6" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[840,1082,1900,1926]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA0FF8DE20EA441FC41BAC6" box="[840,984,1901,1926]" italics="true" pageId="10" pageNumber="551">Baetis majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257CF46FFA0FF8DE506A440FB02BAC7" box="[1088,1179,1900,1927]" pageId="10" pageNumber="551" rank="species">
|
||
<emphasis id="B964723DFFA0FF8DE506A440FB0DBAC6" bold="true" box="[1088,1172,1900,1926]" pageId="10" pageNumber="551">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
inhabits also
|
||
<collectingCountry id="F307EEBFFFA0FF8DE472A440FAEEBAC6" box="[1332,1399,1900,1926]" name="China" pageId="10" pageNumber="551">China</collectingCountry>
|
||
(in provinces of
|
||
<collectingRegion id="49D460CDFFA0FF8DE06FA4BCFEE2BAEA" box="[297,379,1936,1962]" country="China" name="Beijing" pageId="10" pageNumber="551">Beijing</collectingRegion>
|
||
and
|
||
<collectingRegion id="49D460CDFFA0FF8DE0F4A4BCFE63BAEB" box="[434,506,1936,1963]" country="China" name="Hebei" pageId="10" pageNumber="551">Hebei</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAFAE2FFFA0FF8CE181A498FD9FBC5D" blockId="10.[151,1437,1324,2035]" lastBlockId="11.[151,1437,151,465]" lastPageId="11" lastPageNumber="552" pageId="10" pageNumber="551">
|
||
Bayesian phylogeny revealed two well-supported sister clades, one of which includes
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE5D2A499FA0CBA8E" authority="Lehmkuhl" authorityName="Lehmkuhl" box="[1172,1429,1972,1998]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="bundyiae">
|
||
<emphasis id="B964723DFFA0FF8DE5D2A499FA82BA8E" box="[1172,1307,1972,1998]" italics="true" pageId="10" pageNumber="551">B. bundyiae</emphasis>
|
||
Lehmkuhl
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE1D1A4F5FEFFBAB2" authority="Ide" authorityName="Ide" authorityYear="1937" box="[151,358,2008,2034]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="hudsonicus">
|
||
<emphasis id="B964723DFFA0FF8DE1D1A4F5FEAEBAB2" box="[151,311,2008,2034]" italics="true" pageId="10" pageNumber="551">B. hudsonicus</emphasis>
|
||
Ide
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE031A4F5FDF8BAB2" authority="Kimmins" authorityName="Kimmins" authorityYear="1957" box="[375,609,2008,2034]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="macani">
|
||
<emphasis id="B964723DFFA0FF8DE031A4F5FE72BAB2" box="[375,491,2009,2034]" italics="true" pageId="10" pageNumber="551">B. macani</emphasis>
|
||
Kimmins
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE329A4F5FC14BAB3" authority="Bengtsson" authorityName="Bengtsson" authorityYear="1917" box="[623,909,2008,2035]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="subalpinus">
|
||
<emphasis id="B964723DFFA0FF8DE329A4F5FC93BAB2" box="[623,778,2008,2034]" italics="true" pageId="10" pageNumber="551">B. subalpinus</emphasis>
|
||
Bengtsson
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C10D5ACFFA0FF8DE2DDA4F5FB69BAB2" authority="McDunnough" authorityName="McDunnough" authorityYear="1925" box="[923,1264,2008,2034]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="brunneicolor">
|
||
<emphasis id="B964723DFFA0FF8DE2DDA4F5FBD1BAB2" box="[923,1096,2008,2034]" italics="true" pageId="10" pageNumber="551">B brunneicolor</emphasis>
|
||
McDunnough
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C10D5ACFFA0FF8CE5B9A4F5FEB0BDF1" authority="Keffermuller" authorityName="Keffermuller" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" lastPageId="11" lastPageNumber="552" order="Ephemeroptera" pageId="10" pageNumber="551" phylum="Arthropoda" rank="species" species="liebenauae">
|
||
<emphasis id="B964723DFFA0FF8DE5B9A4F5FA05BAB2" box="[1279,1436,2008,2034]" italics="true" pageId="10" pageNumber="551">B. liebenauae</emphasis>
|
||
Keffermüller
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE07CA3BBFE3EBDF0" authorityName="Curtis" authorityYear="1834" box="[314,423,151,176]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="vernus">
|
||
<emphasis id="B964723DFFA1FF8CE07CA3BBFE3EBDF0" box="[314,423,151,176]" italics="true" pageId="11" pageNumber="552">B. vernus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE0FEA3BBFDB5BDF0" authorityName="Kazlauskas" authorityYear="1963" box="[440,556,151,176]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="ursinus">
|
||
<emphasis id="B964723DFFA1FF8CE0FEA3BBFDB5BDF0" box="[440,556,151,176]" italics="true" pageId="11" pageNumber="552">B. ursinus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B964723DFFA1FF8CE378A3BBFD44BDF1" box="[574,733,151,177]" italics="true" pageId="11" pageNumber="552">
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE378A3BBFD40BDF1" box="[574,729,151,177]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="maculosus">B. maculosus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE3A1A3BBFC2BBDF1" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[743,946,151,177]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA1FF8CE3A1A3BBFCD5BDF0" box="[743,844,151,176]" italics="true" pageId="11" pageNumber="552">B. majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFA1FF8CE2FAA3BBFBB9BDF1" bold="true" box="[956,1056,151,177]" pageId="11" pageNumber="552">
|
||
<taxonomicNameLabel id="A257CF46FFA1FF8CE2FAA3BBFB83BDF1" box="[956,1050,151,177]" pageId="11" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE51AA3BBFE84BD95" authority="Selvakumar et al." authorityName="Selvakumar" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="michaelohubbardi">
|
||
<emphasis id="B964723DFFA1FF8CE51AA3BBFAD3BDF1" box="[1116,1354,151,177]" italics="true" pageId="11" pageNumber="552">B. michaelohubbardi</emphasis>
|
||
Selvakumar
|
||
<emphasis id="B964723DFFA1FF8CE1A3A397FE8FBD95" box="[229,278,187,213]" italics="true" pageId="11" pageNumber="552">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
; the second clade includes the remaining species.
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE209A397FB8ABD95" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[847,1043,187,213]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA1FF8CE209A397FC29BD94" box="[847,944,187,212]" italics="true" pageId="11" pageNumber="552">B. majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFA1FF8CE55CA397FBEABD95" bold="true" box="[1050,1139,187,213]" pageId="11" pageNumber="552">
|
||
<taxonomicNameLabel id="A257CF46FFA1FF8CE55CA397FBEABD95" box="[1050,1139,187,213]" pageId="11" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was the earliest branching lineage in the clade and includes
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE356A3F3FD3FBDB9" box="[528,678,223,249]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="maculosus">
|
||
<emphasis id="B964723DFFA1FF8CE356A3F3FD3FBDB9" box="[528,678,223,249]" italics="true" pageId="11" pageNumber="552">B. maculosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B964723DFFA1FF8CE398A3F3FCC0BDB8" box="[734,857,223,248]" italics="true" pageId="11" pageNumber="552">
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE398A3F3FCCABDB8" authorityName="Kazlauskas" authorityYear="1963" box="[734,851,223,248]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="ursinus">B. ursinus</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
however, support for this clade was low (Bayesian posterior probability, PP = 0.52).
|
||
</paragraph>
|
||
<paragraph id="8BAFAE2FFFA1FF8CE181A20BFD85BC91" blockId="11.[151,1437,151,465]" pageId="11" pageNumber="552">
|
||
Morphological and molecular data were used to construct a Bayesian tree, which revealed a polytomy node including
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE041A267FE53BC25" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[263,458,331,357]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA1FF8CE041A267FEF1BC24" box="[263,360,331,356]" italics="true" pageId="11" pageNumber="552">B. majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257CF46FFA1FF8CE096A267FDB3BC25" box="[464,554,331,357]" pageId="11" pageNumber="552" rank="species">
|
||
<emphasis id="B964723DFFA1FF8CE096A267FDBDBC25" bold="true" box="[464,548,331,357]" pageId="11" pageNumber="552">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
as the first lineage;
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE24EA267FC31BC25" authorityName="Ikonomov" authorityYear="1962" box="[776,936,331,357]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="vardarensis">
|
||
<emphasis id="B964723DFFA1FF8CE24EA267FC31BC25" box="[776,936,331,357]" italics="true" pageId="11" pageNumber="552">B. vardarensis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE282A267FBB4BC25" authorityName="Muller-Liebenau" authorityYear="1967" box="[964,1069,331,357]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="lutheri">
|
||
<emphasis id="B964723DFFA1FF8CE282A267FBB4BC25" box="[964,1069,331,357]" italics="true" pageId="11" pageNumber="552">B. lutheri</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE50FA267FB6FBC25" authorityName="Tiunova & Semenchenko" authorityYear="2019" box="[1097,1270,331,357]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="pentaphyllus">
|
||
<emphasis id="B964723DFFA1FF8CE50FA267FB6FBC25" box="[1097,1270,331,357]" italics="true" pageId="11" pageNumber="552">B. pentaphyllus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE443A267FA04BC25" authorityName="Muller-Liebenau" authorityYear="1967" box="[1285,1437,331,357]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="lutheri">
|
||
<emphasis id="B964723DFFA1FF8CE443A267FA04BC25" box="[1285,1437,331,357]" italics="true" pageId="11" pageNumber="552">Baetis lutheri</emphasis>
|
||
</taxonomicName>
|
||
species-group) as the second lineage; and
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE33EA243FD62BCC9" authorityName="Zimmermann" authorityYear="1980" box="[632,763,367,393]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="braaschi">
|
||
<emphasis id="B964723DFFA1FF8CE33EA243FD62BCC9" box="[632,763,367,393]" italics="true" pageId="11" pageNumber="552">B. braaschi</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE25AA243FCFCBCC9" authorityName="Jacob & Zimmermann" authorityYear="1978" box="[796,869,367,393]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="ilex">
|
||
<emphasis id="B964723DFFA1FF8CE25AA243FCFCBCC9" box="[796,869,367,393]" italics="true" pageId="11" pageNumber="552">B. ilex</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE2C1A243FB80BCC9" authorityName="Thomas" authorityYear="1999" box="[903,1049,367,393]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="gadeai">
|
||
<emphasis id="B964723DFFA1FF8CE2C1A243FC06BCC8" box="[903,927,367,392]" italics="true" pageId="11" pageNumber="552">B.</emphasis>
|
||
cf.
|
||
<emphasis id="B964723DFFA1FF8CE28BA243FB80BCC9" box="[973,1049,367,393]" italics="true" pageId="11" pageNumber="552">gadeai</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE57CA243FB2ABCC9" baseAuthorityName="Pictet" baseAuthorityYear="1843" box="[1082,1203,367,393]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="rhodani">
|
||
<emphasis id="B964723DFFA1FF8CE57CA243FB2ABCC9" box="[1082,1203,367,393]" italics="true" pageId="11" pageNumber="552">B. rhodani</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE592A243FADDBCC9" authorityName="Soldan" authorityYear="1977" box="[1236,1348,367,393]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="baksan">
|
||
<emphasis id="B964723DFFA1FF8CE592A243FADDBCC9" box="[1236,1348,367,393]" italics="true" pageId="11" pageNumber="552">B. baksan</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE410A243FF76BCED" baseAuthorityName="Pictet" baseAuthorityYear="1843" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="rhodani">
|
||
<emphasis id="B964723DFFA1FF8CE410A243FF76BCED" italics="true" pageId="11" pageNumber="552">Baetis rhodani</emphasis>
|
||
</taxonomicName>
|
||
species-group) as the third lineage. Thus, the species-group to which
|
||
<taxonomicName id="4C10D5ACFFA1FF8CE556A2BFFB41BCED" authority="Tiunova" authorityName="Tiunova & Semenchenko & Tong" authorityYear="2021" box="[1040,1240,403,429]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="552" phylum="Arthropoda" rank="species" species="majus" status="sp. nov.">
|
||
<emphasis id="B964723DFFA1FF8CE556A2BFFBEDBCEC" box="[1040,1140,403,428]" italics="true" pageId="11" pageNumber="552">B. majus</emphasis>
|
||
Tiunova
|
||
</taxonomicName>
|
||
<emphasis id="B964723DFFA1FF8CE5A7A2BFFAA5BCED" bold="true" box="[1249,1340,403,429]" pageId="11" pageNumber="552">
|
||
<taxonomicNameLabel id="A257CF46FFA1FF8CE5A7A2BFFAA5BCED" box="[1249,1340,403,429]" pageId="11" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
belongs remains unidentified (
|
||
<figureCitation id="132BB2AAFFA1FF8CE0CBA29BFE7DBC91" box="[397,484,439,465]" captionStart="FIGURE 43" captionStartId="13.[151,250,1057,1081]" captionTargetBox="[165,1419,174,1029]" captionTargetId="figure-17@13.[164,1423,166,1033]" captionTargetPageId="13" captionText="FIGURE 43. Bayesian tree based on mitochondrion COI gene for subgenera Baetis and Tenuibaetis form Eastern and Western Palearctic, Nearctic and Oriental regions. Specimens obtained in this study are in bold. Bayesian posterior probabilities (PP) above 0.7 are given above tree nodes." figureDoi="http://doi.org/10.5281/zenodo.4736196" httpUri="https://zenodo.org/record/4736196/files/figure.png" pageId="11" pageNumber="552">Figs 43</figureCitation>
|
||
,
|
||
<figureCitation id="132BB2AAFFA1FF8CE0B6A29BFD89BC91" box="[496,528,439,465]" captionStart="FIGURE 44" captionStartId="13.[151,250,1904,1928]" captionTargetBox="[159,1413,1184,1873]" captionTargetId="figure-62@13.[158,1430,1178,1881]" captionTargetPageId="13" captionText="FIGURE 44. Bayesian tree of the genus Baetis inferred from the cytochrome c oxidase I (COI) nucleotide sequence data (466 bp) and 47 morphological characters. Two species of Labiobaetis were used as outgroup. All data including species-groups are taken from Sroka (2012). Specimen obtained in this study is in bold. Bayesian posterior probabilities (PP) above 0.7 are given above tree nodes." figureDoi="http://doi.org/10.5281/zenodo.4736198" httpUri="https://zenodo.org/record/4736198/files/figure.png" pageId="11" pageNumber="552">44</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |