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<document id="F16466C6EA67C0411E4F7BC78290FD8B" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A86CFFF40CBA14EAFAD8960F" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus ramcharani Perkins 2019, new status" docType="treatment" docVersion="8" lastPageNumber="17" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="16" updateTime="1698775642300" updateUser="ExternalLinkService">
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<mods:title id="F4FA5C71001C3ECF317509E45B59CA4C">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="1AE724E82AA52DC2575E149E2BEADAB7">Perkins, Philip D.</mods:namePart>
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<mods:date id="DBE4CB2FE4228761E18B872AEA015E97">2019</mods:date>
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<mods:number id="BCD432185E4AAC88218351BBCB662685">4708</mods:number>
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<treatment id="03A587E7A86CFFF40CBA14EAFAD8960F" ID-DOI="http://doi.org/10.5281/zenodo.5941786" ID-GBIF-Taxon="160798408" ID-Zenodo-Dep="5941786" LSID="urn:lsid:plazi:treatment:03A587E7A86CFFF40CBA14EAFAD8960F" httpUri="http://treatment.plazi.org/id/03A587E7A86CFFF40CBA14EAFAD8960F" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="C316657AA86CFFEB0CBA14EAFD4E949D" box="[151,687,1710,1737]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BB336F1A86CFFEB0CBA14EAFD4E949D" blockId="15.[151,687,1710,1772]" box="[151,687,1710,1737]" pageId="15" pageNumber="16">
<heading id="D0FB819DA86CFFEB0CBA14EAFD4E949D" bold="true" box="[151,687,1710,1737]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="B978EAE3A86CFFEB0CBA14EAFD4E949D" bold="true" box="[151,687,1710,1737]" pageId="15" pageNumber="16">
<taxonomicName id="4C0C4D72A86CFFEB0CBA14EAFDC4949D" ID-CoL="3N6B2" authority="Makhan" authorityName="Perkins" authorityYear="2019" box="[151,549,1710,1737]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ramcharani" status="stat. nov.">
<emphasis id="B978EAE3A86CFFEB0CBA14EAFE52949C" bold="true" box="[151,435,1710,1737]" italics="true" pageId="15" pageNumber="16">Hydrochus ramcharani</emphasis>
Makhan
</taxonomicName>
<taxonomicNameLabel id="A24B5798A86CFFEB0E0114EAFD4E949D" box="[556,687,1710,1736]" pageId="15" pageNumber="16">new status</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA86CFFEB0CBA1496FEED94B9" box="[151,268,1746,1772]" pageId="15" pageNumber="16" type="description">
<paragraph id="8BB336F1A86CFFEB0CBA1496FEED94B9" blockId="15.[151,687,1710,1772]" box="[151,268,1746,1772]" pageId="15" pageNumber="16">
<figureCitation id="13372A74A86CFFEB0CBA1496FF0494B9" box="[151,229,1746,1772]" captionStart="FIGURE 2" captionStartId="14.[151,250,2009,2034]" captionTargetBox="[199,1400,435,1940]" captionTargetId="figure@14.[151,1435,417,1984]" captionTargetPageId="14" captionText="FIGURE 2. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587185" httpUri="https://zenodo.org/record/3587185/files/figure.png" pageId="15" pageNumber="16">Figs. 2</figureCitation>
,
<figureCitation id="13372A74A86CFFEB0CDC1496FEED94B9" box="[241,268,1746,1772]" captionStart="FIGURE 11" captionStartId="27.[151,250,1957,1982]" captionTargetBox="[211,1391,388,1887]" captionTargetId="figure@27.[173,1415,345,1932]" captionTargetPageId="27" captionText="FIGURE 11. Hydrochus ramcharani Makhan, habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587203" httpUri="https://zenodo.org/record/3587203/files/figure.png" pageId="15" pageNumber="16">11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA86CFFEB0CBA1552FCF69518" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8BB336F1A86CFFEB0CBA1552FDAB957A" blockId="15.[151,791,1814,1869]" box="[151,586,1814,1839]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB1C11DFA86CFFEB0CBA1552FDAB957A" box="[151,586,1814,1839]" pageId="15" pageNumber="16">
<taxonomicName id="4C0C4D72A86CFFEB0CBA1552FDA4957A" ID-CoL="3N6B2" authority="Makhan, 1992: 96" authorityName="Makhan" authorityPageNumber="96" authorityYear="1992" box="[151,581,1814,1839]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ramcharani">
<emphasis id="B978EAE3A86CFFEB0CBA1552FE65957B" box="[151,388,1814,1838]" italics="true" pageId="15" pageNumber="16">Hydrochus ramcharani</emphasis>
<treatmentCitation id="0AAD10E0A86CFFEB0DA71552FDA4957A" author="Makhan, D." box="[394,581,1814,1839]" page="96" pageId="15" pageNumber="16" year="1992">
<bibRefCitation id="EF9D4B00A86CFFEB0DA71552FDA4957A" author="Makhan, D." box="[394,581,1814,1839]" pageId="15" pageNumber="16" pagination="95 - 103" refId="ref30207" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan, 1992: 96</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB336F1A86CFFEB0CBA1571FCF69518" blockId="15.[151,791,1814,1869]" box="[151,791,1844,1869]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB1C11DFA86CFFEB0CBA1571FCF69518" box="[151,791,1844,1869]" pageId="15" pageNumber="16">
<taxonomicName id="4C0C4D72A86CFFEB0CBA1571FE7E9518" ID-CoL="3N67B" authority="Knisch" authorityName="Knisch" box="[151,415,1844,1869]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="drakei">
<emphasis id="B978EAE3A86CFFEB0CBA1571FEAC9518" box="[151,333,1845,1869]" italics="true" pageId="15" pageNumber="16">Hydrochus drakei</emphasis>
Knisch
</taxonomicName>
:
<treatmentCitation id="0AAD10E0A86CFFEB0D851570FD809518" author="Oliva, A." box="[424,609,1844,1869]" page="329" pageId="15" pageNumber="16" year="1996">
<bibRefCitation id="EF9D4B00A86CFFEB0D851570FD809518" author="Oliva, A." box="[424,609,1844,1869]" pageId="15" pageNumber="16" pagination="301 - 341" refId="ref30673" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996: 329)</bibRefCitation>
</treatmentCitation>
; misidentification
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA86CFFF40CBA153EFD4D92AF" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BB336F1A86CFFEB0CBA153EFAF395EF" blockId="15.[151,1437,1914,2014]" pageId="15" pageNumber="16">
<emphasis id="B978EAE3A86CFFEB0CBA153EFE4495C0" bold="true" box="[151,421,1914,1941]" pageId="15" pageNumber="16">Material examined (8):</emphasis>
<materialsCitation id="3B643CACA86CFFEB0D841538FC1095C3" ID-GBIF-Occurrence="2521263703" box="[425,1009,1914,1942]" collectionCode="MIZA" country="Venezuela" location="Anzoategui" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Anzoategui">
<emphasis id="B978EAE3A86CFFEB0D841538FD5F95C0" bold="true" box="[425,702,1914,1942]" pageId="15" pageNumber="16">
<collectingCountry id="F31B7661A86CFFEB0D841538FDC795C3" box="[425,550,1916,1942]" name="Venezuela" pageId="15" pageNumber="16">Venezuela</collectingCountry>
:
<collectingRegion id="49C8F813A86CFFEB0E00153EFD5895C0" box="[557,697,1914,1941]" country="Venezuela" name="Anzoategui" pageId="15" pageNumber="16">Anzoategui</collectingRegion>
:
</emphasis>
VZ10-0204-01A (1
<collectionCode id="ED1DAE34A86CFFEB0FB01538FC0B95C3" box="[925,1002,1916,1942]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="15" pageNumber="16" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA86CFFEB0FD4153EFDC595EC" ID-GBIF-Occurrence="2521263852" collectingDate="1996-06-11" collectionCode="CMN" collectorName="E. Caicara &amp; H. &amp; A. Howden" country="Venezuela" location="Bolivar" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Bolivar">
<emphasis id="B978EAE3A86CFFEB0FD4153EFBBA95C0" bold="true" box="[1017,1115,1914,1941]" pageId="15" pageNumber="16">
<collectingRegion id="49C8F813A86CFFEB0FD4153EFBB795C0" box="[1017,1110,1914,1941]" country="Venezuela" name="Bolivar" pageId="15" pageNumber="16">Bolivar</collectingRegion>
:
</emphasis>
<quantity id="4CF49B14A86CFFEB0872153EFB7495C0" box="[1119,1173,1914,1941]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="km" value="5.0">5 km</quantity>
<collectorName id="26F95327A86CFFEB08B71538FAEF95C0" box="[1178,1294,1915,1942]" pageId="15" pageNumber="16">E. Caicara</collectorName>
,
<date id="FFB21031A86CFFEB093A153FFA7995C0" box="[1303,1432,1915,1941]" pageId="15" pageNumber="16" value="1996-06-11">
<collectingDate id="EFF6E9D9A86CFFEB093A153FFA7995C0" box="[1303,1432,1915,1941]" pageId="15" pageNumber="16" value="1996-06-11">11.VI.1996</collectingDate>
</date>
, at light,
<collectorName id="26F95327A86CFFEB0CD915E4FE5395EC" box="[244,434,1951,1978]" pageId="15" pageNumber="16">H. &amp; A. Howden</collectorName>
(4
<collectionCode id="ED1DAE34A86CFFEB0DF515DBFDFC95EC" box="[472,541,1951,1977]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="15" pageNumber="16" type="Museum">CMN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA86CFFEB0E0215DAFAEF95EF" ID-GBIF-Occurrence="2521263717" box="[559,1294,1950,1978]" collectingDate="1969-02-12" collectionCode="NMNH" collectorName="P. &amp; P. Spangler" country="Venezuela" location="San Fernando" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Guarico">
<emphasis id="B978EAE3A86CFFEB0E0215DAFD7B95EC" bold="true" box="[559,666,1950,1977]" pageId="15" pageNumber="16">
<collectingRegion id="49C8F813A86CFFEB0E0215DAFD7495EC" box="[559,661,1950,1977]" country="Venezuela" name="Guarico" pageId="15" pageNumber="16">Guarico</collectingRegion>
:
</emphasis>
<location id="8ED3602AA86CFFEB0E8C15DBFCDE95EC" LSID="urn:lsid:plazi:treatment:03A587E7A86CFFF40CBA14EAFAD8960F:8ED3602AA86CFFEB0E8C15DBFCDE95EC" box="[673,831,1951,1977]" country="Venezuela" name="San Fernando" pageId="15" pageNumber="16" stateProvince="Guarico">San Fernando</location>
,
<date id="FFB21031A86CFFEB0F6715DBFC2795EC" box="[842,966,1951,1977]" pageId="15" pageNumber="16" value="1969-02-12">
<collectingDate id="EFF6E9D9A86CFFEB0F6715DBFC2795EC" box="[842,966,1951,1977]" pageId="15" pageNumber="16" value="1969-02-12">II-12-1969</collectingDate>
</date>
,
<collectorName id="26F95327A86CFFEB0FFC15E4FB6695EC" box="[977,1159,1951,1978]" pageId="15" pageNumber="16">P. &amp; P. Spangler</collectorName>
(3
<collectionCode id="ED1DAE34A86CFFEB088115E4FAE695EF" box="[1196,1287,1952,1978]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB336F1A86CFFF40CEA1586FD4D92AF" blockId="15.[151,1437,1914,2014]" lastBlockId="16.[151,1437,152,1115]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="B978EAE3A86CFFEB0CEA1586FE309588" bold="true" box="[199,465,1986,2013]" pageId="15" pageNumber="16">Differential Diagnosis.</emphasis>
Among Venezuelan species, recognized by the combination of moderate size (ca. 3.85 4.12 mm), the fine (pronotal) to only moderately coarse (elytral) dorsal punctation, with well separated punctures, and the distinctive male genitalia (
<figureCitation id="13372A74A873FFF40E0512F8FD9C9282" box="[552,637,188,215]" captionStart="FIGURE 11" captionStartId="27.[151,250,1957,1982]" captionTargetBox="[211,1391,388,1887]" captionTargetId="figure@27.[173,1415,345,1932]" captionTargetPageId="27" captionText="FIGURE 11. Hydrochus ramcharani Makhan, habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587203" httpUri="https://zenodo.org/record/3587203/files/figure.png" pageId="16" pageNumber="17">Fig. 11</figureCitation>
; described below).
<taxonomicName id="4C0C4D72A873FFF40F4C12F9FBEB9283" authorityName="Makhan" authorityYear="1992" box="[865,1034,188,214]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ramcharani">
<emphasis id="B978EAE3A873FFF40F4C12F9FBEB9283" box="[865,1034,188,214]" italics="true" pageId="16" pageNumber="17">H. ramcharani</emphasis>
</taxonomicName>
has a generalized habitus; reliable determinations will require dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C316657AA873FFF40CEA1340FB2691BA" pageId="16" pageNumber="17" type="description">
<paragraph id="8BB336F1A873FFF40CEA1340FF379387" blockId="16.[151,1437,152,1115]" pageId="16" pageNumber="17">
<emphasis id="B978EAE3A873FFF40CEA1340FEB0934A" bold="true" box="[199,337,260,287]" pageId="16" pageNumber="17">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="13372A74A873FFF40E4B1340FD5A934A" box="[614,699,260,287]" captionStart="FIGURE 11" captionStartId="27.[151,250,1957,1982]" captionTargetBox="[211,1391,388,1887]" captionTargetId="figure@27.[173,1415,345,1932]" captionTargetPageId="27" captionText="FIGURE 11. Hydrochus ramcharani Makhan, habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587203" httpUri="https://zenodo.org/record/3587203/files/figure.png" pageId="16" pageNumber="17">Fig. 11</figureCitation>
). Size of imaged specimen (length/width, mm): body (length to elytral apices) 4.12/1.44; head width 0.96; pronotum l/w 0.88/0.92; PA 0.84; PB 0.76; elytra 2.52/1.44. Size ca.
<quantity id="4CF49B14A873FFF40CBA1308FEDE9332" box="[151,319,332,359]" metricMagnitude="-3" metricUnit="m" metricValue="3.985" metricValueMax="4.12" metricValueMin="3.85" pageId="16" pageNumber="17" unit="mm" value="3.985" valueMax="4.12" valueMin="3.85">3.854.12 mm</quantity>
. Dorsum grey with very slight iridescence, elytra with black spots. Legs testaceous to dark brown to black. Dorsal punctation small to moderately large, larger on elytra than pronotum. Elytra interstriae ca. 1.5
<date id="FFB21031A873FFF409AD1334FA7D93DE" box="[1408,1436,368,395]" pageId="16" pageNumber="17">2x</date>
strial punctures diameter. Interstria 9
<superScript id="7C799BB9A873FFF40E0313D7FDDA93F4" attach="left" box="[558,571,403,417]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
more convex than others, overhanging 10
<superScript id="7C799BB9A873FFF4082113D7FBF893F4" attach="left" box="[1036,1049,403,417]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
interstria. Callus on 5
<superScript id="7C799BB9A873FFF4092113D7FAF893F4" attach="left" box="[1292,1305,403,417]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
moderately large.
</paragraph>
<paragraph id="8BB336F1A873FFF40CEA1399FD1C904F" blockId="16.[151,1437,152,1115]" pageId="16" pageNumber="17">Pronotum slightly wider than long, widest just behind anterior margin, narrowed at base, sides arcuate or very weakly sinuate, smooth; depressions moderately sized.</paragraph>
<paragraph id="8BB336F1A873FFF40CEA1060FBB69037" blockId="16.[151,1437,152,1115]" pageId="16" pageNumber="17">
Elytral strial punctures ca. 2
<date id="FFB21031A873FFF40E201061FDC6906A" box="[525,551,549,575]" pageId="16" pageNumber="17">3x</date>
size of pronotal punctures, spaces between them about
<date id="FFB21031A873FFF408A11061FB49906A" box="[1164,1192,549,575]" pageId="16" pageNumber="17">2x</date>
their diameter; apices conjointly rounded in dorsal view, in lateral view outer margin with slight angulation.
</paragraph>
<paragraph id="8BB336F1A873FFF40CEA1028FF0890FF" blockId="16.[151,1437,152,1115]" pageId="16" pageNumber="17">Ventral characters: Mentum densely, moderately coarsely punctate, with median fovea. Submentum without foveae.</paragraph>
<paragraph id="8BB336F1A873FFF40CEA10F0FF39910B" blockId="16.[151,1437,152,1115]" pageId="16" pageNumber="17">
Male genitalia (
<figureCitation id="13372A74A873FFF40D5410F0FE28909A" box="[377,457,692,719]" captionStart="FIGURE 11" captionStartId="27.[151,250,1957,1982]" captionTargetBox="[211,1391,388,1887]" captionTargetId="figure@27.[173,1415,345,1932]" captionTargetPageId="27" captionText="FIGURE 11. Hydrochus ramcharani Makhan, habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587203" httpUri="https://zenodo.org/record/3587203/files/figure.png" pageId="16" pageNumber="17">Fig. 11</figureCitation>
) general characters: genitalia very stout, wide in lateral view; paramere tips extending slightly beyond distal end of aedeagus, lateral margins sinuate, medial margins parallel, in lateral view extreme end of tips upturned slightly; aedeagus lanceolate; pseudoparameres large, extending to level of distal end of aedeagus, extending beyond plsa; basal piece with orifice strongly sclerotized, and with strongly sclerotized midline ventrally.
</paragraph>
<paragraph id="8BB336F1A873FFF40CEA112CFE2591F3" blockId="16.[151,1437,152,1115]" pageId="16" pageNumber="17">Dorsal surface: adtl located near midlength of aedeagus; pdmm overlapping lateral margins of aedeagus; adbl large, distinctively shaped.</paragraph>
<paragraph id="8BB336F1A873FFF40CEA11F4FB2691BA" blockId="16.[151,1437,152,1115]" pageId="16" pageNumber="17">Ventral surface: avtl U-shaped, near basal 1/3 of aedeagus; pvmm overlapping lateral margins of aedeagus, but more widely separated than pdmm; pseudoparameres enveloping most of basal ¼ of parameres.</paragraph>
</subSubSection>
<subSubSection id="C316657AA873FFF40CEA11BDFAD8960F" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BB336F1A873FFF40CEA11BDFAD8960F" blockId="16.[151,1437,152,1115]" pageId="16" pageNumber="17">
<emphasis id="B978EAE3A873FFF40CEA11BDFEDA9646" bold="true" box="[199,315,1017,1043]" pageId="16" pageNumber="17">Remarks.</emphasis>
The specimen imaged (
<figureCitation id="13372A74A873FFF40E6711BCFD759646" box="[586,660,1016,1043]" captionStart="FIGURE 11" captionStartId="27.[151,250,1957,1982]" captionTargetBox="[211,1391,388,1887]" captionTargetId="figure@27.[173,1415,345,1932]" captionTargetPageId="27" captionText="FIGURE 11. Hydrochus ramcharani Makhan, habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587203" httpUri="https://zenodo.org/record/3587203/files/figure.png" pageId="16" pageNumber="17">Fig 11</figureCitation>
) is from locality VZ10-0204-01A. I have examined a male
<typeStatus id="54B78853A873FFF4091111BDFA7D9646" box="[1340,1436,1017,1043]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C0C4D72A873FFF40CEA1659FE8F9663" authorityName="Makhan" authorityYear="1992" box="[199,366,1052,1078]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ramcharani">
<emphasis id="B978EAE3A873FFF40CEA1659FE8F9663" box="[199,366,1052,1078]" italics="true" pageId="16" pageNumber="17">H. ramcharani</emphasis>
</taxonomicName>
, including the genitalia, and must disagree with
<bibRefCitation id="EF9D4B00A873FFF40FBA1658FBCA9662" author="Oliva, A." box="[919,1067,1052,1079]" pageId="16" pageNumber="17" pagination="301 - 341" refId="ref30673" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996)</bibRefCitation>
, who placed
<taxonomicName id="4C0C4D72A873FFF408EC1659FA899663" authorityName="Makhan" authorityYear="1992" box="[1217,1384,1052,1078]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ramcharani">
<emphasis id="B978EAE3A873FFF408EC1659FA899663" box="[1217,1384,1052,1078]" italics="true" pageId="16" pageNumber="17">H. ramcharani</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C0C4D72A873FFF40D401605FD82960E" authority="Knisch. See" authorityName="Knisch. See" box="[365,611,1088,1115]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="drakei">
<emphasis id="B978EAE3A873FFF40D401605FE36960F" box="[365,471,1088,1114]" italics="true" pageId="16" pageNumber="17">H. drakei</emphasis>
Knisch. See
</taxonomicName>
also
<bibRefCitation id="EF9D4B00A873FFF40EB21604FCA4960E" author="Perkins, P. D." box="[671,837,1088,1115]" pageId="16" pageNumber="17" pagination="149 - 167" refId="ref30761" refString="Perkins, P. D. (2019) Type designations for Neotropical water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae). Zo otaxa, 47 01 (2), 149 - 167. https: // doi. org / 10.11646 / zootaxa. 4701.6.7" type="journal article" year="2019">Perkins (2019)</bibRefCitation>
on the identity of
<taxonomicName id="4C0C4D72A873FFF4083F1604FAD8960F" authority="Knisch." authorityName="Knisch." box="[1042,1337,1088,1114]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="drakei">
<emphasis id="B978EAE3A873FFF4083F1604FB3A960F" box="[1042,1243,1088,1114]" italics="true" pageId="16" pageNumber="17">Hydrochus drakei</emphasis>
Knisch.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>