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<document id="37F73C98D9A40F8EAC2C39147E4FDC56" ID-DOI="10.5281/zenodo.11067355" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067355" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714052343267" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2021" docId="03AF0B0CFFFBFF93FF9FFEF4A590F953" docLanguage="en" docName="ProcCalifAcadSci.67.4.85-182.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (4)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.1:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="1" docTitle="Broscosoma gaoligongense Deuve and Wrase 2015" docType="treatment" docVersion="2" lastPageNumber="123" masterDocId="FF967374FFDFFFB5FFE4FFCFA106FFEE" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae)." masterLastPageNumber="182" masterPageNumber="85" pageNumber="121" updateTime="1714052772053" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="61361FC445202E3EC5B3F3E7417988F7">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae).</mods:title>
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<mods:namePart id="793210BC6C0CF4C2216CF2F81C6D74B8">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="FBC2E319315CBF534DEA2D4E13F340BC">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:namePart id="2069219416687F9C1A8FE255EA3E985E">Liang, Hongbin</mods:namePart>
<mods:affiliation id="B6E107F029BCF4171F9982A5CD9CEE18">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:title id="9CE85426CF1E790173BF5BF3DFADA15F">Proceedings of the California Academy of Sciences</mods:title>
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<mods:date id="373C414176D081E5371546DEA2CC3436">2021</mods:date>
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6. 
<taxonomicName id="4C06C199FFFBFF91FF72FEF3A3D1FEBD" ID-CoL="NDX3" authority="Deuve and Wrase, 2015" authorityName="Deuve and Wrase" authorityYear="2015" box="[150,727,315,340]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="gaoligongense">
<emphasis id="B9726608FFFBFF91FF72FEF3A0C5FEBD" bold="true" box="[150,451,315,339]" italics="true" pageId="36" pageNumber="121">Broscosoma gaoligongense</emphasis>
<bibRefCitation id="EF97C7EBFFFBFF91FE2EFEF3A3D1FEBD" author="Deuve, T. &amp; D. W. Wrase" box="[458,727,315,340]" pageId="36" pageNumber="121" pagination="25 - 29" refId="ref53183" refString="Deuve, T., and D. W. Wrase. 2015. Description d'un nouveau Broscosoma du Yunnan, Chine (Coleoptera, Caraboidea, Broscidae). Bulletin de la Societe Entomologique de France 120: 25 - 29." type="journal article" year="2015">Deuve and Wrase, 2015</bibRefCitation>
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<paragraph id="8BB9BA1AFFFBFF91FF9FFE93A0C9FE9C" blockId="36.[123,1174,315,1726]" box="[123,463,348,370]" pageId="36" pageNumber="121">
<figureCitation id="133DA69FFFFBFF91FF9FFE93A1E7FE9C" box="[123,225,348,370]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="36" pageNumber="121">Figures 7d</figureCitation>
, 
<figureCitation id="133DA69FFFFBFF91FF0AFE93A014FE9C" box="[238,274,348,370]" captionStart="FIGURE" captionStartId="13.[171,183,1441,1460]" captionTargetBox="[404,932,216,1422]" captionTargetId="figure-66@13.[404,932,216,1422]" captionTargetPageId="13" captionText="FIGURE 10. Head, Broscosoma species, lateral aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote endpoints of genal ridge." figureDoi="http://doi.org/10.5281/zenodo.11067371" httpUri="https://zenodo.org/record/11067371/files/figure.png" pageId="36" pageNumber="121">10b</figureCitation>
, 
<figureCitation id="133DA69FFFFBFF91FEFAFE93A030FE9F" box="[286,310,348,369]" captionStart="FIGURE" captionStartId="37.[171,183,1600,1619]" captionTargetBox="[125,1176,216,1581]" captionTargetId="figure-67@37.[120,1176,216,1586]" captionTargetPageId="37" captionText="FIGURE 20. Broscosoma gaoligongense Deuve and Wrase. a. Habitus (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067393" httpUri="https://zenodo.org/record/11067393/files/figure.png" pageId="36" pageNumber="121">20</figureCitation>
, 
<figureCitation id="133DA69FFFFBFF91FEA7FE93A05DFE9F" box="[323,347,348,369]" captionStart="FIGURE" captionStartId="38.[171,183,927,946]" captionTargetBox="[145,1143,223,907]" captionTargetId="figure-1565@38.[132,1162,218,913]" captionTargetPageId="38" captionText="FIGURE 21. Male genitalia, Broscosoma gaoligongense Deuve and Wrase. (CASENT1027154; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067395" httpUri="https://zenodo.org/record/11067395/files/figure.png" pageId="36" pageNumber="121">21</figureCitation>
, 
<figureCitation id="133DA69FFFFBFF91FE8CFE93A08DFE9F" box="[360,395,348,369]" captionStart="FIGURE" captionStartId="84.[171,183,1585,1604]" captionTargetBox="[147,1149,216,1572]" captionTargetId="figure-57@84.[147,1149,216,1572]" captionTargetPageId="84" captionText="FIGURE 44. Photographs of habitats for Broscosoma species in the Gaoligong Shan region. a. 0.5 km E of Fengxue Yakou, 3150 m, 25.972º/098.683 º, Luzhang Township, Lushui County, Yunnan (habitat for Broscosoma gaoligongense sp. nov.); b. Xiao Jiang at Gangfang Sancha Lukou, 1500 m, 26.122º/98.573º, Lushui County, Yunnan (habitat for Broscosoma holomarginatum sp. nov.). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067447" httpUri="https://zenodo.org/record/11067447/files/figure.png" pageId="36" pageNumber="121">44a</figureCitation>
, 
<figureCitation id="133DA69FFFFBFF91FE73FE93A0C9FE9C" box="[407,463,348,370]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="88.[171,183,995,1014]" captionStartId-1="88.[184,196,1682,1701]" captionStartId-2="89.[171,183,1461,1480]" captionTargetBox-0="[130,1164,233,969]" captionTargetBox-1="[130,1162,1063,1660]" captionTargetBox-2="[140,1154,249,1428]" captionTargetId-0="figure-57@88.[120,1176,215,989]" captionTargetId-1="figure-332@88.[120,1176,1054,1668]" captionTargetId-2="figure-68@89.[120,1176,216,1448]" captionTargetPageId-0="88" captionTargetPageId-1="88" captionTargetPageId-2="89" captionText-0="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." captionText-2="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi-0="http://doi.org/10.5281/zenodo.11067455" figureDoi-1="http://doi.org/10.5281/zenodo.11067457" figureDoi-2="http://doi.org/10.5281/zenodo.11067459" httpUri-0="https://zenodo.org/record/11067455/files/figure.png" httpUri-1="https://zenodo.org/record/11067457/files/figure.png" httpUri-2="https://zenodo.org/record/11067459/files/figure.png" pageId="36" pageNumber="121">48-50</figureCitation>
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<paragraph id="8BB9BA1AFFFBFF91FF9FFE4EA590FE78" blockId="36.[123,1174,315,1726]" box="[123,1174,384,406]" pageId="36" pageNumber="121">
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<treatmentCitation id="0AA79C0BFFFBFF91FF9FFE4EA388FE7B" author="Deuve, T. &amp; D. W. Wrase" box="[123,654,384,406]" page="29" pageId="36" pageNumber="121" year="2015">
<taxonomicName id="4C06C199FFFBFF91FF9FFE4EA388FE7B" ID-CoL="NDX3" authority="Deuve and Wrase, 2015: 29" authorityName="Deuve and Wrase" authorityPageNumber="29" authorityYear="2015" box="[123,654,384,406]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="gaoligongense">
<emphasis id="B9726608FFFBFF91FF9FFE4EA085FE7B" box="[123,387,384,406]" italics="true" pageId="36" pageNumber="121">Broscosoma gaoligongense</emphasis>
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<bibRefCitation id="EF97C7EBFFFBFF91FE6DFE4EA388FE7B" author="Deuve, T. &amp; D. W. Wrase" box="[393,654,384,406]" pageId="36" pageNumber="121" pagination="25 - 29" refId="ref53183" refString="Deuve, T., and D. W. Wrase. 2015. Description d'un nouveau Broscosoma du Yunnan, Chine (Coleoptera, Caraboidea, Broscidae). Bulletin de la Societe Entomologique de France 120: 25 - 29." type="journal article" year="2015">Deuve and Wrase, 2015:29</bibRefCitation>
. 
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, a male, deposited in 
<collectionCode id="ED1722DFFFFBFF91FC2CFE4EA502FE78" box="[968,1028,385,406]" pageId="36" pageNumber="121">DWW</collectionCode>
. Type locality:
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</paragraph>
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<materialsCitation id="3B6EB047FFFBFF91FF4FFE52A51DFE5D" box="[171,1051,412,435]" country="China" elevation="3150" location="Gaoligong Shan" pageId="36" pageNumber="121" specimenCount="1" stateProvince="Yunnan">
<collectingCountry id="F311FA8AFFFBFF91FF4FFE52A1E2FE5D" box="[171,228,413,435]" name="China" pageId="36" pageNumber="121">China</collectingCountry>
, 
<collectingRegion id="49C274F8FFFBFF91FF10FE51A03BFE5D" box="[244,317,414,435]" country="China" name="Yunnan" pageId="36" pageNumber="121">Yunnan</collectingRegion>
, 
<location id="8ED9ECC1FFFBFF91FEAAFE52A0E8FE5D" LSID="urn:lsid:plazi:treatment:03AF0B0CFFFBFF93FF9FFEF4A590F953:8ED9ECC1FFFBFF91FEAAFE52A0E8FE5D" box="[334,494,413,435]" country="China" name="Gaoligong Shan" pageId="36" pageNumber="121" stateProvince="Yunnan">Gaoligong Shan</location>
, pass [Fengxue Yakou] 
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NW of Liuku
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, 
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, 25.97281°/
</paragraph>
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<paragraph id="8BB9BA1AFFFBFF91FF4FFE75A012FE3E" blockId="36.[123,1174,315,1726]" box="[171,276,442,464]" pageId="36" pageNumber="121">98.68339°.</paragraph>
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<subSubSection id="C31CE991FFFBFF91FF4FFE11A2F4FCF6" pageId="36" pageNumber="121" type="diagnosis">
<paragraph id="8BB9BA1AFFFBFF91FF4FFE11A2F4FCF6" blockId="36.[123,1174,315,1726]" pageId="36" pageNumber="121">
<emphasis id="B9726608FFFBFF91FF4FFE11A019FE19" bold="true" box="[171,287,478,503]" pageId="36" pageNumber="121">Diagnosis.</emphasis>
Adults of this species (
<figureCitation id="133DA69FFFFBFF91FDC5FE10A37CFE18" box="[545,634,479,503]" captionStart="FIGURE" captionStartId="37.[171,183,1600,1619]" captionTargetBox="[125,1176,216,1581]" captionTargetId="figure-67@37.[120,1176,216,1586]" captionTargetPageId="37" captionText="FIGURE 20. Broscosoma gaoligongense Deuve and Wrase. a. Habitus (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067393" httpUri="https://zenodo.org/record/11067393/files/figure.png" pageId="36" pageNumber="121">Fig. 20a</figureCitation>
) can be distinguished from those of other species in the region by the following combination of character states: size moderate, BL male = 9.0–10.0 mm, female = 9.6–10. 
<quantity id="4CFE17FFFFFBFF91FE87FDD0A099FDD8" box="[355,415,543,567]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="121" unit="mm" value="2.0">2 mm</quantity>
; dorsum of head and pronotum black or dark piceous, elytra with bright green or blue-green metallic reflection; genal ridge evident basal to post-temporal constriction but extended only to or slightly anterior to the constriction (
<figureCitation id="133DA69FFFFBFF91FD32FD90A229FD99" box="[726,815,607,631]" captionStart="FIGURE" captionStartId="13.[171,183,1441,1460]" captionTargetBox="[404,932,216,1422]" captionTargetId="figure-66@13.[404,932,216,1422]" captionTargetPageId="13" captionText="FIGURE 10. Head, Broscosoma species, lateral aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote endpoints of genal ridge." figureDoi="http://doi.org/10.5281/zenodo.11067371" httpUri="https://zenodo.org/record/11067371/files/figure.png" pageId="36" pageNumber="121">Fig. 10b</figureCitation>
); antennomeres 3 and 4 glabrous except for apical whorl of setae, antennomeres 1 through 4 concolorous, pale rufous; pronotum anterior to sub-basal constriction (
<figureCitation id="133DA69FFFFBFF91FE0FFD6FA331FD56" box="[491,567,672,696]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="36" pageNumber="121">Fig. 7d</figureCitation>
) globose in dorsal view, basolateral setae absent; elytral silhouette ovoid, with humeri indistinct; metatrochanters unisetose; male protarsi with tarsomeres 1 to 3 with ventral pads of adhesive setae; male genitalia as in 
<figureCitation id="133DA69FFFFBFF91FCC2FD2FA275FD16" box="[806,883,736,760]" captionStart="FIGURE" captionStartId="38.[171,183,927,946]" captionTargetBox="[145,1143,223,907]" captionTargetId="figure-1565@38.[132,1162,218,913]" captionTargetPageId="38" captionText="FIGURE 21. Male genitalia, Broscosoma gaoligongense Deuve and Wrase. (CASENT1027154; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067395" httpUri="https://zenodo.org/record/11067395/files/figure.png" pageId="36" pageNumber="121">Fig. 21</figureCitation>
, with median lobe slightly shorter and thicker and apical lamella more symmetrically rounded in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFFBFF93FF4FFCEFA590F953" lastPageId="38" lastPageNumber="123" pageId="36" pageNumber="121" type="discussion">
<paragraph id="8BB9BA1AFFFBFF91FF4FFCEFA30EFA75" blockId="36.[123,1174,315,1726]" pageId="36" pageNumber="121">
<emphasis id="B9726608FFFBFF91FF4FFCEFA020FCD6" bold="true" box="[171,294,800,824]" pageId="36" pageNumber="121">Comments</emphasis>
. As noted in the Comments section for 
<taxonomicName id="4C06C199FFFBFF91FD37FCEEA275FCD6" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[723,883,800,824]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFFBFF91FD37FCEEA275FCD6" box="[723,883,800,824]" italics="true" pageId="36" pageNumber="121">B. bicoloratum</emphasis>
</taxonomicName>
, this is one of four species in the study area members of which have the elytra with distinct metallic reflection but the forebody (head and pronotum) without metallic reflection or with that reflection confined to the pronotal area anterior to the anterior transverse impression. Specimens of 
<taxonomicName id="4C06C199FFFBFF91FCB9FC4DA51EFC77" box="[861,1048,897,921]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="gaoligongense">
<emphasis id="B9726608FFFBFF91FCB9FC4DA51EFC77" box="[861,1048,897,921]" italics="true" pageId="36" pageNumber="121">B. gaoligongense</emphasis>
</taxonomicName>
can be distinguished from those of 
<taxonomicName id="4C06C199FFFBFF91FE60FC6DA323FC57" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[388,549,929,953]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="viridicollare">
<emphasis id="B9726608FFFBFF91FE60FC6DA323FC57" box="[388,549,929,953]" italics="true" pageId="36" pageNumber="121">B. viridicollare</emphasis>
</taxonomicName>
in having the pronotum globose and without a trace metallic reflection (pronotum more elongate and with the area anterior to the anterior transverse impression with a distinct metallic green reflection in 
<taxonomicName id="4C06C199FFFBFF91FD91FC2DA211FC17" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[629,791,993,1017]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="viridicollare">
<emphasis id="B9726608FFFBFF91FD91FC2DA211FC17" box="[629,791,993,1017]" italics="true" pageId="36" pageNumber="121">B. viridicollare</emphasis>
</taxonomicName>
), the elytral microsculpture effaced (finely impressed but distinct in 
<taxonomicName id="4C06C199FFFBFF91FE37FBCDA373FBF4" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[467,629,1026,1050]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="viridicollare">
<emphasis id="B9726608FFFBFF91FE37FBCDA373FBF4" box="[467,629,1026,1050]" italics="true" pageId="36" pageNumber="121">B. viridicollare</emphasis>
</taxonomicName>
), the parascutellare pore puncture inserted near the base of stria 3 (stria 
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<taxonomicName id="4C06C199FFFBFF91FE75FBECA333FBD4" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[401,565,1058,1082]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="viridicollare">
<emphasis id="B9726608FFFBFF91FE75FBECA333FBD4" box="[401,565,1058,1082]" italics="true" pageId="36" pageNumber="121">B. viridicollare</emphasis>
</taxonomicName>
), and the median lobe of the male genitalia shorter and more evenly arcuate (
<figureCitation id="133DA69FFFFBFF91FE8DFB8DA0C4FBB4" box="[361,450,1090,1114]" captionStart="FIGURE" captionStartId="38.[171,183,927,946]" captionTargetBox="[145,1143,223,907]" captionTargetId="figure-1565@38.[132,1162,218,913]" captionTargetPageId="38" captionText="FIGURE 21. Male genitalia, Broscosoma gaoligongense Deuve and Wrase. (CASENT1027154; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067395" httpUri="https://zenodo.org/record/11067395/files/figure.png" pageId="36" pageNumber="121">Fig. 21c</figureCitation>
) (longer and less arcuate in apical third (
<figureCitation id="133DA69FFFFBFF91FC6EFB8DA2E5FBB4" box="[906,995,1090,1114]" captionStart="FIGURE" captionStartId="69.[171,183,922,941]" captionTargetBox="[144,1155,225,900]" captionTargetId="figure-1615@69.[127,1169,215,908]" captionTargetPageId="69" captionText="FIGURE 35. Male genitalia, Broscosoma viridicollare sp. nov. (CASENT1017820; second cirque S of Shibali Yakou, Lumadeng Township, Fugong County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067429" httpUri="https://zenodo.org/record/11067429/files/figure.png" pageId="36" pageNumber="121">Fig. 35c</figureCitation>
) in 
<taxonomicName id="4C06C199FFFBFF91FBF6FB8CA1A3FB94" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="viridicollare">
<emphasis id="B9726608FFFBFF91FBF6FB8CA1A3FB94" italics="true" pageId="36" pageNumber="121">B. viridicollare</emphasis>
</taxonomicName>
). They differ from specimens of 
<taxonomicName id="4C06C199FFFBFF91FDEEFBACA3ACFB94" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[522,682,1122,1146]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFFBFF91FDEEFBACA3ACFB94" box="[522,682,1122,1146]" italics="true" pageId="36" pageNumber="121">B. bicoloratum</emphasis>
</taxonomicName>
in having the metatrochanters unisetose (asetose in most 
<taxonomicName id="4C06C199FFFBFF91FEE1FB4CA0A0FB74" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[261,422,1154,1178]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFFBFF91FEE1FB4CA0A0FB74" box="[261,422,1154,1178]" italics="true" pageId="36" pageNumber="121">B. bicoloratum</emphasis>
</taxonomicName>
, but one seta present unilaterally in a few specimens), the parascutellar setiferous pore inserted near the base of stria 3 (near base of stria 
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most 
<taxonomicName id="4C06C199FFFBFF91FC5CFB6CA551FB55" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[952,1111,1187,1211]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFFBFF91FC5CFB6CA551FB55" box="[952,1111,1187,1211]" italics="true" pageId="36" pageNumber="121">B. bicoloratum</emphasis>
</taxonomicName>
specimens), the eyes (
<figureCitation id="133DA69FFFFBFF91FEDDFB0CA080FB35" box="[313,390,1219,1243]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="36" pageNumber="121">Fig. 7d</figureCitation>
) slightly less convex (more convex in 
<taxonomicName id="4C06C199FFFBFF91FCCEFB0CA2CDFB35" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[810,971,1219,1243]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFFBFF91FCCEFB0CA2CDFB35" box="[810,971,1219,1243]" italics="true" pageId="36" pageNumber="121">B. bicoloratum</emphasis>
</taxonomicName>
(
<figureCitation id="133DA69FFFFBFF91FC38FB0CA521FB34" box="[988,1063,1219,1243]" captionStart="FIGURE" captionStartId="10.[171,183,1484,1503]" captionTargetBox="[120,1176,216,1467]" captionTargetId="figure-55@10.[120,1176,216,1467]" captionTargetPageId="10" captionText="FIGURE 7. Forebody, Broscosoma species, dorsal aspect. a. B. furvum sp. nov. (CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China); b. B. viridicollare sp. nov. (CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China); c. B. bicoloratum sp. nov. (CASENT1026704; slope NW of Heipu Yakou, Dulongjiang Townhip, Gongshan County, Yunnan, China); d. B. gaoligongense Deuve and Wrase (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China); e. B. gongshanense sp. nov. (CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067365" httpUri="https://zenodo.org/record/11067365/files/figure.png" pageId="36" pageNumber="121">Fig. 7c</figureCitation>
)), and the median lobe of male genitalia slightly narrower and with the apical lamella slightly more expand- ed apically (median lobe slightly thicker and with a slightly narrower apical lamella (
<figureCitation id="133DA69FFFFBFF91FBF3FACCA576FAF5" box="[1047,1136,1283,1307]" captionStart="FIGURE" captionStartId="26.[171,183,963,982]" captionTargetBox="[142,1152,219,938]" captionTargetId="figure-1559@26.[120,1176,215,955]" captionTargetPageId="26" captionText="FIGURE 16. Male genitalia, Broscosoma bicoloratum sp. nov. (CASENT1025709; southeast slope of Heipu Yakou, Cikai Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067383" httpUri="https://zenodo.org/record/11067383/files/figure.png" pageId="36" pageNumber="121">Fig. 16c</figureCitation>
) in 
<taxonomicName id="4C06C199FFFBFF91FF9FFAEBA01CFAD5" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[123,282,1315,1339]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFFBFF91FF9FFAEBA01CFAD5" box="[123,282,1315,1339]" italics="true" pageId="36" pageNumber="121">B. bicoloratum</emphasis>
</taxonomicName>
). They differ from members of 
<taxonomicName id="4C06C199FFFBFF91FD8AFAEBA225FAD5" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[622,803,1315,1339]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="gongshanense">
<emphasis id="B9726608FFFBFF91FD8AFAEBA225FAD5" box="[622,803,1315,1339]" italics="true" pageId="36" pageNumber="121">B. gongshanense</emphasis>
</taxonomicName>
in having the elytra with a distinct metallic green reflection (metallic reflection distinctly more bluish in 
<taxonomicName id="4C06C199FFFBFF91FC86FA8BA511FAB5" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[866,1047,1347,1371]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="gongshanense">
<emphasis id="B9726608FFFBFF91FC86FA8BA511FAB5" box="[866,1047,1347,1371]" italics="true" pageId="36" pageNumber="121">B. gongshanense</emphasis>
</taxonomicName>
), the parascutellar seta inserted at the base of stria 3 (stria 
<quantity id="4CFE17FFFFFBFF91FD71FAABA3C7FA92" box="[661,705,1380,1404]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="36" pageNumber="121" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C06C199FFFBFF91FD2EFAABA284FA92" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[714,898,1380,1404]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="gongshanense">
<emphasis id="B9726608FFFBFF91FD2EFAABA284FA92" box="[714,898,1380,1404]" italics="true" pageId="36" pageNumber="121">B. gongshanense</emphasis>
</taxonomicName>
), and the metatrochanter unisetose (asetose 
<taxonomicName id="4C06C199FFFBFF91FEA6FA4AA0FEFA72" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[322,504,1412,1436]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="gongshanense">
<emphasis id="B9726608FFFBFF91FEA6FA4AA0FEFA72" box="[322,504,1412,1436]" italics="true" pageId="36" pageNumber="121">B. gongshanense</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BB9BA1AFFFBFF91FF4FFA6BA1C6F993" blockId="36.[123,1174,315,1726]" pageId="36" pageNumber="121">
As noted above in the Comments section for 
<taxonomicName id="4C06C199FFFBFF91FD54FA6AA252FA52" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[688,852,1444,1468]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="bicoloratum">
<emphasis id="B9726608FFFBFF91FD54FA6AA252FA52" box="[688,852,1444,1468]" italics="true" pageId="36" pageNumber="121">B. bicoloratum</emphasis>
</taxonomicName>
, among 
<taxonomicName id="4C06C199FFFBFF91FC5EFA6AA593FA52" box="[954,1173,1444,1468]" pageId="36" pageNumber="121">
<emphasis id="B9726608FFFBFF91FC5EFA6AA53BFA52" box="[954,1085,1445,1468]" italics="true" pageId="36" pageNumber="121">Broscosoma</emphasis>
species
</taxonomicName>
known from outside the study area, only 
<taxonomicName id="4C06C199FFFBFF91FDACFA0AA3D2FA32" authorityName="Habu" authorityYear="1973" box="[584,724,1476,1500]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B9726608FFFBFF91FDACFA0AA3D2FA32" box="[584,724,1476,1500]" italics="true" pageId="36" pageNumber="121">B. monticola</emphasis>
</taxonomicName>
has members with the forebody lacking metallic reflection and the elytra with markedly metallic reflection as is seen in 
<taxonomicName id="4C06C199FFFBFF91FC3FFA2AA590FA12" box="[987,1174,1508,1532]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="gaoligongense">
<emphasis id="B9726608FFFBFF91FC3FFA2AA590FA12" box="[987,1174,1508,1532]" italics="true" pageId="36" pageNumber="121">B. gaoligongense</emphasis>
</taxonomicName>
specimens. They also also have the parascutellar seta inserted near the base of stria 3; but they have the femora slightly paler and the median lobe of the male genitalia with a large protuberance on the ventral margin (see 
<bibRefCitation id="EF97C7EBFFFBFF91FEB4F98AA0C0F9B3" author="Habu, A." box="[336,454,1605,1629]" pageId="36" pageNumber="121" pagination="81 - 132" refId="ref53285" refString="Habu, A. 1973. On a collection of Carabidae from Nepal made by the Hokkaido University Scientifi c Exedition to Nepal Himalaya, 1968 (I). The Bulletin of the National Institute of Agricultural Sciences (C) 27: 81 - 132." type="journal article" year="1973">Habu 1973</bibRefCitation>
, fig. 6), which is absent from the median lobe of 
<taxonomicName id="4C06C199FFFBFF91FC38F989A590F9B3" box="[988,1174,1605,1629]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="121" phylum="Arthropoda" rank="species" species="gaoligongense">
<emphasis id="B9726608FFFBFF91FC38F989A590F9B3" box="[988,1174,1605,1629]" italics="true" pageId="36" pageNumber="121">B. gaoligongense</emphasis>
</taxonomicName>
males.
</paragraph>
<paragraph id="8BB9BA1AFFFBFF93FF4FF94AA1E6FB73" blockId="36.[123,1174,315,1726]" lastBlockId="38.[123,1174,1029,1726]" lastPageId="38" lastPageNumber="123" pageId="36" pageNumber="121">
<emphasis id="B9726608FFFBFF91FF4FF94AA090F973" bold="true" box="[171,406,1669,1693]" pageId="36" pageNumber="121">Habitat distribution.</emphasis>
A few members of this species were found under stones on organic substrate along the edges of the roadcut through bamboo and 
<taxonomicName id="4C06C199FFFBFF91FD38F96AA27CF953" box="[732,890,1701,1725]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="36" pageNumber="121" phylum="Tracheophyta" rank="genus">
<emphasis id="B9726608FFFBFF91FD38F96AA27CF953" box="[732,890,1701,1725]" italics="true" pageId="36" pageNumber="121">Rhododendron</emphasis>
</taxonomicName>
thickets (
<figureCitation id="133DA69FFFFBFF91FC05F96AA531F953" box="[993,1079,1701,1726]" captionStart="FIGURE" captionStartId="84.[171,183,1585,1604]" captionTargetBox="[147,1149,216,1572]" captionTargetId="figure-57@84.[147,1149,216,1572]" captionTargetPageId="84" captionText="FIGURE 44. Photographs of habitats for Broscosoma species in the Gaoligong Shan region. a. 0.5 km E of Fengxue Yakou, 3150 m, 25.972º/098.683 º, Luzhang Township, Lushui County, Yunnan (habitat for Broscosoma gaoligongense sp. nov.); b. Xiao Jiang at Gangfang Sancha Lukou, 1500 m, 26.122º/98.573º, Lushui County, Yunnan (habitat for Broscosoma holomarginatum sp. nov.). Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067447" httpUri="https://zenodo.org/record/11067447/files/figure.png" pageId="36" pageNumber="121">Fig. 44a</figureCitation>
) on both sides of the pass (Fengxue Yakou) during the day. The majority of specimens were collected at night, with the aid of headlamps, where they were found active on the surface of the ground, especially on mosses, in thickets and on the banks of the roadcut. This species is found at moderately high elevations, with our records documenting its occurrence in the 
<quantity id="4CFE17FFFFF9FF93FC75FBAAA54BFB93" box="[913,1101,1125,1149]" metricMagnitude="3" metricUnit="m" metricValue="3.135" metricValueMax="3.15" metricValueMin="3.12" pageId="38" pageNumber="123" unit="m" value="3135.0" valueMax="3150.0" valueMin="3120.0">3120 to 3150 m</quantity>
range (
<figureCitation id="133DA69FFFF9FF93FF60FB4AA1D6FB73" box="[132,208,1157,1181]" captionStart="FIGURE" captionStartId="88.[184,196,1682,1701]" captionTargetBox="[130,1162,1063,1660]" captionTargetId="figure-332@88.[120,1176,1054,1668]" captionTargetPageId="88" captionText="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." figureDoi="http://doi.org/10.5281/zenodo.11067457" httpUri="https://zenodo.org/record/11067457/files/figure.png" pageId="38" pageNumber="123">Fig. 49</figureCitation>
).
</paragraph>
<caption id="DF79EA92FFFAFF90FF4FF98FA3E1F968" ID-DOI="http://doi.org/10.5281/zenodo.11067393" ID-Zenodo-Dep="11067393" httpUri="https://zenodo.org/record/11067393/files/figure.png" pageId="37" pageNumber="122" startId="37.[171,183,1600,1619]" targetBox="[125,1176,216,1581]" targetPageId="37" targetType="figure">
<paragraph id="8BB9BA1AFFFAFF90FF4FF98FA3E1F968" blockId="37.[123,1173,1599,1670]" pageId="37" pageNumber="122">
<smallCapsWord id="8D5F2CC6FFFAFF90FF4FF98FA1EEF9BF" baselines="1614,1613" box="[171,232,1600,1619]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="37" pageNumber="122">FIGURE</smallCapsWord>
20. 
<taxonomicName id="4C06C199FFFAFF90FEF4F98FA3A4F9BD" authority="Deuve and Wrase." authorityName="Deuve and Wrase." box="[272,674,1599,1619]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="122" phylum="Arthropoda" rank="species" species="gaoligongense">
<emphasis id="B9726608FFFAFF90FEF4F98FA0FAF9BC" box="[272,508,1599,1618]" italics="true" pageId="37" pageNumber="122">Broscosoma gaoligongense</emphasis>
Deuve and Wrase.
</taxonomicName>
a. Habitus (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<caption id="DF79EA92FFF9FF93FF4FFC50A265FC0B" ID-DOI="http://doi.org/10.5281/zenodo.11067395" ID-Zenodo-Dep="11067395" httpUri="https://zenodo.org/record/11067395/files/figure.png" pageId="38" pageNumber="123" startId="38.[171,183,927,946]" targetBox="[145,1143,223,907]" targetPageId="38" targetType="figure">
<paragraph id="8BB9BA1AFFF9FF93FF4FFC50A265FC0B" blockId="38.[123,1174,926,997]" pageId="38" pageNumber="123">
<smallCapsWord id="8D5F2CC6FFF9FF93FF4FFC50A1EEFC5E" baselines="941,940" box="[171,232,927,946]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="38" pageNumber="123">FIGURE</smallCapsWord>
21. Male genitalia, 
<taxonomicName id="4C06C199FFF9FF93FE45FC50A23DFC5C" authority="Deuve and Wrase." authorityName="Deuve and Wrase." box="[417,827,926,946]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="123" phylum="Arthropoda" rank="species" species="gaoligongense">
<emphasis id="B9726608FFF9FF93FE45FC50A388FC5F" box="[417,654,926,945]" italics="true" pageId="38" pageNumber="123">Broscosoma gaoligongense</emphasis>
Deuve and Wrase.
</taxonomicName>
(CASENT1027154; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB9BA1AFFF9FF93FF4FFB6AA1C6FA33" blockId="38.[123,1174,1029,1726]" pageId="38" pageNumber="123">
<emphasis id="B9726608FFF9FF93FF4FFB6AA216FB53" bold="true" box="[171,784,1189,1213]" pageId="38" pageNumber="123">
Geographical distribution within the Gaoligong 
<materialsCitation id="3B6EB047FFF9FF93FD2BFB6AA326FB33" location="We" pageId="38" pageNumber="123" specimenCount="70" specimenCount-female="12" specimenCount-male="23" stateProvince="Shan">
<collectingRegion id="49C274F8FFF9FF93FD2BFB6AA20FFB53" box="[719,777,1189,1213]" country="Myanmar" name="Shan" pageId="38" pageNumber="123">Shan</collectingRegion>
. 
<figureCitation id="133DA69FFFF9FF93FCF2FB6AA269FB53" box="[790,879,1189,1213]" captionStart="FIGURE" captionStartId="37.[171,183,1600,1619]" captionTargetBox="[125,1176,216,1581]" captionTargetId="figure-67@37.[120,1176,216,1586]" captionTargetPageId="37" captionText="FIGURE 20. Broscosoma gaoligongense Deuve and Wrase. a. Habitus (CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067393" httpUri="https://zenodo.org/record/11067393/files/figure.png" pageId="38" pageNumber="123">Fig. 20b</figureCitation>
. 
<location id="8ED9ECC1FFF9FF93FC98FB69A299FB53" LSID="urn:lsid:plazi:treatment:03AF0B0CFFFBFF93FF9FFEF4A590F953:8ED9ECC1FFF9FF93FC98FB69A299FB53" box="[892,927,1190,1213]" name="We" pageId="38" pageNumber="123" stateProvince="Shan">We</location>
examined a total of 
<specimenCount id="9D007193FFF9FF93FB9FFB6AA1ECFB33" count="35" pageId="38" pageNumber="123" type="generic">35 specimens</specimenCount>
(
<specimenCount id="9D007193FFF9FF93FF1AFB0AA067FB33" box="[254,353,1221,1245]" count="23" pageId="38" pageNumber="123" type="male">23 males</specimenCount>
and 
<specimenCount id="9D007193FFF9FF93FE7AFB0AA311FB33" box="[414,535,1221,1245]" count="12" pageId="38" pageNumber="123" type="female">12 females</specimenCount>
)
</materialsCitation>
</emphasis>
<materialsCitation id="3B6EB047FFF9FF93FDCFFB0AA1BFFA33" collectingDate="2005-05-11" collectingDateMax="2005-05-19" collectingDateMin="2005-05-11" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; H. B. Liang &amp; C E. Griswold, D. Z &amp; K. J. Guo &amp; Fengxue Yakou &amp; Pianma Road &amp; D. Kavanaugh &amp; C. E. Griswold &amp; D. Z. Dong" county="Lushui County" elevation="3150" location="Pianma" municipality="Luzhang Township" pageId="38" pageNumber="123" specimenCount="35" specimenCount-female="12" specimenCount-male="23" stateProvince="Shan">
from the following localities: 
<emphasis id="B9726608FFF9FF93FC9FFB0AA52BFB33" bold="true" box="[891,1069,1221,1245]" italics="true" pageId="38" pageNumber="123">
<collectingCounty id="62D8C296FFF9FF93FC9FFB0AA522FB33" box="[891,1060,1221,1245]" pageId="38" pageNumber="123">Lushui County</collectingCounty>
:
</emphasis>
<collectingMunicipality id="6BDD2060FFF9FF93FBDCFB0AA1E2FB13" pageId="38" pageNumber="123">Luzhang Township</collectingMunicipality>
(
<location id="8ED9ECC1FFF9FF93FF13FB2AA041FB13" LSID="urn:lsid:plazi:treatment:03AF0B0CFFFBFF93FF9FFEF4A590F953:8ED9ECC1FFF9FF93FF13FB2AA041FB13" box="[247,327,1253,1277]" county="Lushui County" municipality="Luzhang Township" name="Pianma" pageId="38" pageNumber="123" stateProvince="Shan">Pianma</location>
road at 
<location id="8ED9ECC1FFF9FF93FE4CFB29A357FB13" LSID="urn:lsid:plazi:treatment:03AF0B0CFFFBFF93FF9FFEF4A590F953:8ED9ECC1FFF9FF93FE4CFB29A357FB13" box="[424,593,1253,1278]" county="Lushui County" municipality="Luzhang Township" name="Fengxue Yakou" pageId="38" pageNumber="123" stateProvince="Shan">Fengxue Yakou</location>
, 25.97228º/098.68336º, 
<quantity id="4CFE17FFFFF9FF93FC85FB2AA2ADFB13" box="[865,939,1253,1277]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="38" pageNumber="123" unit="m" value="3150.0">
<elevation id="002B5D29FFF9FF93FC85FB2AA2ADFB13" box="[865,939,1253,1277]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="38" pageNumber="123" unit="m" value="3150.0">3150m</elevation>
</quantity>
, 
<date id="FFB89CDAFFF9FF93FC58FB2AA556FB13" box="[956,1104,1253,1278]" pageId="38" pageNumber="123" value="2005-05-11">
<collectingDate id="EFFC6532FFF9FF93FC58FB2AA556FB13" box="[956,1104,1253,1278]" pageId="38" pageNumber="123" value="2005-05-11">11 May 2005</collectingDate>
</date>
, 
<collectorName id="26F3DFCCFFF9FF93FB85FB29A1F2FAF3" pageId="38" pageNumber="123">D.H. Kavanaugh</collectorName>
, 
<collectorName id="26F3DFCCFFF9FF93FEE2FAC9A084FAF3" box="[262,386,1285,1309]" pageId="38" pageNumber="123">H.B. Liang</collectorName>
, 
<collectorName id="26F3DFCCFFF9FF93FE70FACAA375FAF3" box="[404,627,1285,1309]" pageId="38" pageNumber="123">C E. Griswold, D.Z</collectorName>
. Dong &amp; 
<collectorName id="26F3DFCCFFF9FF93FD09FAC9A257FAF3" box="[749,849,1285,1309]" pageId="38" pageNumber="123">K.J. Guo</collectorName>
collectors [
<specimenCount id="9D007193FFF9FF93FC3CFAC9A538FAF3" box="[984,1086,1285,1309]" count="1" pageId="38" pageNumber="123" type="male">one male</specimenCount>
; 
<collectionCode id="ED1722DFFFF9FF93FBB5FACAA583FAF3" box="[1105,1157,1285,1309]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="38" pageNumber="123" type="Museum">CAS</collectionCode>
]; 
<collectorName id="26F3DFCCFFF9FF93FF9FFAE9A019FAD3" box="[123,287,1317,1342]" pageId="38" pageNumber="123">Fengxue Yakou</collectorName>
to 
<quantity id="4CFE17FFFFF9FF93FEA4FAEAA08FFAD3" box="[320,393,1317,1341]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="123" unit="km" value="0.5">0.5 km</quantity>
E on 
<collectorName id="26F3DFCCFFF9FF93FE22FAEAA352FAD3" box="[454,596,1317,1341]" pageId="38" pageNumber="123">Pianma Road</collectorName>
, 25.97288°/98.68336° to 25.97347°/98.68780°, 
<quantity id="4CFE17FFFFF9FF93FBB3FAEAA1C8FAB3" metricMagnitude="3" metricUnit="m" metricValue="3.1399999999999997" metricValueMax="3.15" metricValueMin="3.13" pageId="38" pageNumber="123" unit="m" value="3140.0" valueMax="3150.0" valueMin="3130.0">
<elevation id="002B5D29FFF9FF93FBB3FAEAA1C8FAB3" metricMagnitude="3" metricUnit="m" metricValue="3.1399999999999997" metricValueMax="3.15" metricValueMin="3.13" pageId="38" pageNumber="123" unit="m" value="3140.0" valueMax="3150.0" valueMin="3130.0">3130- 3150 m</elevation>
</quantity>
, 
<date id="FFB89CDAFFF9FF93FF38FA8AA068FAB3" box="[220,366,1349,1374]" pageId="38" pageNumber="123" value="2005-05-17">
<collectingDate id="EFFC6532FFF9FF93FF38FA8AA068FAB3" box="[220,366,1349,1374]" pageId="38" pageNumber="123" value="2005-05-17">17 May 2005</collectingDate>
</date>
, 
<collectorName id="26F3DFCCFFF9FF93FE9AFA89A31FFAB3" box="[382,537,1349,1373]" pageId="38" pageNumber="123">D. Kavanaugh</collectorName>
, 
<collectorName id="26F3DFCCFFF9FF93FDCDFA8AA3C2FAB3" box="[553,708,1349,1373]" pageId="38" pageNumber="123">C.E. Griswold</collectorName>
, 
<collectorName id="26F3DFCCFFF9FF93FD37FA89A24AFAB3" box="[723,844,1349,1373]" pageId="38" pageNumber="123">H.B. Liang</collectorName>
, &amp; 
<collectorName id="26F3DFCCFFF9FF93FC9DFA89A2EBFAB0" box="[889,1005,1350,1374]" pageId="38" pageNumber="123">D.Z. Dong</collectorName>
collectors [
<specimenCount id="9D007193FFF9FF93FB8BFA89A1C0FA93" count="1" pageId="38" pageNumber="123" type="female">one female</specimenCount>
, 
<collectionCode id="ED1722DFFFF9FF93FF05FAAAA008FA93" box="[225,270,1381,1405]" pageId="38" pageNumber="123">IOZ</collectionCode>
]); 
<location id="8ED9ECC1FFF9FF93FEDAFAAAA317FA93" LSID="urn:lsid:plazi:treatment:03AF0B0CFFFBFF93FF9FFEF4A590F953:8ED9ECC1FFF9FF93FEDAFAAAA317FA93" box="[318,529,1381,1405]" county="Lushui County" municipality="Luzhang Township" name="Pianma Township" pageId="38" pageNumber="123" stateProvince="Shan">Pianma Township</location>
(
<collectorName id="26F3DFCCFFF9FF93FDCAFAA9A3E1FA93" box="[558,743,1381,1406]" pageId="38" pageNumber="123">Fengxue Yakou</collectorName>
to 
<quantity id="4CFE17FFFFF9FF93FCC0FAAAA278FA93" box="[804,894,1381,1405]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="38" pageNumber="123" unit="km" value="0.6">0.6 km</quantity>
W on 
<collectorName id="26F3DFCCFFF9FF93FC09FAAAA588FA93" box="[1005,1166,1381,1405]" pageId="38" pageNumber="123">Pianma Road</collectorName>
, 25.97228º/98.68336º to 25.974108º/98.67716º, 
<quantity id="4CFE17FFFFF9FF93FD67FA4AA210FA73" box="[643,790,1413,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.135" metricValueMax="3.15" metricValueMin="3.12" pageId="38" pageNumber="123" unit="m" value="3135.0" valueMax="3150.0" valueMin="3120.0">
<elevation id="002B5D29FFF9FF93FD67FA4AA210FA73" box="[643,790,1413,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.135" metricValueMax="3.15" metricValueMin="3.12" pageId="38" pageNumber="123" unit="m" value="3135.0" valueMax="3150.0" valueMin="3120.0">3120-3150 m</elevation>
</quantity>
, 
<date id="FFB89CDAFFF9FF93FCC2FA4AA2D9FA73" box="[806,991,1413,1438]" pageId="38" pageNumber="123" value="2005-05-17" valueMax="2005-05-19" valueMin="2005-05-17">
<collectingDate id="EFFC6532FFF9FF93FCC2FA4AA2D9FA73" box="[806,991,1413,1438]" pageId="38" pageNumber="123" value="2005-05-17" valueMax="2005-05-19" valueMin="2005-05-17">17-19 May 2005</collectingDate>
</date>
, 
<collectorName id="26F3DFCCFFF9FF93FC15FA49A588FA73" box="[1009,1166,1413,1437]" pageId="38" pageNumber="123">D. Kavanaugh</collectorName>
, 
<collectorName id="26F3DFCCFFF9FF93FF9FFA6AA011FA53" box="[123,279,1445,1469]" pageId="38" pageNumber="123">C.E. Griswold</collectorName>
, 
<collectorName id="26F3DFCCFFF9FF93FEC3FA69A0A4FA53" box="[295,418,1445,1469]" pageId="38" pageNumber="123">H.B. Liang</collectorName>
, 
<collectorName id="26F3DFCCFFF9FF93FE56FA69A32EFA50" box="[434,552,1446,1470]" pageId="38" pageNumber="123">D.Z. Dong</collectorName>
&amp; 
<collectorName id="26F3DFCCFFF9FF93FDB4FA69A3B5FA53" box="[592,691,1445,1469]" pageId="38" pageNumber="123">K.J. Guo</collectorName>
collectors [
<specimenCount id="9D007193FFF9FF93FCD3FA6AA29FFA53" box="[823,921,1445,1469]" count="22" pageId="38" pageNumber="123" type="male">22 males</specimenCount>
and 
<specimenCount id="9D007193FFF9FF93FC37FA6AA54CFA53" box="[979,1098,1445,1469]" count="11" pageId="38" pageNumber="123" type="female">11 females</specimenCount>
; 
<collectionCode id="ED1722DFFFF9FF93FBBFFA6AA589FA53" box="[1115,1167,1445,1469]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="38" pageNumber="123" type="Museum">CAS</collectionCode>
, 
<collectionCode id="ED1722DFFFF9FF93FF9FFA0AA1A1FA33" box="[123,167,1477,1501]" pageId="38" pageNumber="123">IOZ</collectionCode>
])
</materialsCitation>
.
</paragraph>
<paragraph id="8BB9BA1AFFF9FF93FF4FFA2AA595F9F3" blockId="38.[123,1174,1029,1726]" pageId="38" pageNumber="123">
<materialsCitation id="3B6EB047FFF9FF93FF4FFA2AA58AF9F3" county="Lushui County" location="Core Areas" municipality="Gaoligong" pageId="38" pageNumber="123" specimenCount="1" stateProvince="Shan">
Members of this species were collected only in 
<collectingCounty id="62D8C296FFF9FF93FD54FA2AA257FA13" box="[688,849,1509,1533]" pageId="38" pageNumber="123">Lushui County</collectingCounty>
on the crest of the 
<collectingMunicipality id="6BDD2060FFF9FF93FBC1FA2AA590FA13" box="[1061,1174,1509,1533]" pageId="38" pageNumber="123">Gaoligong</collectingMunicipality>
<collectingRegion id="49C274F8FFF9FF93FF9FF9CAA1B6F9F3" box="[123,176,1541,1565]" country="Myanmar" name="Shan" pageId="38" pageNumber="123">Shan</collectingRegion>
in the southcentral part of the study area, an area that staddles 
<location id="8ED9ECC1FFF9FF93FCB3F9CAA2C9F9F3" LSID="urn:lsid:plazi:treatment:03AF0B0CFFFBFF93FF9FFEF4A590F953:8ED9ECC1FFF9FF93FCB3F9CAA2C9F9F3" box="[855,975,1541,1565]" county="Lushui County" municipality="Gaoligong" name="Core Areas" pageId="38" pageNumber="123" stateProvince="Shan">Core Areas</location>
4 and 5 (
<figureCitation id="133DA69FFFF9FF93FBD3F9CAA585F9F3" box="[1079,1155,1541,1565]" captionStart="FIGURE" captionStartId="88.[171,183,995,1014]" captionTargetBox="[130,1164,233,969]" captionTargetId="figure-57@88.[120,1176,215,989]" captionTargetPageId="88" captionText="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." figureDoi="http://doi.org/10.5281/zenodo.11067455" httpUri="https://zenodo.org/record/11067455/files/figure.png" pageId="38" pageNumber="123">Fig. 48</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB9BA1AFFF9FF93FF4FF9EAA3BEF993" blockId="38.[123,1174,1029,1726]" pageId="38" pageNumber="123">
<emphasis id="B9726608FFF9FF93FF4FF9EAA328F9D3" bold="true" box="[171,558,1573,1597]" pageId="38" pageNumber="123">Overall geographical distribution.</emphasis>
<materialsCitation id="3B6EB047FFF9FF93FDD0F9EAA3B7F993" country="China" county="Lushui County" location="Gaoligong Shan" pageId="38" pageNumber="123" specimenCount="1" stateProvince="Yunnan">
This species currently is known only from the 
<typeStatus id="54BD04B8FFF9FF93FBC0F9E9A554F9D0" box="[1060,1106,1574,1598]" pageId="38" pageNumber="123">type</typeStatus>
locality and its immediate environs in 
<collectingCounty id="62D8C296FFF9FF93FE1BF98AA3A2F9B3" box="[511,676,1605,1629]" pageId="38" pageNumber="123">Lushui County</collectingCounty>
on the crest of the southcentral part of the 
<location id="8ED9ECC1FFF9FF93FF9FF9AAA02EF993" LSID="urn:lsid:plazi:treatment:03AF0B0CFFFBFF93FF9FFEF4A590F953:8ED9ECC1FFF9FF93FF9FF9AAA02EF993" box="[123,296,1637,1661]" country="China" county="Lushui County" name="Gaoligong Shan" pageId="38" pageNumber="123" stateProvince="Yunnan">Gaoligong Shan</location>
, in western 
<collectingRegion id="49C274F8FFF9FF93FE4FF9A9A365F993" box="[427,611,1637,1661]" country="China" name="Yunnan" pageId="38" pageNumber="123">Yunnan Province</collectingRegion>
, 
<collectingCountry id="F311FA8AFFF9FF93FD95F9AAA3B7F993" box="[625,689,1637,1661]" name="China" pageId="38" pageNumber="123">China</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB9BA1AFFF9FF93FF4FF94AA590F953" blockId="38.[123,1174,1029,1726]" pageId="38" pageNumber="123">
<emphasis id="B9726608FFF9FF93FF4FF94AA24DF973" bold="true" box="[171,843,1669,1693]" pageId="38" pageNumber="123">
Geographical relationships with other 
<taxonomicName id="4C06C199FFF9FF93FD83F949A24DF973" box="[615,843,1669,1693]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="123" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9726608FFF9FF93FD83F949A3E8F973" bold="true" box="[615,750,1670,1693]" italics="true" pageId="38" pageNumber="123">Broscosoma</emphasis>
species.
</taxonomicName>
</emphasis>
Members of this species have been found syntopic only with those of 
<taxonomicName id="4C06C199FFF9FF93FDF9F969A3E0F953" authorityName="Putzeys. We" authorityYear="1877" box="[541,742,1701,1725]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="123" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFF9FF93FDF9F969A3E0F953" box="[541,742,1701,1725]" italics="true" pageId="38" pageNumber="123">Broscosoma ribbei</emphasis>
</taxonomicName>
at and around the 
<typeStatus id="54BD04B8FFF9FF93FC4CF969A2D0F950" box="[936,982,1702,1726]" pageId="38" pageNumber="123">type</typeStatus>
locality (
<figureCitation id="133DA69FFFF9FF93FBDFF96AA580F953" box="[1083,1158,1701,1725]" captionStart="FIGURE" captionStartId="89.[171,183,1461,1480]" captionTargetBox="[140,1154,249,1428]" captionTargetId="figure-68@89.[120,1176,216,1448]" captionTargetPageId="89" captionText="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi="http://doi.org/10.5281/zenodo.11067459" httpUri="https://zenodo.org/record/11067459/files/figure.png" pageId="38" pageNumber="123">Fig. 50</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>