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<document id="CBEA8CC336281BC6C8AAEBE293A151B6" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A840FFC20CBA16B1FB029332" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus piroei Makhan" docType="treatment" docVersion="8" lastPageNumber="39" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="36" updateTime="1698775642300" updateUser="ExternalLinkService">
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<mods:title id="1A446E98B61990AAAD0656C7F45D5846">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
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<treatment id="03A587E7A840FFC20CBA16B1FB029332" ID-DOI="http://doi.org/10.5281/zenodo.5941799" ID-GBIF-Taxon="160798423" ID-Zenodo-Dep="5941799" LSID="urn:lsid:plazi:treatment:03A587E7A840FFC20CBA16B1FB029332" httpUri="http://treatment.plazi.org/id/03A587E7A840FFC20CBA16B1FB029332" lastPageId="38" lastPageNumber="39" pageId="35" pageNumber="36">
<subSubSection id="C316657AA840FFC70CBA16B1FE3D975A" box="[151,476,1269,1296]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BB336F1A840FFC70CBA16B1FE3D975A" blockId="35.[151,476,1269,1331]" box="[151,476,1269,1296]" pageId="35" pageNumber="36">
<heading id="D0FB819DA840FFC70CBA16B1FE3D975A" bold="true" box="[151,476,1269,1296]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<taxonomicName id="4C0C4D72A840FFC70CBA16B1FE3D975A" ID-CoL="6MJWP" authority="Makhan" authorityName="Makhan" box="[151,476,1269,1296]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="piroei">
<emphasis id="B978EAE3A840FFC70CBA16B1FE3D975A" bold="true" box="[151,476,1269,1296]" pageId="35" pageNumber="36">
<emphasis id="B978EAE3A840FFC70CBA16B1FE8B975A" bold="true" box="[151,362,1269,1296]" italics="true" pageId="35" pageNumber="36">Hydrochus piroei</emphasis>
Makhan
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA840FFC70CBA175DFEEC9766" box="[151,269,1305,1331]" pageId="35" pageNumber="36" type="description">
<paragraph id="8BB336F1A840FFC70CBA175DFEEC9766" blockId="35.[151,476,1269,1331]" box="[151,269,1305,1331]" pageId="35" pageNumber="36">
<figureCitation id="13372A74A840FFC70CBA175DFF049766" box="[151,229,1305,1331]" captionStart="FIGURE 2" captionStartId="14.[151,250,2009,2034]" captionTargetBox="[199,1400,435,1940]" captionTargetId="figure@14.[151,1435,417,1984]" captionTargetPageId="14" captionText="FIGURE 2. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587185" httpUri="https://zenodo.org/record/3587185/files/figure.png" pageId="35" pageNumber="36">Figs. 2</figureCitation>
,
<figureCitation id="13372A74A840FFC70CDC175DFEEC9766" box="[241,269,1305,1331]" captionStart="FIGURE 21" captionStartId="39.[151,250,1779,1804]" captionTargetBox="[223,1387,186,1752]" captionTargetId="figure@39.[151,1435,181,1753]" captionTargetPageId="39" captionText="FIGURE 21. Hydrochus piroei Makhan, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587225" httpUri="https://zenodo.org/record/3587225/files/figure.png" pageId="35" pageNumber="36">21</figureCitation>
</paragraph>
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<paragraph id="8BB336F1A840FFC70CBA171CFDEF9725" blockId="35.[151,526,1367,1392]" box="[151,526,1367,1392]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="AB1C11DFA840FFC70CBA171CFDEF9725" box="[151,526,1367,1392]" pageId="35" pageNumber="36">
<taxonomicName id="4C0C4D72A840FFC70CBA171CFDE89725" ID-CoL="6MJWP" authority="Makhan, 1992: 97" authorityName="Makhan" authorityPageNumber="97" authorityYear="1992" box="[151,521,1367,1392]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="piroei">
<emphasis id="B978EAE3A840FFC70CBA171CFEA99725" box="[151,328,1368,1392]" italics="true" pageId="35" pageNumber="36">Hydrochus piroei</emphasis>
<treatmentCitation id="0AAD10E0A840FFC70D631713FDE89725" author="Makhan, D." box="[334,521,1367,1392]" page="97" pageId="35" pageNumber="36" year="1992">
<bibRefCitation id="EF9D4B00A840FFC70D631713FDE89725" author="Makhan, D." box="[334,521,1367,1392]" pageId="35" pageNumber="36" pagination="95 - 103" refId="ref30207" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan, 1992: 97</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="C316657AA840FFC70CBA17DCFA7A9432" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BB336F1A840FFC70CBA17DCFD7497AE" blockId="35.[151,1437,1432,2035]" pageId="35" pageNumber="36">
<emphasis id="B978EAE3A840FFC70CBA17DCFE5597E6" bold="true" box="[151,436,1432,1459]" pageId="35" pageNumber="36">Material examined (10).</emphasis>
<materialsCitation id="3B643CACA840FFC70D9717DDFC2A97E6" ID-GBIF-Occurrence="2521263680" box="[442,971,1432,1459]" collectionCode="SEMC" country="Venezuela" location="Sucre" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Sucre">
<emphasis id="B978EAE3A840FFC70D9717DDFD6F97E6" bold="true" box="[442,654,1432,1459]" pageId="35" pageNumber="36">
<collectingCountry id="F31B7661A840FFC70D9717DDFDD697E6" box="[442,567,1433,1459]" name="Venezuela" pageId="35" pageNumber="36">Venezuela</collectingCountry>
:
<collectingRegion id="49C8F813A840FFC70E6C17DCFD6897E6" box="[577,649,1432,1459]" country="Venezuela" name="Sucre" pageId="35" pageNumber="36">Sucre</collectingRegion>
:
</emphasis>
VZ10-0129-04A (2
<collectionCode id="ED1DAE34A840FFC70F5F17DDFC2297E6" box="[882,963,1433,1459]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="35" pageNumber="36" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA840FFC70FF817DCFEE99782" ID-GBIF-Occurrence="2521263663" collectionCode="SEMC, MALUZ" country="Venezuela" location="Anzoategui" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Anzoategui">
<emphasis id="B978EAE3A840FFC70FF817DCFB8497E6" bold="true" box="[981,1125,1432,1459]" pageId="35" pageNumber="36">
<collectingRegion id="49C8F813A840FFC70FF817DCFB8097E6" box="[981,1121,1432,1459]" country="Venezuela" name="Anzoategui" pageId="35" pageNumber="36">Anzoategui</collectingRegion>
:
</emphasis>
VZ10-0204-01A (3
<collectionCode id="ED1DAE34A840FFC7096417DDFA7997E6" box="[1353,1432,1433,1459]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="35" pageNumber="36" type="Museum">SEMC</collectionCode>
,
<collectionCode id="ED1DAE34A840FFC70CBA17F9FEE19782" box="[151,256,1469,1495]" pageId="35" pageNumber="36">MALUZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA840FFC70D3F17F8FA789782" ID-GBIF-Occurrence="2521263692" box="[274,1433,1468,1495]" collectingDate="1989-08-03" collectionCode="CMN" collectorName="S. &amp; J. Peck" country="Venezuela" elevation="1" location="Lake Maracaibo" municipality="Palmarito" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Merida">
<emphasis id="B978EAE3A840FFC70D3F17F8FE959782" bold="true" box="[274,372,1468,1495]" pageId="35" pageNumber="36">
<collectingRegion id="49C8F813A840FFC70D3F17F8FE8E9782" box="[274,367,1468,1495]" country="Venezuela" name="Merida" pageId="35" pageNumber="36">Merida</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BD7AC8BA840FFC70D5717F9FE0B9782" box="[378,490,1469,1495]" pageId="35" pageNumber="36">Palmarito</collectingMunicipality>
,
<location id="8ED3602AA840FFC70DD817F9FD4F9782" LSID="urn:lsid:plazi:treatment:03A587E7A840FFC20CBA16B1FB029332:8ED3602AA840FFC70DD817F9FD4F9782" box="[501,686,1469,1495]" country="Venezuela" municipality="Palmarito" name="Lake Maracaibo" pageId="35" pageNumber="36" stateProvince="Merida">Lake Maracaibo</location>
,
<quantity id="4CF49B14A840FFC70E9517F9FD3F9782" box="[696,734,1469,1495]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="35" pageNumber="36" unit="m" value="1.0">
<elevation id="0021D1C2A840FFC70E9517F9FD3F9782" box="[696,734,1469,1495]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="35" pageNumber="36" unit="m" value="1.0">1m</elevation>
</quantity>
,
<date id="FFB21031A840FFC70EC717F9FC919782" box="[746,880,1469,1495]" pageId="35" pageNumber="36" value="1989-08-03">
<collectingDate id="EFF6E9D9A840FFC70EC717F9FC919782" box="[746,880,1469,1495]" pageId="35" pageNumber="36" value="1989-08-03">3.VIII.1989</collectingDate>
</date>
,
<collectorName id="26F95327A840FFC70F5617F9FC189782" box="[891,1017,1469,1495]" pageId="35" pageNumber="36">S.&amp;J. Peck</collectorName>
, lakeshore, ex: UV, 89-285 (2
<collectionCode id="ED1DAE34A840FFC7096117F9FA709782" box="[1356,1425,1469,1495]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="35" pageNumber="36" type="Museum">CMN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA840FFC70CBA17A4FD7097AE" ID-GBIF-Occurrence="2521263659" box="[151,657,1504,1531]" collectionCode="SEMC, MIZA" country="Venezuela" location="Trujillo" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Trujillo">
<emphasis id="B978EAE3A840FFC70CBA17A4FF1D97AE" bold="true" box="[151,252,1504,1531]" pageId="35" pageNumber="36">
<collectingRegion id="49C8F813A840FFC70CBA17A4FF1697AE" box="[151,247,1504,1531]" country="Venezuela" name="Trujillo" pageId="35" pageNumber="36">Trujillo</collectingRegion>
:
</emphasis>
VZ12-0128-06A (3
<collectionCode id="ED1DAE34A840FFC70DCF17A5FDD097AE" box="[482,561,1505,1531]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="35" pageNumber="36" type="Museum">SEMC</collectionCode>
,
<collectionCode id="ED1DAE34A840FFC70E1117A5FD6897AE" box="[572,649,1505,1531]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="35" pageNumber="36" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB336F1A840FFC70CEA1440FA7A9432" blockId="35.[151,1437,1432,2035]" pageId="35" pageNumber="36">
<emphasis id="B978EAE3A840FFC70CEA1440FE30944A" bold="true" box="[199,465,1540,1567]" pageId="35" pageNumber="36">Differential Diagnosis.</emphasis>
Among Venezuelan species, recognized by the combination of moderate size (ca.
<quantity id="4CF49B14A840FFC709701440FEE09416" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.86" metricValueMin="2.54" pageId="35" pageNumber="36" unit="mm" value="2.7" valueMax="2.86" valueMin="2.54">2.54 2.86 mm</quantity>
), the moderately low and relatively punctate pronotal reliefs, and the male genitalia (
<figureCitation id="13372A74A840FFC708E3146DFAC29416" box="[1230,1315,1577,1603]" captionStart="FIGURE 21" captionStartId="39.[151,250,1779,1804]" captionTargetBox="[223,1387,186,1752]" captionTargetId="figure@39.[151,1435,181,1753]" captionTargetPageId="39" captionText="FIGURE 21. Hydrochus piroei Makhan, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587225" httpUri="https://zenodo.org/record/3587225/files/figure.png" pageId="35" pageNumber="36">Fig. 21</figureCitation>
; described below). This species has a generalized
<taxonomicName id="4C0C4D72A840FFC70E661409FD249432" authorityName="Leach" authorityYear="1817" box="[587,709,1613,1639]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAE3A840FFC70E661409FD249432" box="[587,709,1613,1639]" italics="true" pageId="35" pageNumber="36">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C316657AA840FFC20CEA1434FF28934A" lastPageId="38" lastPageNumber="39" pageId="35" pageNumber="36" type="description">
<paragraph id="8BB336F1A840FFC70CEA1434FD44956A" blockId="35.[151,1437,1432,2035]" pageId="35" pageNumber="36">
<emphasis id="B978EAE3A840FFC70CEA1434FEB094DE" bold="true" box="[199,337,1648,1675]" pageId="35" pageNumber="36">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="13372A74A840FFC70E4B1435FD5D94DE" box="[614,700,1649,1675]" captionStart="FIGURE 21" captionStartId="39.[151,250,1779,1804]" captionTargetBox="[223,1387,186,1752]" captionTargetId="figure@39.[151,1435,181,1753]" captionTargetPageId="39" captionText="FIGURE 21. Hydrochus piroei Makhan, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587225" httpUri="https://zenodo.org/record/3587225/files/figure.png" pageId="35" pageNumber="36">Fig. 21</figureCitation>
). Size of imaged specimen (length/width, mm): body (length to elytral apices) 2.86/1.03; head width 0.70; pronotum l/w 0.65/0.68; PA 0.65; PB 0.48; elytra 1.73/1.03. Body size moderate (ca.
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). Dorsum grey, with slight iridescence, elytra with black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 1.5
<date id="FFB21031A840FFC70ED61498FCF694A2" box="[763,791,1756,1783]" pageId="35" pageNumber="36">2x</date>
that of pronotum. Elytra interstriae ca. 11.5x strial puncture diameter. Interstria 9
<superScript id="7C799BB9A840FFC70D941544FE27955B" attach="left" box="[441,454,1792,1806]" fontSize="6" pageId="35" pageNumber="36">th</superScript>
more convex than others, overhanging 10
<superScript id="7C799BB9A840FFC70F861544FC59955B" attach="left" box="[939,952,1792,1806]" fontSize="6" pageId="35" pageNumber="36">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C799BB9A840FFC709781544FA83955B" attach="left" box="[1365,1378,1792,1806]" fontSize="6" pageId="35" pageNumber="36">th</superScript>
very slightly raised, indistinct, moderately elongate.
</paragraph>
<paragraph id="8BB336F1A840FFC70CEA150DFDDA95FE" blockId="35.[151,1437,1432,2035]" pageId="35" pageNumber="36">Pronotum slightly wider than long, widest slightly behind anterior margin, narrowed at base, sides arcuate or very weakly sinuate, smooth; anterior margin slightly bisinuate; depressions shallow, much more densely punctate than low reliefs between depressions.</paragraph>
<paragraph id="8BB336F1A840FFC70CEA15F1FC9A95A6" blockId="35.[151,1437,1432,2035]" pageId="35" pageNumber="36">
Elytra with spaces between strial punctures ca. 0.5
<date id="FFB21031A840FFC70F2215F0FCC6959A" box="[783,807,1972,1999]" pageId="35" pageNumber="36">1x</date>
their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with slight angulation.
</paragraph>
<paragraph id="8BB336F1A847FFC00CEA12DDFE529282" blockId="36.[151,1436,153,215]" pageId="36" pageNumber="37">Ventral characters: Mentum moderately coarsely punctate, with median fovea. Submentum with two very large, almost confluent, foveae.</paragraph>
<caption id="DF736679A847FFC00CBA1519FCB49523" ID-DOI="http://doi.org/10.5281/zenodo.3587219" ID-Zenodo-Dep="3587219" box="[151,853,1885,1910]" httpUri="https://zenodo.org/record/3587219/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1885,1910]" targetBox="[226,1371,295,1841]" targetPageId="36">
<paragraph id="8BB336F1A847FFC00CBA1519FCB49523" blockId="36.[151,853,1885,1910]" box="[151,853,1885,1910]" pageId="36" pageNumber="37">
<emphasis id="B978EAE3A847FFC00CBA1519FEC19523" bold="true" box="[151,288,1885,1910]" pageId="36" pageNumber="37">FIGURE 18.</emphasis>
<taxonomicName id="4C0C4D72A847FFC00D0B151AFDDB9523" authority="Sharp" authorityName="Sharp" box="[294,570,1885,1910]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B978EAE3A847FFC00D0B151AFE759523" box="[294,404,1886,1910]" italics="true" pageId="36" pageNumber="37">Hydrochus</emphasis>
<emphasis id="B978EAE3A847FFC00DB6151AFE179523" box="[411,502,1886,1910]" italics="true" pageId="36" pageNumber="37">obscurus</emphasis>
Sharp
</taxonomicName>
, habitus and male genitalia.
</paragraph>
</caption>
<caption id="DF736679A846FFC10CBA1546FC14954E" ID-DOI="http://doi.org/10.5281/zenodo.3587221" ID-Zenodo-Dep="3587221" box="[151,1013,1794,1819]" httpUri="https://zenodo.org/record/3587221/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,250,1794,1819]" targetBox="[199,1384,188,1730]" targetPageId="37">
<paragraph id="8BB336F1A846FFC10CBA1546FC14954E" blockId="37.[151,1013,1794,1819]" box="[151,1013,1794,1819]" pageId="37" pageNumber="38">
<emphasis id="B978EAE3A846FFC10CBA1546FEC1954E" bold="true" box="[151,288,1794,1819]" pageId="37" pageNumber="38">FIGURE 19.</emphasis>
<taxonomicName id="4C0C4D72A846FFC10D0B1546FDC0954F" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[294,545,1794,1818]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="variabiloides" status="sp. nov.">
<emphasis id="B978EAE3A846FFC10D0B1546FDC0954F" box="[294,545,1794,1818]" italics="true" pageId="37" pageNumber="38">Hydrochus variabiloides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5798A846FFC10E0A1547FDBE954F" box="[551,607,1795,1818]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<paragraph id="8BB336F1A846FFC10CEA150DFA9695A6" blockId="37.[151,1437,1865,2035]" pageId="37" pageNumber="38">
Male genitalia (
<figureCitation id="13372A74A846FFC10D54150DFE2A9536" box="[377,459,1865,1891]" captionStart="FIGURE 21" captionStartId="39.[151,250,1779,1804]" captionTargetBox="[223,1387,186,1752]" captionTargetId="figure@39.[151,1435,181,1753]" captionTargetPageId="39" captionText="FIGURE 21. Hydrochus piroei Makhan, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587225" httpUri="https://zenodo.org/record/3587225/files/figure.png" pageId="37" pageNumber="38">Fig. 21</figureCitation>
) general characters: genitalia rather narrow, elongate; parameres slightly but distinctly longer than basal piece, narrow in dorsal/ventral views, in lateral view much wider and slightly arcuate, tips extending very slightly beyond distal apex of aedeagus, tips rounded, medial margins straight, almost parallel; aedeagus lanceolate, with sclerotized lateral margins; basal piece distinctive appearing laterally compressed over distal ½, lateral margins strongly sclerotized in this area, rim of orifice strongly sclerotized and apicoventrally produced.
</paragraph>
<paragraph id="8BB336F1A845FFC20CEA12DDFE7C9282" blockId="38.[151,1437,153,359]" pageId="38" pageNumber="39">Dorsal surface: adtl subbasal, hence membranous area of aedeagus large; pdmm slightly overlapping aedeagus in basal 1/3; adbl large.</paragraph>
<paragraph id="8BB336F1A845FFC20CEA12A5FF28934A" blockId="38.[151,1437,153,359]" pageId="38" pageNumber="39">Ventral surface: avtl subbasal, hence membranous area of aedeagus large; pvmm more or less contiguous with alm.</paragraph>
</subSubSection>
<subSubSection id="C316657AA845FFC20CEA136DFB029332" pageId="38" pageNumber="39" type="discussion">
<paragraph id="8BB336F1A845FFC20CEA136DFB029332" blockId="38.[151,1437,153,359]" pageId="38" pageNumber="39">
<emphasis id="B978EAE3A845FFC20CEA136DFEDA9316" bold="true" box="[199,315,297,323]" pageId="38" pageNumber="39">Remarks.</emphasis>
The specimen imaged (
<figureCitation id="13372A74A845FFC20E65136DFD789316" box="[584,665,297,323]" captionStart="FIGURE 21" captionStartId="39.[151,250,1779,1804]" captionTargetBox="[223,1387,186,1752]" captionTargetId="figure@39.[151,1435,181,1753]" captionTargetPageId="39" captionText="FIGURE 21. Hydrochus piroei Makhan, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587225" httpUri="https://zenodo.org/record/3587225/files/figure.png" pageId="38" pageNumber="39">Fig. 21</figureCitation>
) is from locality VZ10-0129-04A. I was not able to examine the
<typeStatus id="54B78853A845FFC2095A136DFF3C9332" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C0C4D72A845FFC20D2F130AFE869332" box="[258,359,334,359]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="piroei">
<emphasis id="B978EAE3A845FFC20D2F130AFE869332" box="[258,359,334,359]" italics="true" pageId="38" pageNumber="39">H. piroei</emphasis>
</taxonomicName>
; its location is unknown. My concept is based on the examination of
<typeStatus id="54B78853A845FFC2085D1309FB3E9332" box="[1136,1247,333,359]" pageId="38" pageNumber="39" type="paratype">paratypes</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>