treatments-xml/data/03/A5/87/03A587E7A84FFFC80CBA1101FB13958A.xml
2024-06-21 12:22:17 +02:00

229 lines
25 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="2DB127A778D3DEA7BC33BA91DA044CE1" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A84FFFC80CBA1101FB13958A" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus argutissimus Perkins 2019, new species" docType="treatment" docVersion="8" lastPageNumber="45" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="45" updateTime="1698775642300" updateUser="ExternalLinkService">
<mods:mods id="B4439D09B241F9E1F89680B471C9D308" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AB50026E3D76B4590F77DC61ACD40204">
<mods:title id="57F485AE006C490BB3033748ABFE501A">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
<mods:name id="B89795C927762079997FCDCAA67DBE84" type="personal">
<mods:role id="ADB4536484BE33D546E3024E43A34172">
<mods:roleTerm id="8ABE6B11F8AA3FFCF3D046DB08E8F5A6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="68D6774741A437646F3C85CC773A22F8">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="368C300D34E4646F4DB15EF109E1F130">text</mods:typeOfResource>
<mods:relatedItem id="08F09EC02D2FB7AE9600427088772315" type="host">
<mods:titleInfo id="A39844666FA7D00B60AA6F2E04D82195">
<mods:title id="866BA183622964E88C7B1EA66CEF1F5D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="115DD666B1D0DCABC012B468CFCD2357">
<mods:date id="775D234CF39504476D489BAC547F56D7">2019</mods:date>
<mods:detail id="4B25E7F5DC310EA970B899AA68720A7A" type="pubDate">
<mods:number id="69BC6260E7A63D3334E2BFB1DA7082EC">2019-12-12</mods:number>
</mods:detail>
<mods:detail id="E7C3D4E32C25D49AF59BD42B6E82C52B" type="volume">
<mods:number id="6FEAEBD51F2D4BD5DDCDFDF5F63E47F7">4708</mods:number>
</mods:detail>
<mods:detail id="3227336D4717F4F43EBAB9D4204AA67F" type="issue">
<mods:number id="1C892E9E2E76BA0C0D46A5E701222E7F">1</mods:number>
</mods:detail>
<mods:extent id="97BF280971F5EA9226F47B45CE696A5F" unit="page">
<mods:start id="6AAF26566C9C7FC99D97A252654A960D">1</mods:start>
<mods:end id="B9B4857AD9A5B1246DFC2A5BFBC24392">59</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4F6596D5BE502BC792EF9188CBC452AD">journal article</mods:classification>
<mods:identifier id="7DF35220115FC2785B69B6429EA1C4EC" type="CLB-Dataset">24540</mods:identifier>
<mods:identifier id="F1829188E116F5B247DB7F8EDE1EE60E" type="DOI">10.11646/zootaxa.4708.1.1</mods:identifier>
<mods:identifier id="BD118B932C0745B377CD46043818B6A6" type="GBIF-Dataset">653d580d-e1c7-499c-8604-4b5a7b1920ef</mods:identifier>
<mods:identifier id="C4D13D883BF1DAA8DE4CCB55F0BA854A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="69FD41831576B16108EA9A9FB68928D0" type="Zenodo-Dep">3587181</mods:identifier>
<mods:identifier id="05951B7B7A8925224235FFEC2AD95C4E" type="ZooBank">09F24F0B-5090-4089-A525-8E5150C91EF7</mods:identifier>
</mods:mods>
<treatment id="03A587E7A84FFFC80CBA1101FB13958A" ID-DOI="http://doi.org/10.5281/zenodo.5941807" ID-GBIF-Taxon="160798416" ID-Zenodo-Dep="5941807" LSID="urn:lsid:plazi:treatment:03A587E7A84FFFC80CBA1101FB13958A" httpUri="http://treatment.plazi.org/id/03A587E7A84FFFC80CBA1101FB13958A" lastPageNumber="45" pageId="44" pageNumber="45">
<subSubSection id="C316657AA84FFFC80CBA1101FDB5910A" box="[151,596,837,864]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8BB336F1A84FFFC80CBA1101FDB5910A" blockId="44.[151,596,837,899]" box="[151,596,837,864]" pageId="44" pageNumber="45">
<heading id="D0FB819DA84FFFC80CBA1101FDB5910A" bold="true" box="[151,596,837,864]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<emphasis id="B978EAE3A84FFFC80CBA1101FDB5910A" bold="true" box="[151,596,837,864]" italics="true" pageId="44" pageNumber="45">
<taxonomicName id="4C0C4D72A84FFFC80CBA1101FE5E910A" ID-CoL="8Y4P3" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[151,447,837,864]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="argutissimus" status="sp. nov.">Hydrochus argutissimus</taxonomicName>
<taxonomicNameLabel id="A24B5798A84FFFC80DEB1101FDB5910A" box="[454,596,837,863]" pageId="44" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA84FFFC80CBA112DFEEC91D6" box="[151,269,872,899]" pageId="44" pageNumber="45" type="description">
<paragraph id="8BB336F1A84FFFC80CBA112DFEEC91D6" blockId="44.[151,596,837,899]" box="[151,269,872,899]" pageId="44" pageNumber="45">
<figureCitation id="13372A74A84FFFC80CBA112DFF0491D6" box="[151,229,873,899]" captionStart="FIGURE 2" captionStartId="14.[151,250,2009,2034]" captionTargetBox="[199,1400,435,1940]" captionTargetId="figure@14.[151,1435,417,1984]" captionTargetPageId="14" captionText="FIGURE 2. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587185" httpUri="https://zenodo.org/record/3587185/files/figure.png" pageId="44" pageNumber="45">Figs. 2</figureCitation>
,
<figureCitation id="13372A74A84FFFC80CDC112CFEEC91D6" box="[241,269,872,899]" captionStart="FIGURE 25" captionStartId="45.[151,250,1794,1819]" captionTargetBox="[227,1381,186,1751]" captionTargetId="figure@45.[178,1410,181,1768]" captionTargetPageId="45" captionText="FIGURE 25. Hydrochus argutissimus n. sp., habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587233" httpUri="https://zenodo.org/record/3587233/files/figure.png" pageId="44" pageNumber="45">25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA84FFFC80CBA11F5FD08975A" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8BB336F1A84FFFC80CBA11F5FCDA960E" blockId="44.[151,1437,944,2015]" pageId="44" pageNumber="45">
<collectingRegion id="49C8F813A84FFFC80CBA11F5FEA8919E" box="[151,329,944,971]" pageId="44" pageNumber="45">
<emphasis id="B978EAE3A84FFFC80CBA11F5FEA8919E" bold="true" box="[151,329,944,971]" pageId="44" pageNumber="45">Type Material.</emphasis>
</collectingRegion>
<materialsCitation id="3B643CACA84FFFC80D7D11F5FEDD9646" ID-GBIF-Occurrence="2521263668" collectingDate="2009-07-16" collectionCode="MIZA" collectorName="Short &amp; Sites &amp; Garcia &amp; Inciarte &amp; Gustafson &amp; Camacho" country="Venezuela" latitude="7.5839667" location="El Tama National Park" longLatPrecision="1" longitude="-72.17233" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Tachira" typeStatus="holotype">
<emphasis id="B978EAE3A84FFFC80D7D11F5FDFB919E" bold="true" box="[336,538,944,971]" pageId="44" pageNumber="45">
<typeStatus id="54B78853A84FFFC80D7D11F5FE5C919E" box="[336,445,945,971]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="F31B7661A84FFFC80E0111F5FD3B919E" box="[556,730,945,971]" name="Venezuela" pageId="44" pageNumber="45">VENEZUELA</collectingCountry>
:
<collectingRegion id="49C8F813A84FFFC80EC911F5FC9B919E" box="[740,890,945,971]" country="Venezuela" name="Tachira" pageId="44" pageNumber="45">Tachira State</collectingRegion>
,
<geoCoordinate id="EE385036A84FFFC80FAB11F5FBFB919E" box="[902,1050,944,971]" degrees="7" direction="north" minutes="35.038" orientation="latitude" pageId="44" pageNumber="45" precision="1" value="7.5839667">7° 35.038 N</geoCoordinate>
,
<geoCoordinate id="EE385036A84FFFC8080811F5FB2A919E" box="[1061,1227,945,971]" degrees="72" direction="west" minutes="10.340" orientation="longitude" pageId="44" pageNumber="45" precision="1" value="-72.17233">72° 10.340 W</geoCoordinate>
,
<location id="8ED3602AA84FFFC808FB11F5FF2C91BA" LSID="urn:lsid:plazi:treatment:03A587E7A84FFFC80CBA1101FB13958A:8ED3602AA84FFFC808FB11F5FF2C91BA" country="Venezuela" latitude="7.5839667" longLatPrecision="1" longitude="-72.17233" name="El Tama National Park" pageId="44" pageNumber="45" stateProvince="Tachira">El Tama National Park</location>
,
<date id="FFB21031A84FFFC80CF41191FEBD91BA" box="[217,348,981,1007]" pageId="44" pageNumber="45" value="2009-07-16">
<collectingDate id="EFF6E9D9A84FFFC80CF41191FEBD91BA" box="[217,348,981,1007]" pageId="44" pageNumber="45" value="2009-07-16">
16.vii.
<quantity id="4CF49B14A84FFFC80D301191FEBD91BA" box="[285,348,981,1007]" metricMagnitude="6" metricUnit="m" metricValue="2.009472" pageId="44" pageNumber="45" unit="m" value="2009472.0">2009</quantity>
</collectingDate>
</date>
, 472m, leg.
<collectorName id="26F95327A84FFFC80DCA1191FDC791BA" box="[487,550,981,1007]" pageId="44" pageNumber="45">Short</collectorName>
,
<collectorName id="26F95327A84FFFC80E1E1191FD8D91BA" box="[563,620,981,1007]" pageId="44" pageNumber="45">Sites</collectorName>
,
<collectorName id="26F95327A84FFFC80E551191FD2491BA" box="[632,709,981,1007]" pageId="44" pageNumber="45">Garcia</collectorName>
,
<collectorName id="26F95327A84FFFC80EFF1191FCCB91BA" box="[722,810,981,1007]" pageId="44" pageNumber="45">Inciarte</collectorName>
,
<collectorName id="26F95327A84FFFC80F1B1191FC4991BA" box="[822,936,981,1007]" pageId="44" pageNumber="45">Gustafson</collectorName>
&amp;
<collectorName id="26F95327A84FFFC80FE01191FBDB91BA" box="[973,1082,981,1007]" pageId="44" pageNumber="45">Camacho</collectorName>
,, HG-vapor light; VZ09-0716- 07A” (
<collectionCode id="ED1DAE34A84FFFC80CCB11BDFED59646" box="[230,308,1017,1043]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="44" pageNumber="45" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA84FFFC80D6411BDFF1B9662" ID-GBIF-Occurrence="2521263819" collectingDate="1982-11-01" collectionCode="NMNH" collectorName="El Dorado &amp; John L. Hellman" country="Venezuela" location="Bolivar" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Bolivar" typeStatus="paratype">
<emphasis id="B978EAE3A84FFFC80D6411BDFD1B9646" bold="true" box="[329,762,1016,1043]" pageId="44" pageNumber="45">
<typeStatus id="54B78853A84FFFC80D6411BDFE209646" box="[329,449,1017,1043]" pageId="44" pageNumber="45" type="paratype">Paratypes</typeStatus>
(41): Venezuela:
<collectingRegion id="49C8F813A84FFFC80EB511BCFD149646" box="[664,757,1016,1043]" country="Venezuela" name="Bolivar" pageId="44" pageNumber="45">Bolivar</collectingRegion>
:
</emphasis>
<collectorName id="26F95327A84FFFC80F2911BDFC9B9646" box="[772,890,1017,1043]" pageId="44" pageNumber="45">El Dorado</collectorName>
(
<quantity id="4CF49B14A84FFFC80FA011BCFC399646" box="[909,984,1016,1043]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="44" pageNumber="45" unit="km" value="65.0">65 km</quantity>
S),
<date id="FFB21031A84FFFC8082611BDFB7F9646" box="[1035,1182,1017,1043]" pageId="44" pageNumber="45" value="1982-11-01">
<collectingDate id="EFF6E9D9A84FFFC8082611BDFB7F9646" box="[1035,1182,1017,1043]" pageId="44" pageNumber="45" value="1982-11-01">1 Nov. 1982</collectingDate>
</date>
,
<collectorName id="26F95327A84FFFC8088111BDFA8C9646" box="[1196,1389,1017,1043]" pageId="44" pageNumber="45">John L. Hellman</collectorName>
(33
<collectionCode id="ED1DAE34A84FFFC80CBA1659FF139662" box="[151,242,1053,1079]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="44" pageNumber="45" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA84FFFC80D291659FC5A9662" ID-GBIF-Occurrence="2521263702" box="[260,955,1053,1079]" collectionCode="NMNH" country="Venezuela" location="Guanare" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Portuguesa" typeStatus="paratype">
<emphasis id="B978EAE3A84FFFC80D291659FE759662" bold="true" box="[260,404,1053,1079]" pageId="44" pageNumber="45">
<collectingRegion id="49C8F813A84FFFC80D291659FE719662" box="[260,400,1053,1079]" country="Venezuela" name="Portuguesa" pageId="44" pageNumber="45">Portuguesa</collectingRegion>
:
</emphasis>
<location id="8ED3602AA84FFFC80DB61659FE1C9662" LSID="urn:lsid:plazi:treatment:03A587E7A84FFFC80CBA1101FB13958A:8ED3602AA84FFFC80DB61659FE1C9662" box="[411,509,1053,1079]" country="Venezuela" name="Guanare" pageId="44" pageNumber="45" stateProvince="Portuguesa">Guanare</location>
, at light, Borys Malkin, leg. (1
<collectionCode id="ED1DAE34A84FFFC80F751659FC529662" box="[856,947,1053,1079]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="44" pageNumber="45" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA84FFFC80FE81659FE69960E" ID-GBIF-Occurrence="2521263707" collectingDate="1957-09-10" collectingDateMax="1957-09-13" collectingDateMin="1957-09-10" collectionCode="NMNH" collectorName="Borys Malkin Collector" country="Venezuela" location="Guanare" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Venezuela" typeStatus="paratype">
<location id="8ED3602AA84FFFC80FE81659FBC69662" LSID="urn:lsid:plazi:treatment:03A587E7A84FFFC80CBA1101FB13958A:8ED3602AA84FFFC80FE81659FBC69662" box="[965,1063,1053,1079]" country="Venezuela" name="Guanare" pageId="44" pageNumber="45" stateProvince="Venezuela">Guanare</location>
,
<date id="FFB21031A84FFFC8081C1659FB169662" box="[1073,1271,1052,1079]" pageId="44" pageNumber="45" value="1957-09-10" valueMax="1957-09-13" valueMin="1957-09-10">
<collectingDate id="EFF6E9D9A84FFFC8081C1659FB169662" box="[1073,1271,1052,1079]" pageId="44" pageNumber="45" value="1957-09-10" valueMax="1957-09-13" valueMin="1957-09-10">IX-10 to 13-1957</collectingDate>
</date>
,
<collectorName id="26F95327A84FFFC8092F1659FF1E960E" pageId="44" pageNumber="45">Borys Malkin Collector</collectorName>
(1
<collectionCode id="ED1DAE34A84FFFC80D081605FE61960E" box="[293,384,1089,1115]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="44" pageNumber="45" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA84FFFC80DBF1604FCD6960E" ID-GBIF-Occurrence="2521263878" box="[402,823,1088,1115]" collectionCode="SEMC" country="Venezuela" location="Tachira" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Tachira" typeStatus="paratype">
<emphasis id="B978EAE3A84FFFC80DBF1604FE16960E" bold="true" box="[402,503,1088,1115]" pageId="44" pageNumber="45">
<collectingRegion id="49C8F813A84FFFC80DBF1604FE12960E" box="[402,499,1088,1115]" country="Venezuela" name="Tachira" pageId="44" pageNumber="45">Tachira</collectingRegion>
:
</emphasis>
VZ09-0716-07A (6
<collectionCode id="ED1DAE34A84FFFC80EF31605FCCE960E" box="[734,815,1089,1115]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="44" pageNumber="45" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB336F1A84FFFC80CEA1620FD08975A" blockId="44.[151,1437,944,2015]" pageId="44" pageNumber="45">
<emphasis id="B978EAE3A84FFFC80CEA1620FE34962A" bold="true" box="[199,469,1124,1151]" pageId="44" pageNumber="45">Differential Diagnosis.</emphasis>
Among Venezuelan species, recognized by the combination of small size (ca.
<quantity id="4CF49B14A84FFFC809701621FEE096F6" metricMagnitude="-3" metricUnit="m" metricValue="2.57" metricValueMax="2.85" metricValueMin="2.29" pageId="44" pageNumber="45" unit="mm" value="2.57" valueMax="2.85" valueMin="2.29">2.29 2.85 mm</quantity>
), the pronotum with moderately deep, densely punctate depressions, the pronotal shape, the elytral disc with very large punctures and very narrow, zig-zag shaped interstriae, the large punctures of the elytral apices, and the male genitalia (
<figureCitation id="13372A74A84FFFC80D461695FE5F96BE" box="[363,446,1232,1259]" captionStart="FIGURE 25" captionStartId="45.[151,250,1794,1819]" captionTargetBox="[227,1381,186,1751]" captionTargetId="figure@45.[178,1410,181,1768]" captionTargetPageId="45" captionText="FIGURE 25. Hydrochus argutissimus n. sp., habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587233" httpUri="https://zenodo.org/record/3587233/files/figure.png" pageId="44" pageNumber="45">Fig. 25</figureCitation>
; described below). Differentiation from other similarly small sized species with similarly shaped pronotum should include dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C316657AA84FFFC80CEA175CFC8F95C2" pageId="44" pageNumber="45" type="description">
<paragraph id="8BB336F1A84FFFC80CEA175CFAFB97B2" blockId="44.[151,1437,944,2015]" pageId="44" pageNumber="45">
<emphasis id="B978EAE3A84FFFC80CEA175CFEB09766" bold="true" box="[199,337,1304,1331]" pageId="44" pageNumber="45">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="13372A74A84FFFC80E49175DFD5B9766" box="[612,698,1304,1331]" captionStart="FIGURE 25" captionStartId="45.[151,250,1794,1819]" captionTargetBox="[227,1381,186,1751]" captionTargetId="figure@45.[178,1410,181,1768]" captionTargetPageId="45" captionText="FIGURE 25. Hydrochus argutissimus n. sp., habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587233" httpUri="https://zenodo.org/record/3587233/files/figure.png" pageId="44" pageNumber="45">Fig. 25</figureCitation>
). Size:
<typeStatus id="54B78853A84FFFC80F3D175DFC939766" box="[784,882,1305,1331]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.29 /0.81; head width 0.54; pronotum l/w 0.52/0.54; PA 0.54; PB 0.42; elytra 1.48 /0.81. Body size small (ca.
<quantity id="4CF49B14A84FFFC80CBA1724FEDC972E" box="[151,317,1376,1403]" metricMagnitude="-3" metricUnit="m" metricValue="2.57" metricValueMax="2.85" metricValueMin="2.29" pageId="44" pageNumber="45" unit="mm" value="2.57" valueMax="2.85" valueMin="2.29">2.292.85 mm</quantity>
). Dorsum dark brown to reddish brown, with very slight iridescence, elytra without black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra deep, ca. 2
<date id="FFB21031A84FFFC80FDC17C1FBEC97CA" box="[1009,1037,1413,1439]" pageId="44" pageNumber="45">4x</date>
that of pronotum. Elytra interstriae narrow ridges to ca.
<date id="FFB21031A84FFFC80D5A17EDFE729796" box="[375,403,1449,1475]" pageId="44" pageNumber="45">1x</date>
strial puncture diameter, often with zig-zag pattern due to large, alternately spaced punctures. Interstria 9
<superScript id="7C799BB9A84FFFC80D3C1788FEFF978F" attach="left" box="[273,286,1484,1498]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
more convex than others, overhanging 10
<superScript id="7C799BB9A84FFFC80EDB1788FCE2978F" attach="left" box="[758,771,1484,1498]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C799BB9A84FFFC808BC1788FB7F978F" attach="left" box="[1169,1182,1484,1498]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
not raised.
</paragraph>
<paragraph id="8BB336F1A84FFFC80CEA17B5FECE9406" blockId="44.[151,1437,944,2015]" pageId="44" pageNumber="45">Pronotum slightly wider than long (as ca. 54/52), widest at anterior margin, markedly narrowed at base, sides straight, not denticulate; anterior margin bisinuate; depressions shallow, densely punctate; reliefs very low, generally punctate.</paragraph>
<paragraph id="8BB336F1A84FFFC80CEA1419FE8694EA" blockId="44.[151,1437,944,2015]" pageId="44" pageNumber="45">Elytra with spaces between strial punctures narrow ridges; suture raised, especially at summit of posterior declivity; apices slightly widened, with submarginal groove having large translucent punctures and, apically, one large elongate puncture.</paragraph>
<paragraph id="8BB336F1A84FFFC80CEA148DFB1E94B6" blockId="44.[151,1437,944,2015]" box="[199,1279,1737,1763]" pageId="44" pageNumber="45">Ventral characters: Mentum coarsely punctate, with median fovea. Submentum with two foveae.</paragraph>
<paragraph id="8BB336F1A84FFFC80CEA14A9FC67951A" blockId="44.[151,1437,944,2015]" pageId="44" pageNumber="45">
Male genitalia (
<figureCitation id="13372A74A84FFFC80D5814A9FE279552" box="[373,454,1772,1799]" captionStart="FIGURE 25" captionStartId="45.[151,250,1794,1819]" captionTargetBox="[227,1381,186,1751]" captionTargetId="figure@45.[178,1410,181,1768]" captionTargetPageId="45" captionText="FIGURE 25. Hydrochus argutissimus n. sp., habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587233" httpUri="https://zenodo.org/record/3587233/files/figure.png" pageId="44" pageNumber="45">Fig. 25</figureCitation>
) general characters: paramere tips appearing sagittate,
<emphasis id="B978EAE3A84FFFC8080A14A9FAC79552" box="[1063,1318,1773,1799]" italics="true" pageId="44" pageNumber="45">but only in lateral view</emphasis>
; in dorsal/ ventral views tips have contiguous medial margins, which extend beyond distal end of aedeagus; basal piece long, proximal 1/2 strongly arcuate ventrad, with terminal sclerotization.
</paragraph>
<paragraph id="8BB336F1A84FFFC80CEA151DFC389526" blockId="44.[151,1437,944,2015]" box="[199,985,1881,1907]" pageId="44" pageNumber="45">Dorsal surface: adtl and adbl lacking; pdmm slightly overlapping alm.</paragraph>
<paragraph id="8BB336F1A84FFFC80CEA1539FC8F95C2" blockId="44.[151,1437,944,2015]" box="[199,878,1917,1943]" pageId="44" pageNumber="45">Ventral surface: pvmm closer to one another than are pdmm.</paragraph>
</subSubSection>
<subSubSection id="C316657AA84FFFC80CEA15E5FB9795EE" box="[199,1142,1953,1979]" pageId="44" pageNumber="45" type="etymology">
<paragraph id="8BB336F1A84FFFC80CEA15E5FB9795EE" blockId="44.[151,1437,944,2015]" box="[199,1142,1953,1979]" pageId="44" pageNumber="45">
<emphasis id="B978EAE3A84FFFC80CEA15E5FEAF95EE" bold="true" box="[199,334,1953,1979]" pageId="44" pageNumber="45">Etymology.</emphasis>
Named in reference to the relationship with
<taxonomicName id="4C0C4D72A84FFFC80F6F15E5FB9795EE" authority="Knisch." authorityName="Knisch." box="[834,1142,1953,1979]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="argutus">
<emphasis id="B978EAE3A84FFFC80F6F15E5FBF995EE" box="[834,1048,1953,1979]" italics="true" pageId="44" pageNumber="45">Hydrochus argutus</emphasis>
Knisch.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA84FFFC80CEA1581FB13958A" box="[199,1266,1989,2015]" pageId="44" pageNumber="45" type="discussion">
<paragraph id="8BB336F1A84FFFC80CEA1581FB13958A" blockId="44.[151,1437,944,2015]" box="[199,1266,1989,2015]" pageId="44" pageNumber="45">
<emphasis id="B978EAE3A84FFFC80CEA1581FEDA958A" bold="true" box="[199,315,1989,2015]" pageId="44" pageNumber="45">Remarks.</emphasis>
Images of the
<typeStatus id="54B78853A84FFFC80DCC1581FDA9958A" box="[481,584,1989,2015]" pageId="44" pageNumber="45" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C0C4D72A84FFFC80E401581FC78958A" authority="Knisch" authorityName="Knisch" box="[621,921,1989,2015]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="argutus">
<emphasis id="B978EAE3A84FFFC80E401581FCA2958A" box="[621,835,1989,2015]" italics="true" pageId="44" pageNumber="45">Hydrochus argutus</emphasis>
Knisch
</taxonomicName>
can be seen in
<bibRefCitation id="EF9D4B00A84FFFC808681581FB0F958A" author="Perkins, P. D." box="[1093,1262,1989,2015]" pageId="44" pageNumber="45" pagination="149 - 167" refId="ref30761" refString="Perkins, P. D. (2019) Type designations for Neotropical water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae). Zo otaxa, 47 01 (2), 149 - 167. https: // doi. org / 10.11646 / zootaxa. 4701.6.7" type="journal article" year="2019">Perkins (2019)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>