391 lines
58 KiB
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391 lines
58 KiB
XML
<document id="1BF121BFB1B9B2CAA50142059970FE23" ID-DOI="10.5281/zenodo.11067355" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067355" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714052343267" checkinUser="felipe" docAuthor="Kavanaugh, David H. & Liang, Hongbin" docDate="2021" docId="03AF0B0CFFC3FFAAFF9FFAB3A590F9DD" docLanguage="en" docName="ProcCalifAcadSci.67.4.85-182.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (4)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.1:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="1" docTitle="Broscosoma danzhuense Kavanaugh and Liang 2021, sp. nov." docType="treatment" docVersion="2" lastPageNumber="116" masterDocId="FF967374FFDFFFB5FFE4FFCFA106FFEE" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae)." masterLastPageNumber="182" masterPageNumber="85" pageNumber="113" updateTime="1714052772053" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="7E663E3A5472BA62D13BFC39C7550DE9">
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<mods:title id="94B742FB6569DAACCAAB7EA899A4D486">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae).</mods:title>
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<mods:name id="328862CF29FC14A39C5199CC6449BA0E" type="personal">
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<mods:role id="FC69658327797B058276305277479878">
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<mods:roleTerm id="8A76839FDC9C7C1E1F3541633CEAFED3">Author</mods:roleTerm>
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<mods:namePart id="C3765CD8CAFA8EAF08A7BD787C90069F">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="FA08E3A30DD352C5CF7E510915F9F953">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:name id="962175C0549945C84C82DE81FC704FFC" type="personal">
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<mods:roleTerm id="B0A8A5F898B6BB9FF25373321DE41440">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="913F39CA79CC2B71325AB8E49443D9CF">Liang, Hongbin</mods:namePart>
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<mods:affiliation id="FF0778EEB75F510D160CEF38F036293B">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:typeOfResource id="AF3F6FC8DBA63A78363D6D7987F6048B">text</mods:typeOfResource>
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<mods:title id="16A18922A9875C3096F762FB384436CA">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="0200A48668DFCF102682BB1CFD28DB55">
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<mods:date id="8CB3C751FCAF3922B1E53AF5D160FEA7">2021</mods:date>
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<mods:number id="1A05071CEC4C253F7D6D4414E3058BCD">2021-09-30</mods:number>
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<mods:number id="DE9F6F04B1BFB436A30CC0F3B60323BE">67</mods:number>
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<mods:number id="4F773A3961DCCA89F19A0C21C636EC22">4</mods:number>
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<mods:start id="2C66C4E19F86604D9DE610F75B28B558">85</mods:start>
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<mods:identifier id="F55239029D9D6BC5BD327B8BFFA256E5" type="DOI">10.5281/zenodo.11067355</mods:identifier>
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<mods:identifier id="5794F01F6081C665CF65931A790A3529" type="ISSN">0068-547X</mods:identifier>
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<treatment id="03AF0B0CFFC3FFAAFF9FFAB3A590F9DD" ID-DOI="http://doi.org/10.5281/zenodo.11067403" ID-Zenodo-Dep="11067403" LSID="urn:lsid:plazi:treatment:03AF0B0CFFC3FFAAFF9FFAB3A590F9DD" httpUri="http://treatment.plazi.org/id/03AF0B0CFFC3FFAAFF9FFAB3A590F9DD" lastPageId="31" lastPageNumber="116" pageId="28" pageNumber="113">
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<subSubSection id="C31CE991FFC3FFA9FF9FFAB3A214FA7A" box="[123,786,1403,1428]" pageId="28" pageNumber="113" type="nomenclature">
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<heading id="D0F10D76FFC3FFA9FF9FFAB3A214FA7A" box="[123,786,1403,1428]" fontSize="10" level="2" pageId="28" pageNumber="113" reason="7">
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<emphasis id="B9726608FFC3FFA9FF9FFAB3A214FA7A" bold="true" box="[123,786,1403,1428]" pageId="28" pageNumber="113">
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4.
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<taxonomicName id="4C06C199FFC3FFA9FF72FAB3A3A8FA7A" authority="Kavanaugh and Liang" authorityName="Kavanaugh and Liang" authorityYear="2021" box="[150,686,1403,1428]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="113" phylum="Arthropoda" rank="species" species="danzhuense" status="sp. nov.">
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<emphasis id="B9726608FFC3FFA9FF72FAB3A0A0FA7D" bold="true" box="[150,422,1403,1427]" italics="true" pageId="28" pageNumber="113">Broscosoma danzhuense</emphasis>
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Kavanaugh and Liang
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</taxonomicName>
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,
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<taxonomicNameLabel id="A241DB73FFC3FFA9FD58FAB3A214FA7A" box="[700,786,1404,1428]" pageId="28" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C31CE991FFC3FFA9FF9FFA53A068FA5C" box="[123,366,1436,1458]" pageId="28" pageNumber="113" type="description">
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<paragraph id="8BB9BA1AFFC3FFA9FF9FFA53A068FA5C" blockId="28.[123,786,1403,1458]" box="[123,366,1436,1458]" pageId="28" pageNumber="113">
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<figureCitation id="133DA69FFFC3FFA9FF9FFA53A1E6FA5F" box="[123,224,1436,1458]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="28" pageNumber="113">Figures 6e</figureCitation>
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,
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<figureCitation id="133DA69FFFC3FFA9FF09FA53A003FA5F" box="[237,261,1436,1457]" captionStart="FIGURE" captionStartId="29.[171,183,1612,1631]" captionTargetBox="[120,1176,216,1596]" captionTargetId="figure-68@29.[120,1176,216,1598]" captionTargetPageId="29" captionText="FIGURE 17. Broscosoma danzhuense sp. nov. a. Habitus (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067385" httpUri="https://zenodo.org/record/11067385/files/figure.png" pageId="28" pageNumber="113">17</figureCitation>
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,
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<figureCitation id="133DA69FFFC3FFA9FEF5FA53A02FFA5F" box="[273,297,1436,1457]" captionStart="FIGURE" captionStartId="83.[171,183,1575,1594]" captionTargetBox="[197,1099,216,1560]" captionTargetId="figure-67@83.[197,1099,216,1560]" captionTargetPageId="83" captionText="FIGURE 43. Photographs of habitat for broscine species in the Gaoligong Shan region; a. Danzhu He drainage, 2840 m, 27.633°/98.604°, Cikai Township, Gongshan County, Yunnan (habitat for Broscosoma danzhuense sp. nov. and Eobroscus bhtanensis Morvan). Photo by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067445" httpUri="https://zenodo.org/record/11067445/files/figure.png" pageId="28" pageNumber="113">43</figureCitation>
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,
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<figureCitation id="133DA69FFFC3FFA9FED2FA53A068FA5C" box="[310,366,1436,1458]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="88.[171,183,995,1014]" captionStartId-1="88.[184,196,1682,1701]" captionStartId-2="89.[171,183,1461,1480]" captionTargetBox-0="[130,1164,233,969]" captionTargetBox-1="[130,1162,1063,1660]" captionTargetBox-2="[140,1154,249,1428]" captionTargetId-0="figure-57@88.[120,1176,215,989]" captionTargetId-1="figure-332@88.[120,1176,1054,1668]" captionTargetId-2="figure-68@89.[120,1176,216,1448]" captionTargetPageId-0="88" captionTargetPageId-1="88" captionTargetPageId-2="89" captionText-0="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." captionText-2="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi-0="http://doi.org/10.5281/zenodo.11067455" figureDoi-1="http://doi.org/10.5281/zenodo.11067457" figureDoi-2="http://doi.org/10.5281/zenodo.11067459" httpUri-0="https://zenodo.org/record/11067455/files/figure.png" httpUri-1="https://zenodo.org/record/11067457/files/figure.png" httpUri-2="https://zenodo.org/record/11067459/files/figure.png" pageId="28" pageNumber="113">48-50</figureCitation>
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</paragraph>
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<subSubSection id="C31CE991FFC3FFABFF4FFA0AA590FEFD" lastPageId="30" lastPageNumber="115" pageId="28" pageNumber="113" type="materials_examined">
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<paragraph id="8BB9BA1AFFC3FFA9FF4FFA0AA3E9F993" blockId="28.[123,1173,1477,1726]" pageId="28" pageNumber="113">
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<emphasis id="B9726608FFC3FFA9FF4FFA0AA056FA33" bold="true" box="[171,336,1477,1501]" pageId="28" pageNumber="113">Type material.</emphasis>
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<materialsCitation id="3B6EB047FFC3FFA9FEB3FA0AA3E9F993" collectingDate="2000-06-30" collectingDateMax="2000-07-05" collectingDateMin="2000-06-30" collectionCode="IOZ" collectorName="C. E. Groswold & D. - Z. Collectors" country="China" county="Gongshan County" elevation="2780" latitude="27.63267" location="Nujiang Prefecture" longLatPrecision="1" longitude="98.60861" municipality="Gaoligong Shan" pageId="28" pageNumber="113" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="54BD04B8FFC3FFA9FEB3FA0AA0BCFA33" box="[343,442,1477,1501]" pageId="28" pageNumber="113" type="holotype">Holotype</typeStatus>
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, a female, in
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<collectionCode id="ED1722DFFFC3FFA9FDAFFA0AA371FA33" box="[587,631,1477,1501]" pageId="28" pageNumber="113">IOZ</collectionCode>
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, labeled: “ CASENT 1004637”/ “
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<collectingCountry id="F311FA8AFFC3FFA9FC07FA0AA531FA33" box="[995,1079,1477,1501]" name="China" pageId="28" pageNumber="113">CHINA</collectingCountry>
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,
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<collectingRegion id="49C274F8FFC3FFA9FBA0FA09A1DCFA13" country="China" name="Yunnan" pageId="28" pageNumber="113">Yunnan Province</collectingRegion>
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,
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<collectingMunicipality id="6BDD2060FFC3FFA9FF03FA2AA095FA13" box="[231,403,1509,1533]" pageId="28" pageNumber="113">Gaoligong Shan</collectingMunicipality>
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,
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<location id="8ED9ECC1FFC3FFA9FE45FA2AA36AFA13" LSID="urn:lsid:plazi:treatment:03AF0B0CFFC3FFAAFF9FFAB3A590F9DD:8ED9ECC1FFC3FFA9FE45FA2AA36AFA13" box="[417,620,1509,1533]" country="China" county="Gongshan County" latitude="27.63267" longLatPrecision="1" longitude="98.60861" municipality="Gaoligong Shan" name="Nujiang Prefecture" pageId="28" pageNumber="113" stateProvince="Yunnan">Nujiang Prefecture</location>
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,
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<collectingCounty id="62D8C296FFC3FFA9FD9DFA2AA23FFA13" box="[633,825,1509,1533]" pageId="28" pageNumber="113">Gongshan County</collectingCounty>
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,
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<location id="8ED9ECC1FFC3FFA9FCA2FA2AA2B9FA13" LSID="urn:lsid:plazi:treatment:03AF0B0CFFC3FFAAFF9FFAB3A590F9DD:8ED9ECC1FFC3FFA9FCA2FA2AA2B9FA13" box="[838,959,1509,1533]" country="China" county="Gongshan County" latitude="27.63267" longLatPrecision="1" longitude="98.60861" municipality="Gaoligong Shan" name="Danzhu He" pageId="28" pageNumber="113" stateProvince="Yunnan">Danzhu He</location>
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drainage, 13.5-13.8 airkm
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<locationDeviation id="9A0BA338FFC3FFA9FF32F9CAA0CFF9F3" box="[214,457,1541,1565]" location="Gongshan" pageId="28" pageNumber="113">SSW of Gongshan</locationDeviation>
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[
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<location id="8ED9ECC1FFC3FFA9FE09F9CAA32FF9F3" LSID="urn:lsid:plazi:treatment:03AF0B0CFFC3FFAAFF9FFAB3A590F9DD:8ED9ECC1FFC3FFA9FE09F9CAA32FF9F3" box="[493,553,1541,1565]" country="China" county="Gongshan County" latitude="27.63267" longLatPrecision="1" longitude="98.60861" municipality="Gaoligong Shan" name="Cikai" pageId="28" pageNumber="113" stateProvince="Yunnan">Cikai</location>
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],
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<quantity id="4CFE17FFFFC3FFA9FDB0F9CAA3FAF9F3" box="[596,764,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="2.78" metricValueMax="2.84" metricValueMin="2.7199999999999998" pageId="28" pageNumber="113" unit="m" value="2780.0" valueMax="2840.0" valueMin="2720.0">
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<elevation id="002B5D29FFC3FFA9FDB0F9CAA3FAF9F3" box="[596,764,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="2.78" metricValueMax="2.84" metricValueMin="2.7199999999999998" pageId="28" pageNumber="113" unit="m" value="2780.0" valueMax="2840.0" valueMin="2720.0">2720-2840 m</elevation>
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</quantity>
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,”/ “
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<geoCoordinate id="EE32DCDDFFC3FFA9FCDAF9C9A2C3F9F3" box="[830,965,1541,1565]" degrees="27. 63267" direction="north" orientation="latitude" pageId="28" pageNumber="113" precision="1" value="27.63267">N27.63267°</geoCoordinate>
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/
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<geoCoordinate id="EE32DCDDFFC3FFA9FC29F9C9A557F9F3" box="[973,1105,1541,1565]" degrees="98. 60861" direction="east" orientation="longitude" pageId="28" pageNumber="113" precision="1" value="98.60861">E98.60861°</geoCoordinate>
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to”/ “
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<geoCoordinate id="EE32DCDDFFC3FFA9FF63F9E9A00FF9D3" box="[135,265,1573,1597]" degrees="27. 63331" direction="north" orientation="latitude" pageId="28" pageNumber="113" precision="1" value="27.63331">N27.63331°</geoCoordinate>
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/
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<geoCoordinate id="EE32DCDDFFC3FFA9FEF4F9E9A089F9D3" box="[272,399,1573,1597]" degrees="98. 60356" direction="east" orientation="longitude" pageId="28" pageNumber="113" precision="1" value="98.60356">E98.60356°</geoCoordinate>
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,
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<date id="FFB89CDAFFC3FFA9FE79F9EAA38FF9D3" box="[413,649,1573,1597]" pageId="28" pageNumber="113" value="2000-06-30" valueMax="2000-07-05" valueMin="2000-06-30">
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<collectingDate id="EFFC6532FFC3FFA9FE79F9EAA38FF9D3" box="[413,649,1573,1597]" pageId="28" pageNumber="113" value="2000-06-30" valueMax="2000-07-05" valueMin="2000-06-30">30 June – 5 July 2000</collectingDate>
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</date>
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, Stop#00-17E, D.H. Kavanaugh,
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<collectorName id="26F3DFCCFFC3FFA9FC11F9EAA593F9D3" box="[1013,1173,1573,1597]" pageId="28" pageNumber="113">C.E. Groswold</collectorName>
|
||
[sic], Liang H.-B., D. Ubick, & Dong
|
||
<collectorName id="26F3DFCCFFC3FFA9FDF8F989A3CAF9B3" box="[540,716,1605,1629]" pageId="28" pageNumber="113">D.-Z. Collectors</collectorName>
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”./ “
|
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<typeStatus id="54BD04B8FFC3FFA9FD1DF98AA280F9B3" box="[761,902,1605,1629]" pageId="28" pageNumber="113" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4C06C199FFC3FFA9FC6AF989A050F993" authority="Kavanaugh & Liang" authorityName="Kavanaugh & Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="113" phylum="Arthropoda" rank="species" species="danzhuense" status="sp. nov.">
|
||
<emphasis id="B9726608FFC3FFA9FC6AF989A593F9B3" box="[910,1173,1605,1629]" italics="true" pageId="28" pageNumber="113">Broscosoma danzhuense</emphasis>
|
||
Kavanaugh & Liang
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A241DB73FFC3FFA9FEB9F9A9A0B6F993" box="[349,432,1638,1661]" pageId="28" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
|
||
designated 2021” [red label].
|
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</materialsCitation>
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</paragraph>
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<paragraph id="8BB9BA1AFFC3FFA9FF4FF949A1D5F953" blockId="28.[123,1173,1477,1726]" pageId="28" pageNumber="113">
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<materialsCitation id="3B6EB047FFC3FFA9FF4FF949A1CAF953" country="China" county="Gongshan County" elevation="2780" location="Danzhu He" municipality="Cikai Township" pageId="28" pageNumber="113" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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||
<emphasis id="B9726608FFC3FFA9FF4FF949A03AF973" bold="true" box="[171,316,1669,1694]" pageId="28" pageNumber="113">
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<typeStatus id="54BD04B8FFC3FFA9FF4FF949A1E5F970" box="[171,227,1670,1694]" pageId="28" pageNumber="113">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<collectingCountry id="F311FA8AFFC3FFA9FEADF94AA08FF973" box="[329,393,1669,1693]" name="China" pageId="28" pageNumber="113">China</collectingCountry>
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||
,
|
||
<collectingRegion id="49C274F8FFC3FFA9FE71F949A0E0F973" box="[405,486,1670,1693]" country="China" name="Yunnan" pageId="28" pageNumber="113">Yunnan</collectingRegion>
|
||
,
|
||
<collectingCounty id="62D8C296FFC3FFA9FE10F94AA3B5F970" box="[500,691,1669,1694]" pageId="28" pageNumber="113">Gongshan County</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BDD2060FFC3FFA9FD24F94AA26FF973" box="[704,873,1669,1693]" pageId="28" pageNumber="113">Cikai Township</collectingMunicipality>
|
||
,
|
||
<location id="8ED9ECC1FFC3FFA9FC92F94AA2E8F973" LSID="urn:lsid:plazi:treatment:03AF0B0CFFC3FFAAFF9FFAB3A590F9DD:8ED9ECC1FFC3FFA9FC92F94AA2E8F973" box="[886,1006,1669,1693]" country="China" county="Gongshan County" municipality="Cikai Township" name="Danzhu He" pageId="28" pageNumber="113" stateProvince="Yunnan">Danzhu He</location>
|
||
drainage
|
||
<quantity id="4CFE17FFFFC3FFA9FBB3F949A1CAF953" metricMagnitude="3" metricUnit="m" metricValue="2.78" metricValueMax="2.84" metricValueMin="2.7199999999999998" pageId="28" pageNumber="113" unit="m" value="2780.0" valueMax="2840.0" valueMin="2720.0">
|
||
<elevation id="002B5D29FFC3FFA9FBB3F949A1CAF953" metricMagnitude="3" metricUnit="m" metricValue="2.78" metricValueMax="2.84" metricValueMin="2.7199999999999998" pageId="28" pageNumber="113" unit="m" value="2780.0" valueMax="2840.0" valueMin="2720.0">2720- 2840 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF79EA92FFC2FFA8FF4FF983A334F97C" ID-DOI="http://doi.org/10.5281/zenodo.11067385" ID-Zenodo-Dep="11067385" httpUri="https://zenodo.org/record/11067385/files/figure.png" pageId="29" pageNumber="114" startId="29.[171,183,1612,1631]" targetBox="[120,1176,216,1596]" targetPageId="29" targetType="figure">
|
||
<paragraph id="8BB9BA1AFFC2FFA8FF4FF983A334F97C" blockId="29.[123,1174,1612,1682]" pageId="29" pageNumber="114">
|
||
<smallCapsWord id="8D5F2CC6FFC2FFA8FF4FF983A1EEF9B3" baselines="1626,1625" box="[171,232,1612,1631]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="29" pageNumber="114">FIGURE</smallCapsWord>
|
||
17.
|
||
<taxonomicName id="4C06C199FFC2FFA8FEE8F983A0D8F9B1" authority="Kavanaugh & Liang, 2021" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[268,478,1612,1631]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="114" phylum="Arthropoda" rank="species" species="danzhuense" status="sp. nov.">
|
||
<emphasis id="B9726608FFC2FFA8FEE8F983A0D8F9B1" box="[268,478,1612,1631]" italics="true" pageId="29" pageNumber="114">Broscosoma danzhuense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A241DB73FFC2FFA8FE06F983A325F9B1" box="[482,547,1612,1631]" pageId="29" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
|
||
a. Habitus (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB9BA1AFFC1FFABFF4FFF14A590FEFD" blockId="30.[123,1174,219,1726]" pageId="30" pageNumber="115">
|
||
<emphasis id="B9726608FFC1FFABFF4FFF14A0E7FF1D" bold="true" box="[171,481,219,243]" pageId="30" pageNumber="115">Derivation of species name.</emphasis>
|
||
The species epithet,
|
||
<taxonomicName id="4C06C199FFC1FFABFD5BFF14A23DFF1D" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[703,827,219,243]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="115" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC1FFABFD5BFF14A23DFF1D" box="[703,827,219,243]" italics="true" pageId="30" pageNumber="115">danzhuense</emphasis>
|
||
</taxonomicName>
|
||
, is derived from the name of the river valley in which the
|
||
<typeStatus id="54BD04B8FFC1FFABFE6CFF33A0B0FEFA" box="[392,438,252,276]" pageId="30" pageNumber="115">type</typeStatus>
|
||
specimen was collected, and the Latin suffix,
|
||
<emphasis id="B9726608FFC1FFABFC42FF33A2E2FEFD" box="[934,996,252,275]" italics="true" pageId="30" pageNumber="115">-ensis</emphasis>
|
||
, denoting place.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CE991FFC1FFABFF4FFED4A310FDDB" pageId="30" pageNumber="115" type="diagnosis">
|
||
<paragraph id="8BB9BA1AFFC1FFABFF4FFED4A310FDDB" blockId="30.[123,1174,219,1726]" pageId="30" pageNumber="115">
|
||
<emphasis id="B9726608FFC1FFABFF4FFED4A019FEDA" bold="true" box="[171,287,283,308]" pageId="30" pageNumber="115">Diagnosis.</emphasis>
|
||
Adults of this species (
|
||
<figureCitation id="133DA69FFFC1FFABFDC5FED4A37CFEDD" box="[545,634,283,308]" captionStart="FIGURE" captionStartId="29.[171,183,1612,1631]" captionTargetBox="[120,1176,216,1596]" captionTargetId="figure-68@29.[120,1176,216,1598]" captionTargetPageId="29" captionText="FIGURE 17. Broscosoma danzhuense sp. nov. a. Habitus (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067385" httpUri="https://zenodo.org/record/11067385/files/figure.png" pageId="30" pageNumber="115">Fig. 17a</figureCitation>
|
||
) can be distinguished from those of other species in the region by the following combination of character states: size moderate, BL female =
|
||
<quantity id="4CFE17FFFFC1FFABFBD9FEF3A58BFEBD" box="[1085,1165,316,340]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="30" pageNumber="115" unit="mm" value="9.3">9.3 mm</quantity>
|
||
; entire dorsum of body dark green, legs with femora darker (brown to piceous) than tibiae (reddish-brown); antennomeres 3 and 4 glabrous except for apical whorl of setae; eyes (
|
||
<figureCitation id="133DA69FFFC1FFABFC2AFEB3A51FFE7A" box="[974,1049,380,404]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="30" pageNumber="115">Fig. 6e</figureCitation>
|
||
) moderately convex; pronotum with lateral margination absent, basolateral setae absent; elytral silhouette subovoid, greatest width distinctly posterior to midlength, with humeri rounded yet slightly evident, elytral microsculpture faintly impressed but evident, only striae 1 impressed, all other elytral striae represented only be punctures diminishing in size and depth laterally and posteriorly, elytral intervals flat; metatrochanters asetose.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CE991FFC1FFAAFF4FFDF2A0FDFEBD" lastPageId="31" lastPageNumber="116" pageId="30" pageNumber="115" type="description">
|
||
<paragraph id="8BB9BA1AFFC1FFABFF4FFDF2A28EFD58" blockId="30.[123,1174,219,1726]" pageId="30" pageNumber="115">
|
||
<emphasis id="B9726608FFC1FFABFF4FFDF2A030FDB8" bold="true" box="[171,310,573,598]" pageId="30" pageNumber="115">Description.</emphasis>
|
||
<figureCitation id="133DA69FFFC1FFABFEDBFDF2A09CFDBB" box="[319,410,573,598]" captionStart="FIGURE" captionStartId="29.[171,183,1612,1631]" captionTargetBox="[120,1176,216,1596]" captionTargetId="figure-68@29.[120,1176,216,1598]" captionTargetPageId="29" captionText="FIGURE 17. Broscosoma danzhuense sp. nov. a. Habitus (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067385" httpUri="https://zenodo.org/record/11067385/files/figure.png" pageId="30" pageNumber="115">Fig. 17a</figureCitation>
|
||
. Size medium, BL female =
|
||
<quantity id="4CFE17FFFFC1FFABFD04FDF1A232FDBB" box="[736,820,574,598]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="30" pageNumber="115" unit="mm" value="9.3">9.3 mm</quantity>
|
||
, ratio EL/PL = 2.3. Body color piceous. Last visible abdominal sternite dark or rufous brown, antennae with all antennomeres rufous brown, femora brown or piceous, tibiae and tarsi reddish brown. Head, Pronotum, elytra with distinct green metallic reflection, venter without metallic reflection.
|
||
</paragraph>
|
||
<paragraph id="8BB9BA1AFFC1FFABFF4FFD71A00FFBF6" blockId="30.[123,1174,219,1726]" pageId="30" pageNumber="115">
|
||
Head.
|
||
<figureCitation id="133DA69FFFC1FFABFF15FD71A03DFD38" box="[241,315,702,726]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="30" pageNumber="115">Fig. 6e</figureCitation>
|
||
. Eyes small, their diameter slightly less than length of tempora. Frontal furrows deeply impressed, narrow, arcuate, slightly convergent anteriorly and posteriorly, long, extended posteriorly beyond middle of eyes, faintly rugulose, impunctate. Vertex with post-temporal transverse sulcus deep, sharply and narrowly defined, narrowly punctulate only along bottom of sulcus. Tempora nearly straight, subparallel. Genal ridges present, restricted to head region distinctly behind post-temporal constriction. Clypeus with one pair of setae. Supraorbital setae present, one pair. Eustipes of maxilla with two setae, the dorsobasal seta distinctly more than half as long as ventrobasal seta.
|
||
<collectingRegion id="49C274F8FFC1FFABFED5FC6FA08DFC59" box="[305,395,928,951]" country="Myanmar" name="Chin" pageId="30" pageNumber="115">Mentum</collectingRegion>
|
||
with tooth present, simple, paramedial region shallowly foveate, one pair of mental setae present. Glosal sclerite (ligula) with one pair of setae. Submentum with two or three pairs of setae. Gula without transverse grooves. Antennomeres 3 and 4 without pubescence, ratio A3/A5 = 1.3.
|
||
</paragraph>
|
||
<paragraph id="8BB9BA1AFFC1FFABFF4FFBEEA25CFAF4" blockId="30.[123,1174,219,1726]" pageId="30" pageNumber="115">
|
||
Pronotum.
|
||
<figureCitation id="133DA69FFFC1FFABFEC1FBEEA072FBD6" box="[293,372,1057,1081]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="30" pageNumber="115">Fig. 6e</figureCitation>
|
||
. Discal region anterior to sub-basal constriction slightly longitudinally ovoid, slightly narrowed anteriorly with base distinctly pedunculate, widest slightly posterior to middle of discal region, ratio PL/PW = 1.2, disk markedly convex; apical, lateral, and basal margination absent; anterior transverse impression shallow, narrowly delineated, punctate in bottom of impression only; median longitudinal impression distinct, but fine and shallowly impressed; posterior transverse impression deep, sharply delineated; anterior region smooth; pronotal base coarsely and densely punctate; one pair of midlateral pronotal setae present, inserted slightly anterior to mid-length of discal region; basolateral pronotal setae absent.
|
||
</paragraph>
|
||
<paragraph id="8BB9BA1AFFC1FFABFF4FFAECA3D5F992" blockId="30.[123,1174,219,1726]" pageId="30" pageNumber="115">Elytra. Elytral silhouette subovoid, slightly narrowed in basal half, widest distinctly posterior to middle, ratio EL/EW = 1.6, greatest elevation above lateral margin (in lateral view) posterior to middle; humeri distinctly present, roundly obtuse; lateral margins moderately arcuate and nearly straight in anterior half sub-basally; parascutellar striole present, short, distally merged in a distinct jog with free base of stria 1. Eight elytral striae present; stria 1 moderately deeply and sharply impressed, striae 2 to 8 indistinct, obliterated or represented by extremely fine and shallow linear depressions, successively less evident laterally and apically, striae 8 merged with lateral groove throughout its length or nearly so; stria 1 moderately punctate at least in basal half, striae 2 to 8 successively more finely and sparsely punctate or impunctate laterally and apically. Elytral microsculpture effaced. Parascutellar seta at base of stria 2, discal setae absent, umbilicate series comprised of one post-humeral and two preapical setae.</paragraph>
|
||
<paragraph id="8BB9BA1AFFC1FFABFF4FF94AA237F973" blockId="30.[123,1174,219,1726]" box="[171,817,1669,1693]" pageId="30" pageNumber="115">Hindwings. Slightly reduced, incapable of supporting flight.</paragraph>
|
||
<paragraph id="8BB9BA1AFFC1FFABFF4FF96AA3E1F953" blockId="30.[123,1174,219,1726]" box="[171,743,1701,1726]" pageId="30" pageNumber="115">Thoracic venter. Metepisternum ratio ML/MW = 2.4.</paragraph>
|
||
<paragraph id="8BB9BA1AFFC0FFAAFF4FFF14A0D6FEFD" blockId="31.[123,1174,219,1588]" pageId="31" pageNumber="116">Legs. Hind trochanter asetose. Tarsomeres without dorsal pubescence; tarsomere 5 with two or more pairs of setae ventrally.</paragraph>
|
||
<paragraph id="8BB9BA1AFFC0FFAAFF4FFED4A291FEDD" blockId="31.[123,1174,219,1588]" box="[171,919,283,307]" pageId="31" pageNumber="116">Abdomen. Female sternite VI with one pair of apicoparamedial setae.</paragraph>
|
||
<paragraph id="8BB9BA1AFFC0FFAAFF4FFEF4A0FDFEBD" blockId="31.[123,1174,219,1588]" box="[171,507,315,339]" pageId="31" pageNumber="116">Male genitalia. Male unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CE991FFC0FFAAFF4FFE94A590F9DD" pageId="31" pageNumber="116" type="discussion">
|
||
<paragraph id="8BB9BA1AFFC0FFAAFF4FFE94A3CEFC7D" blockId="31.[123,1174,219,1588]" pageId="31" pageNumber="116">
|
||
<emphasis id="B9726608FFC0FFAAFF4FFE94A02BFE9D" bold="true" box="[171,301,347,371]" pageId="31" pageNumber="116">Comments.</emphasis>
|
||
This is one of six species in the study area members of which have the elytral humeri rounded or slightly angulate but distinct and both the forebody and the elytra with distinct metallic reflection. In two of these,
|
||
<taxonomicName id="4C06C199FFC0FFAAFE1DFE53A3CEFE5D" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[505,712,411,435]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="holomarginatum">
|
||
<emphasis id="B9726608FFC0FFAAFE1DFE53A3CEFE5D" box="[505,712,411,435]" italics="true" pageId="31" pageNumber="116">B. holomarginatum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C06C199FFC0FFAAFD19FE53A297FE5D" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[765,913,412,435]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="purpureum">
|
||
<emphasis id="B9726608FFC0FFAAFD19FE53A297FE5D" box="[765,913,412,435]" italics="true" pageId="31" pageNumber="116">B. purpureum</emphasis>
|
||
</taxonomicName>
|
||
, the reflection is mainly blue or purple in color, respectively, with some greenish tinges in most specimens of
|
||
<taxonomicName id="4C06C199FFC0FFAAFBF0FE73A1D4FE1D" authorityName="Kavanaugh & Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="holomarginatum">
|
||
<emphasis id="B9726608FFC0FFAAFBF0FE73A1D4FE1D" italics="true" pageId="31" pageNumber="116">B. holomarginatum</emphasis>
|
||
</taxonomicName>
|
||
. In addition to
|
||
<taxonomicName id="4C06C199FFC0FFAAFE9FFE13A30DFE1D" box="[379,523,475,499]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuens">
|
||
<emphasis id="B9726608FFC0FFAAFE9FFE13A30DFE1D" box="[379,523,475,499]" italics="true" pageId="31" pageNumber="116">B. danzhuens</emphasis>
|
||
</taxonomicName>
|
||
e, three species have the full dorsum of the body with green metallic reflection.
|
||
<taxonomicName id="4C06C199FFC0FFAAFEA9FE33A32EFDFD" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[333,552,508,531]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="parvum">
|
||
<emphasis id="B9726608FFC0FFAAFEA9FE33A32EFDFD" box="[333,552,508,531]" italics="true" pageId="31" pageNumber="116">Broscosoma parvum</emphasis>
|
||
</taxonomicName>
|
||
members are smaller in size, have eyes that are less convex (
|
||
<figureCitation id="133DA69FFFC0FFAAFF56FDD4A1FDFDDD" box="[178,251,539,563]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="31" pageNumber="116">Fig. 6f</figureCitation>
|
||
), and have the elytral silhouette slightly narrower across the humeri and the greatest width at or very near midlength (distinctly posterior to midlength in
|
||
<taxonomicName id="4C06C199FFC0FFAAFC84FDF3A2FDFDBD" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[864,1019,571,595]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC0FFAAFC84FDF3A2FDFDBD" box="[864,1019,571,595]" italics="true" pageId="31" pageNumber="116">B. danzhuense</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C06C199FFC0FFAAFBF6FDF3A1D2FD9D" authorityName="Kavanaugh & Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="resbecqi">
|
||
<emphasis id="B9726608FFC0FFAAFBF6FDF3A1D2FD9D" italics="true" pageId="31" pageNumber="116">Broscosoma resbecqi</emphasis>
|
||
</taxonomicName>
|
||
members have the pronotum with the discal portion relatively shorter and more globose (
|
||
<figureCitation id="133DA69FFFC0FFAAFF60FDB4A1D6FD7D" box="[132,208,635,659]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="31" pageNumber="116">Fig. 6d</figureCitation>
|
||
) and lateral margination present and nearly complete, interrupted only briefly near basal two-thirds length (
|
||
<figureCitation id="133DA69FFFC0FFAAFEA0FD54A097FD5D" box="[324,401,667,691]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="31" pageNumber="116">Fig. 9b</figureCitation>
|
||
) (absent from
|
||
<taxonomicName id="4C06C199FFC0FFAAFDD7FD53A3C8FD5D" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[563,718,667,691]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC0FFAAFDD7FD53A3C8FD5D" box="[563,718,667,691]" italics="true" pageId="31" pageNumber="116">B. danzhuense</emphasis>
|
||
</taxonomicName>
|
||
), the elytra with their greatest width at or very near midlength, and femora pale rufous (darker in
|
||
<taxonomicName id="4C06C199FFC0FFAAFD37FD73A268FD3D" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[723,878,699,723]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC0FFAAFD37FD73A268FD3D" box="[723,878,699,723]" italics="true" pageId="31" pageNumber="116">B. danzhuense</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C06C199FFC0FFAAFC61FD73A556FD3D" authorityName="Putzeys. We" authorityYear="1877" box="[901,1104,699,723]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="ribbei">
|
||
<emphasis id="B9726608FFC0FFAAFC61FD73A556FD3D" box="[901,1104,699,723]" italics="true" pageId="31" pageNumber="116">Broscosoma ribbei</emphasis>
|
||
</taxonomicName>
|
||
members have the eyes distinctly more convex, the post-temporal transverse sulcus not or more sparsely punctate, the pronotum with the discal portion relatively shorter and more globose (
|
||
<figureCitation id="133DA69FFFC0FFAAFBF0FD34A559FCFD" box="[1044,1119,763,787]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="31" pageNumber="116">Fig. 6b</figureCitation>
|
||
) and lateral margination incomplete but present in most specimens, at least in the vicinity of the midlateral setae and more extensively in some specimens, elytral microsculpture distinct, deeply to moderately impressed (effaced in
|
||
<taxonomicName id="4C06C199FFC0FFAAFE08FC93A381FC9D" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[492,647,859,883]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC0FFAAFE08FC93A381FC9D" box="[492,647,859,883]" italics="true" pageId="31" pageNumber="116">B. danzhuense</emphasis>
|
||
</taxonomicName>
|
||
), and the metatrochanters unisetose, unilaterally asetose in a few specimens (asetose in
|
||
<taxonomicName id="4C06C199FFC0FFAAFDF9FCB3A3BEFC7D" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[541,696,891,915]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC0FFAAFDF9FCB3A3BEFC7D" box="[541,696,891,915]" italics="true" pageId="31" pageNumber="116">B. danzhuense</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB9BA1AFFC0FFAAFF4FFC53A305FB5D" blockId="31.[123,1174,219,1588]" pageId="31" pageNumber="116">
|
||
Among
|
||
<taxonomicName id="4C06C199FFC0FFAAFEE6FC53A0DCFC5D" box="[258,474,923,947]" pageId="31" pageNumber="116">
|
||
<emphasis id="B9726608FFC0FFAAFEE6FC53A083FC5D" box="[258,389,924,947]" italics="true" pageId="31" pageNumber="116">Broscosoma</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
known from outside the study area, only two species,
|
||
<taxonomicName id="4C06C199FFC0FFAAFBCEFC53A0ADFC3D" authority="Deuve, 2006" authorityName="Deuve" authorityYear="2006" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="montreuili">
|
||
<emphasis id="B9726608FFC0FFAAFBCEFC53A01CFC3D" italics="true" pageId="31" pageNumber="116">Broscosoma montreuili</emphasis>
|
||
Deuve, 2006
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C06C199FFC0FFAAFE06FC73A236FC3D" authority="Deuve, 2006" authorityName="Deuve" authorityYear="2006" box="[482,816,955,979]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="tiani">
|
||
<emphasis id="B9726608FFC0FFAAFE06FC73A398FC3D" box="[482,670,956,979]" italics="true" pageId="31" pageNumber="116">Broscosoma tiani</emphasis>
|
||
Deuve, 2006
|
||
</taxonomicName>
|
||
have members in which the elytral humeri are at least faintly evident (although in both they are less evident than in
|
||
<taxonomicName id="4C06C199FFC0FFAAFC17FC13A58AFC1D" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[1011,1164,987,1011]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC0FFAAFC17FC13A58AFC1D" box="[1011,1164,987,1011]" italics="true" pageId="31" pageNumber="116">B. danzhuense</emphasis>
|
||
</taxonomicName>
|
||
) and the entire dorsum exhibits metallic green reflection. In members of
|
||
<taxonomicName id="4C06C199FFC0FFAAFC92FC33A507FBFA" authorityName="Deuve" authorityYear="2006" box="[886,1025,1020,1044]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="montreuili">
|
||
<emphasis id="B9726608FFC0FFAAFC92FC33A507FBFA" box="[886,1025,1020,1044]" italics="true" pageId="31" pageNumber="116">B. montreuili</emphasis>
|
||
</taxonomicName>
|
||
, the forebody is a darker, less vividly metallic green than the elytra (both parts equally vivid in
|
||
<taxonomicName id="4C06C199FFC0FFAAFC0FFBD3A583FBDA" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[1003,1157,1052,1076]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC0FFAAFC0FFBD3A583FBDA" box="[1003,1157,1052,1076]" italics="true" pageId="31" pageNumber="116">B. danzhuense</emphasis>
|
||
</taxonomicName>
|
||
), and in both
|
||
<taxonomicName id="4C06C199FFC0FFAAFF1EFBF3A080FBBA" authorityName="Deuve" authorityYear="2006" box="[250,390,1084,1108]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="montreuili">
|
||
<emphasis id="B9726608FFC0FFAAFF1EFBF3A080FBBA" box="[250,390,1084,1108]" italics="true" pageId="31" pageNumber="116">B. montreuili</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C06C199FFC0FFAAFE5EFBF3A30FFBBD" authorityName="Deuve" authorityYear="2006" box="[442,521,1084,1107]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="tiani">
|
||
<emphasis id="B9726608FFC0FFAAFE5EFBF3A30FFBBD" box="[442,521,1084,1107]" italics="true" pageId="31" pageNumber="116">B. tiani</emphasis>
|
||
</taxonomicName>
|
||
members the discal portion of the pronotum is more globose (slightly narrower and more elongate in
|
||
<taxonomicName id="4C06C199FFC0FFAAFDCFFB93A3C3FB9A" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[555,709,1116,1140]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC0FFAAFDCFFB93A3C3FB9A" box="[555,709,1116,1140]" italics="true" pageId="31" pageNumber="116">B. danzhuense</emphasis>
|
||
</taxonomicName>
|
||
), the elytra are widest at or near midlength and with striae 2 to 8 more faintly defined and finely punctate, and the femora are darker (lighter piceous or brown in
|
||
<taxonomicName id="4C06C199FFC0FFAAFEBDFB53A0F5FB5A" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[345,499,1180,1204]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC0FFAAFEBDFB53A0F5FB5A" box="[345,499,1180,1204]" italics="true" pageId="31" pageNumber="116">B. danzhuense</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB9BA1AFFC0FFAAFF4FFB74A57DFADD" blockId="31.[123,1174,219,1588]" pageId="31" pageNumber="116">
|
||
<emphasis id="B9726608FFC0FFAAFF4FFB74A090FB3A" bold="true" box="[171,406,1211,1236]" pageId="31" pageNumber="116">Habitat distribution.</emphasis>
|
||
The unique specimen of this species was found under a stone at the edge of a moderate-sized cascading stream descending through mixed coniferous/ broadleaf evergreen forest and shaded during much of the day. The site of collection was at some point within the moderate elevational range of
|
||
<quantity id="4CFE17FFFFC0FFAAFE76FAD3A33BFADD" box="[402,573,1308,1332]" metricMagnitude="3" metricUnit="m" metricValue="2.78" metricValueMax="2.84" metricValueMin="2.7199999999999998" pageId="31" pageNumber="116" unit="m" value="2780.0" valueMax="2840.0" valueMin="2720.0">2720 to 2840 m</quantity>
|
||
(
|
||
<figureCitation id="133DA69FFFC0FFAAFDA9FAD3A39FFADA" box="[589,665,1308,1332]" captionStart="FIGURE" captionStartId="88.[184,196,1682,1701]" captionTargetBox="[130,1162,1063,1660]" captionTargetId="figure-332@88.[120,1176,1054,1668]" captionTargetPageId="88" captionText="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." figureDoi="http://doi.org/10.5281/zenodo.11067457" httpUri="https://zenodo.org/record/11067457/files/figure.png" pageId="31" pageNumber="116">Fig. 49</figureCitation>
|
||
), but the precise elevation was not recorded.
|
||
</paragraph>
|
||
<paragraph id="8BB9BA1AFFC0FFAAFF4FFAF4A2A7FA7D" blockId="31.[123,1174,219,1588]" pageId="31" pageNumber="116">
|
||
<emphasis id="B9726608FFC0FFAAFF4FFAF4A20AFABA" bold="true" box="[171,780,1339,1364]" pageId="31" pageNumber="116">
|
||
Geographical distribution within the Gaoligong
|
||
<materialsCitation id="3B6EB047FFC0FFAAFD2FFAF4A29CFA7A" county="Gongshan County" location="Core Area" pageId="31" pageNumber="116" specimenCount="2" stateProvince="Shan">
|
||
<collectingRegion id="49C274F8FFC0FFAAFD2FFAF4A203FABD" box="[715,773,1339,1363]" country="Myanmar" name="Shan" pageId="31" pageNumber="116">Shan</collectingRegion>
|
||
.
|
||
<figureCitation id="133DA69FFFC0FFAAFCF6FAF3A26CFABA" box="[786,874,1340,1364]" captionStart="FIGURE" captionStartId="29.[171,183,1612,1631]" captionTargetBox="[120,1176,216,1596]" captionTargetId="figure-68@29.[120,1176,216,1598]" captionTargetPageId="29" captionText="FIGURE 17. Broscosoma danzhuense sp. nov. a. Habitus (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067385" httpUri="https://zenodo.org/record/11067385/files/figure.png" pageId="31" pageNumber="116">Fig. 17b</figureCitation>
|
||
. This species is known only from the
|
||
<typeStatus id="54BD04B8FFC0FFAAFF3AFA93A00AFA9A" box="[222,268,1372,1396]" pageId="31" pageNumber="116">type</typeStatus>
|
||
locality in
|
||
<collectingCounty id="62D8C296FFC0FFAAFE6CFA93A34CFA9A" box="[392,586,1372,1396]" pageId="31" pageNumber="116">Gongshan County</collectingCounty>
|
||
(see
|
||
<typeStatus id="54BD04B8FFC0FFAAFD60FA93A3BFFA9A" box="[644,697,1372,1396]" pageId="31" pageNumber="116">Type</typeStatus>
|
||
material above for exact collection data) on the eastern slope of the range in the southern part of
|
||
<location id="8ED9ECC1FFC0FFAAFD57FAB3A226FA7A" LSID="urn:lsid:plazi:treatment:03AF0B0CFFC3FFAAFF9FFAB3A590F9DD:8ED9ECC1FFC0FFAAFD57FAB3A226FA7A" box="[691,800,1404,1428]" county="Gongshan County" name="Core Area" pageId="31" pageNumber="116" stateProvince="Shan">Core Area</location>
|
||
2 (
|
||
<figureCitation id="133DA69FFFC0FFAAFCA1FAB3A294FA7A" box="[837,914,1404,1428]" captionStart="FIGURE" captionStartId="88.[171,183,995,1014]" captionTargetBox="[130,1164,233,969]" captionTargetId="figure-57@88.[120,1176,215,989]" captionTargetPageId="88" captionText="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." figureDoi="http://doi.org/10.5281/zenodo.11067455" httpUri="https://zenodo.org/record/11067455/files/figure.png" pageId="31" pageNumber="116">Fig. 48</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB9BA1AFFC0FFAAFF4FFA53A228FA3D" blockId="31.[123,1174,219,1588]" pageId="31" pageNumber="116">
|
||
<emphasis id="B9726608FFC0FFAAFF4FFA53A334FA5A" bold="true" box="[171,562,1436,1460]" pageId="31" pageNumber="116">Overall geographical distribution.</emphasis>
|
||
This species currently is known only from the northern part of the Gaoligong Shan, in western
|
||
<collectingRegion id="49C274F8FFC0FFAAFDC6FA73A3DFFA3A" box="[546,729,1468,1492]" country="China" name="Yunnan" pageId="31" pageNumber="116">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F311FA8AFFC0FFAAFD03FA73A221FA3A" box="[743,807,1468,1492]" name="China" pageId="31" pageNumber="116">China</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB9BA1AFFC0FFAAFF4FFA13A590F9DD" blockId="31.[123,1174,219,1588]" pageId="31" pageNumber="116">
|
||
<emphasis id="B9726608FFC0FFAAFF4FFA13A24AFA1A" bold="true" box="[171,844,1500,1524]" pageId="31" pageNumber="116">
|
||
Geographical relationships with other
|
||
<taxonomicName id="4C06C199FFC0FFAAFD8CFA13A24AFA1A" box="[616,844,1500,1524]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B9726608FFC0FFAAFD8CFA13A3E9FA1D" bold="true" box="[616,751,1500,1523]" italics="true" pageId="31" pageNumber="116">Broscosoma</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</emphasis>
|
||
No other
|
||
<taxonomicName id="4C06C199FFC0FFAAFC59FA12A593FA1A" box="[957,1173,1500,1524]" pageId="31" pageNumber="116">
|
||
<emphasis id="B9726608FFC0FFAAFC59FA12A546FA1A" box="[957,1088,1501,1524]" italics="true" pageId="31" pageNumber="116">Broscosoma</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
was found syntopic with the
|
||
<typeStatus id="54BD04B8FFC0FFAAFE4EFA33A0DEF9FA" box="[426,472,1532,1556]" pageId="31" pageNumber="116">type</typeStatus>
|
||
of this species, although
|
||
<taxonomicName id="4C06C199FFC0FFAAFD07FA32A239F9FA" authorityName="Putzeys. We" authorityYear="1877" box="[739,831,1532,1556]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="ribbei">
|
||
<emphasis id="B9726608FFC0FFAAFD07FA32A239F9FA" box="[739,831,1532,1556]" italics="true" pageId="31" pageNumber="116">B. ribbei</emphasis>
|
||
</taxonomicName>
|
||
occurs in the same general area.
|
||
<taxonomicName id="4C06C199FFC0FFAAFF9FF9D3A069F9DA" box="[123,367,1564,1588]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="B9726608FFC0FFAAFF9FF9D3A069F9DA" box="[123,367,1564,1588]" italics="true" pageId="31" pageNumber="116">Eobroscus bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
also has been found in the same area and habitat as
|
||
<taxonomicName id="4C06C199FFC0FFAAFC77F9D2A52AF9DA" authorityName="Kavanaugh & Liang" authorityYear="2021" box="[915,1068,1564,1588]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="danzhuense">
|
||
<emphasis id="B9726608FFC0FFAAFC77F9D2A52AF9DA" box="[915,1068,1564,1588]" italics="true" pageId="31" pageNumber="116">B. danzhuense</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133DA69FFFC0FFAAFBDFF9D3A580F9DA" box="[1083,1158,1564,1588]" captionStart="FIGURE" captionStartId="89.[171,183,1461,1480]" captionTargetBox="[140,1154,249,1428]" captionTargetId="figure-68@89.[120,1176,216,1448]" captionTargetPageId="89" captionText="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi="http://doi.org/10.5281/zenodo.11067459" httpUri="https://zenodo.org/record/11067459/files/figure.png" pageId="31" pageNumber="116">Fig. 50</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |