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<document id="879C1E57A64E5D54DD41041C07CCBCA5" ID-CLB-Dataset="8756" ID-DOI="10.5852/ejt.2021.731.1209" ID-GBIF-Dataset="3cf2c6ad-4274-4301-9cd0-e8295eace8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4452219" ID-ZooBank="070B1EBB-976A-4EBB-B37D-6CB4A10D09FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611143662020" checkinUser="plazi" docAuthor="Vargas, Héctor A." docDate="2021" docId="03A587BDBB2B5E6761DD61369F876559" docLanguage="en" docName="ejt-731_117-134.pdf" docOrigin="European Journal of Taxonomy 731" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Helioandesia Vargas 2021, gen. nov." docType="treatment" docUuid="2ACDF84E-8E2D-43C6-90D5-B271DC332483" docUuidSource="ZooBank" docVersion="8" lastPageNumber="125" masterDocId="FF9CFFC5BB2D5E6C634C625F9D00641F" masterDocTitle="Systematics of Helioandesia tarregai gen. et sp. nov. (Lepidoptera: Yponomeutoidea: Heliodinidae) from the Andes of Northern Chile" masterLastPageNumber="134" masterPageNumber="117" pageNumber="123" updateTime="1698924879965" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4E5B2AF08FD773FC785E114CC88006C0">Systematics of Helioandesia tarregai gen. et sp. nov. (Lepidoptera: Yponomeutoidea: Heliodinidae) from the Andes of Northern Chile</mods:title>
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<mods:name id="031AF02831944E3BAD85EF7048418876" type="personal">
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<mods:namePart id="0C551EDED38999B145E719F8BE57B5FD">Vargas, Héctor A.</mods:namePart>
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<mods:nameIdentifier id="3F0A4C085B44D3C5A418B69A49B2E98A" type="ORCID">0000-0002-5355-3157</mods:nameIdentifier>
<mods:affiliation id="B98B4208A9BB0F08CBEF596621662DB7">Departamento de Recursos Ambientales, Facultad de Ciencias Agronómicas, Universidad de Tarapacá, Arica, Chile.</mods:affiliation>
<mods:nameIdentifier id="2B1EA82F5E86545BADC641DBC0426701" type="email">havargas@academicos.uta.cl; lepvargas@gmail.com</mods:nameIdentifier>
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<mods:title id="12B435B6E3BF73772DA563E61C74ADDD">European Journal of Taxonomy</mods:title>
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<mods:part id="051B61C77F43506E85971EE97FD51E74">
<mods:date id="6D5636B9D1531163C1D3CEE71B872AE0">2021</mods:date>
<mods:detail id="2FB5660D29106405F3A8153C561ACC3F" type="pubDate">
<mods:number id="D91C0B3A375149E126A97D92AF132981">2021-01-20</mods:number>
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<mods:number id="FC33A6EBE77DA1EAB130724F1AC71CC9">731</mods:number>
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<mods:classification id="66A5B7F99204EBF7667C8DFB99C67A0D">journal article</mods:classification>
<mods:identifier id="F4F2062345CD78A5044A8A48B1B86947" type="CLB-Dataset">8756</mods:identifier>
<mods:identifier id="1F4E043C72F6E3D78895CD3276CF9ED8" type="DOI">10.5852/ejt.2021.731.1209</mods:identifier>
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<treatment id="03A587BDBB2B5E6761DD61369F876559" ID-DOI="http://doi.org/10.5281/zenodo.4452221" ID-GBIF-Taxon="177010520" ID-Zenodo-Dep="4452221" LSID="urn:lsid:zoobank.org:act:2ACDF84E-8E2D-43C6-90D5-B271DC332483" httpUri="http://treatment.plazi.org/id/03A587BDBB2B5E6761DD61369F876559" lastPageId="11" lastPageNumber="125" pageId="6" pageNumber="123">
<subSubSection id="C3166520BB2B5E6A61DD61369EA3679B" box="[657,931,873,900]" pageId="6" pageNumber="123" type="nomenclature">
<paragraph id="8BB336ABBB2B5E6A61DD61369EA3679B" blockId="6.[363,1225,873,970]" box="[657,931,873,900]" pageId="6" pageNumber="123">
<heading id="D0FB81C7BB2B5E6A61DD61369EA3679B" box="[657,931,873,900]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="123" reason="2">
<taxonomicName id="4C0C4D28BB2B5E6A61DD61369E31679B" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[657,817,873,900]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B978EAB9BB2B5E6A61DD61369E31679B" bold="true" box="[657,817,873,900]" italics="true" pageId="6" pageNumber="123">Helioandesia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB2B5E6A607461359EA3679B" box="[824,931,874,900]" pageId="6" pageNumber="123" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3166520BB2B5E6A622761D39E4C67D5" pageId="6" pageNumber="123" type="description">
<paragraph id="8BB336ABBB2B5E6A622761D399C967B8" blockId="6.[363,1225,873,970]" box="[363,1225,908,935]" pageId="6" pageNumber="123">
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urn:lsid:zoobank.org:act:
<uuid id="FFAA0C7EBB2B5E6A61C661D399C967B8" box="[650,1225,908,935]" pageId="6" pageNumber="123">2ACDF84E-8E2D-43C6-90D5-B271DC332483</uuid>
</uri>
</paragraph>
<paragraph id="8BB336ABBB2B5E6A61AB61EF9E4C67D5" blockId="6.[363,1225,873,970]" box="[743,844,944,970]" pageId="6" pageNumber="123">
<figureCitation id="13372A2EBB2B5E6A61AB61EF9E4C67D5" box="[743,844,944,970]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="7.[189,232,1928,1954]" captionStartId-1="8.[188,231,1963,1989]" captionStartId-2="9.[189,232,1857,1883]" captionStartId-3="10.[191,234,1927,1953]" captionTargetBox-0="[189,1398,757,1889]" captionTargetBox-1="[190,1397,1272,1927]" captionTargetBox-2="[327,1261,319,1818]" captionTargetBox-3="[189,1398,559,1861]" captionTargetId-0="figure-163@7.[189,1398,757,1889]" captionTargetId-1="figure-364@8.[190,1397,1272,1927]" captionTargetId-2="figure-11@9.[326,1261,319,1818]" captionTargetId-3="figure-92@10.[189,1398,559,1861]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Fig. 3. Helioandesia tarregai gen. et sp. nov., ♂, holotype (IDEA-LEPI-2020-015). A. Dorsal view. B. Head, lateral. C. Basal flagellomeres of right antenna. D. Three bulging metallic spots on the basal half of forewing. E. Hair pencil from base of subcosta, ventral. Scale bar: 1 mm." captionText-1="Fig. 4. Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024), wing venation showing frenulum with two bristles of unequal length. Scale bar: 1 mm." captionText-2="Fig. 5. Male genitalia and eighth segment of Helioandesia tarregai gen. et sp. nov., ♂, paratype (IDEA- LEPI-2020-021). A. Male genitalia, ventral view, phallus removed. B. Phallus, lateral view. C. Tergum and pleural lobes of eighth segment. D. Socius, closed arrow in A. E. Small spine-like cornuti on the vesica, open arrow in A. F. Medial part of the cornutus. G. Coremata between pleural lobes of segment VIII and genitalia. Scale bars: 0.1 mm." captionText-3="Fig. 6. Female genitalia of Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024) A. Female genitalia, ventral view. B. Papillae analis. C. Ornamentation of the proximal part of the signum. D. Ornamentation of the distal part of the signum. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4452227" figureDoi-1="http://doi.org/10.5281/zenodo.4452229" figureDoi-2="http://doi.org/10.5281/zenodo.4452231" figureDoi-3="http://doi.org/10.5281/zenodo.4452233" httpUri-0="https://zenodo.org/record/4452227/files/figure.png" httpUri-1="https://zenodo.org/record/4452229/files/figure.png" httpUri-2="https://zenodo.org/record/4452231/files/figure.png" httpUri-3="https://zenodo.org/record/4452233/files/figure.png" pageId="6" pageNumber="123">Figs 3–6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3166520BB2B5E6A63F161AE9E456025" pageId="6" pageNumber="123" type="type_taxon">
<paragraph id="8BB336ABBB2B5E6A63F161AE9C586014" blockId="6.[189,344,1009,1035]" box="[189,344,1009,1035]" pageId="6" pageNumber="123">
<heading id="D0FB81C7BB2B5E6A63F161AE9C586014" bold="true" box="[189,344,1009,1035]" fontSize="11" level="3" pageId="6" pageNumber="123" reason="3">
<emphasis id="B978EAB9BB2B5E6A63F161AE9C586014" bold="true" box="[189,344,1009,1035]" pageId="6" pageNumber="123">
<typeStatus id="54B78809BB2B5E6A63F161AE9DFA6014" box="[189,250,1009,1035]" pageId="6" pageNumber="123">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336ABBB2B5E6A63F1667F9E456025" blockId="6.[189,837,1056,1082]" box="[189,837,1056,1082]" pageId="6" pageNumber="123">
<taxonomicName id="4C0C4D28BB2B5E6A63F1667F9CC06026" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[189,448,1056,1082]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB2B5E6A63F1667F9CC06026" box="[189,448,1056,1082]" italics="true" pageId="6" pageNumber="123">Helioandesia tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB2B5E6A628B667F9F7A6025" box="[455,634,1056,1082]" pageId="6" pageNumber="123" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, designated here.
</paragraph>
</subSubSection>
<subSubSection id="C3166520BB2B5E6A63F1663E9C2E6359" pageId="6" pageNumber="123" type="diagnosis">
<paragraph id="8BB336ABBB2B5E6A63F1663E9C366064" blockId="6.[189,310,1121,1147]" box="[189,310,1121,1147]" pageId="6" pageNumber="123">
<heading id="D0FB81C7BB2B5E6A63F1663E9C366064" bold="true" box="[189,310,1121,1147]" fontSize="11" level="3" pageId="6" pageNumber="123" reason="3">
<emphasis id="B978EAB9BB2B5E6A63F1663E9C366064" bold="true" box="[189,310,1121,1147]" pageId="6" pageNumber="123">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336ABBB2B5E6A63F166CF9C2E6359" blockId="6.[188,1401,1168,1862]" pageId="6" pageNumber="123">
<taxonomicName id="4C0C4D28BB2B5E6A63F166CF9C5960B5" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[189,345,1168,1194]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B978EAB9BB2B5E6A63F166CF9C5960B5" box="[189,345,1168,1194]" italics="true" pageId="6" pageNumber="123">Helioandesia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB2B5E6A622E66CF9CCF60B5" box="[354,463,1168,1194]" pageId="6" pageNumber="123" rank="genus">gen. nov.</taxonomicNameLabel>
can be recognized by the following combination of morphological characters: (1) presence of CuP on forewing of female and male; (2) two bristles of unequal length in the female frenulum; (3) forewing mostly gray, ornamented with strongly bulging metallic spots, (4) two triangular processes on anterior margin of tergum VIII in the male abdomen; (5) sclerotization of the widened proximal part of the ductus bursae not reaching the ostium; (6) signum with proximal portion coarser than distal portion; (7) presence of an appendix bursae; (8) slightly swollen distal end of the saccus; and (9) a well-developed cornutus on the vesica. The mostly gray forewing pattern of
<taxonomicName id="4C0C4D28BB2B5E6A6796673C98766162" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[1242,1398,1379,1405]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B978EAB9BB2B5E6A6796673C98766162" box="[1242,1398,1379,1405]" italics="true" pageId="6" pageNumber="123">Helioandesia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB2B5E6A63F167D89C2F61BE" box="[189,303,1415,1441]" pageId="6" pageNumber="123" rank="genus">gen. nov.</taxonomicNameLabel>
, ornamented with strongly bulging metallic spots, resembles that of the mainly Nearctic
<taxonomicName id="4C0C4D28BB2B5E6A63F167F69C6661DC" authorityName="Chambers" authorityYear="1876" box="[189,358,1449,1475]" class="Insecta" family="Heliodinidae" genus="Lithariapteryx" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB9BB2B5E6A63F167F69C6661DC" box="[189,358,1449,1475]" italics="true" pageId="6" pageNumber="123">Lithariapteryx</emphasis>
</taxonomicName>
. However, the latter lacks CuP in the forewing of female and male, has a single bristle in the female frenulum, and lacks a well-developed cornutus on the vesica. The female genitalia of
<taxonomicName id="4C0C4D28BB2B5E6A63F167AF9C406216" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[189,320,1520,1545]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB2B5E6A63F167AF9C406216" box="[189,320,1520,1545]" italics="true" pageId="6" pageNumber="123">H. tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB2B5E6A620B67AF9CF76215" box="[327,503,1520,1546]" pageId="6" pageNumber="123" rank="species">gen. et sp. nov.</taxonomicNameLabel>
are very similar to those of the Bolivian
<taxonomicName id="4C0C4D28BB2B5E6A609A67AF9C016232" authority="(Meyrick, 1932)" baseAuthorityName="Meyrick" baseAuthorityYear="1932" class="Insecta" family="Heliodinidae" genus="Lithariapteryx" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="loriculata">
<emphasis id="B978EAB9BB2B5E6A609A67AF99FB6215" box="[982,1275,1520,1546]" italics="true" pageId="6" pageNumber="123">Lithariapteryx loriculata</emphasis>
(Meyrick, 1932)
</taxonomicName>
in the shape of the transverse bridge, signum and antrum. However,
<taxonomicName id="4C0C4D28BB2B5E6A6774644B99BC6232" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[1080,1212,1556,1581]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB2B5E6A6774644B99BC6232" box="[1080,1212,1556,1581]" italics="true" pageId="6" pageNumber="123">H. tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB2B5E6A678F644C98776232" box="[1219,1399,1555,1581]" pageId="6" pageNumber="123" rank="species">gen. et sp. nov.</taxonomicNameLabel>
lacks a V-shaped mark from costa on the forewing, has a sclerotization on the distal part of the ductus bursae and has the signum on the left margin of the corpus bursae, while
<taxonomicName id="4C0C4D28BB2B5E6A6765640599BD626C" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[1065,1213,1625,1651]" class="Insecta" family="Heliodinidae" genus="Lithariapteryx" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="loriculata">
<emphasis id="B978EAB9BB2B5E6A6765640599BD626C" box="[1065,1213,1625,1651]" italics="true" pageId="6" pageNumber="123">L. loriculata</emphasis>
</taxonomicName>
has a V-shaped mark from the costa on the forewing, lacks sclerotization on the distal part of the ductus bursae and has the signum on the middle of the ventral part of the corpus bursae. The male of
<taxonomicName id="4C0C4D28BB2B5E6A673164FF981362A5" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[1149,1299,1696,1722]" class="Insecta" family="Heliodinidae" genus="Lithariapteryx" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="loriculata">
<emphasis id="B978EAB9BB2B5E6A673164FF981362A5" box="[1149,1299,1696,1722]" italics="true" pageId="6" pageNumber="123">L. loriculata</emphasis>
</taxonomicName>
remains unknown, impeding comparison with
<taxonomicName id="4C0C4D28BB2B5E6A6133649B9E0262C2" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[639,770,1732,1757]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB2B5E6A6133649B9E0262C2" box="[639,770,1732,1757]" italics="true" pageId="6" pageNumber="123">H. tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB2B5E6A6047649C9EBD62C2" box="[779,957,1731,1757]" pageId="6" pageNumber="123" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Two bristles of unequal length in the female frenulum of
<taxonomicName id="4C0C4D28BB2B5E6A62FC64B99F4C631F" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[432,588,1766,1792]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B978EAB9BB2B5E6A62FC64B99F4C631F" box="[432,588,1766,1792]" italics="true" pageId="6" pageNumber="123">Helioandesia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB2B5E6A611B64B89FC5631E" box="[599,709,1767,1793]" pageId="6" pageNumber="123" rank="genus">gen. nov.</taxonomicNameLabel>
resemble those of
<taxonomicName id="4C0C4D28BB2B5E6A60E264B899E1631F" authority="Meyrick, 1907" authorityName="Meyrick" authorityYear="1907" box="[942,1249,1766,1792]" class="Insecta" family="Heliodinidae" genus="Epicroesa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB9BB2B5E6A60E264B89924631F" box="[942,1060,1767,1792]" italics="true" pageId="6" pageNumber="123">Epicroesa</emphasis>
Meyrick, 1907
</taxonomicName>
,
<taxonomicName id="4C0C4D28BB2B5E6A67BC64B9986F631F" authorityName="Stainton" authorityYear="1854" box="[1264,1391,1766,1792]" class="Insecta" family="Heliodinidae" genus="Heliodines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB9BB2B5E6A67BC64B9986F631F" box="[1264,1391,1766,1792]" italics="true" pageId="6" pageNumber="123">Heliodines</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0C4D28BB2B5E6A63F165569F00633C" authority="Busck, 1900" authorityName="Busck" authorityYear="1900" box="[189,512,1801,1827]" class="Insecta" family="Heliodinidae" genus="Lamprolophus" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB9BB2B5E6A63F165569C67633C" box="[189,359,1801,1827]" italics="true" pageId="6" pageNumber="123">Lamprolophus</emphasis>
Busck, 1900
</taxonomicName>
and
<taxonomicName id="4C0C4D28BB2B5E6A617465569E85633C" authority="Meyrick, 1927" authorityName="Meyrick" authorityYear="1927" box="[568,901,1801,1827]" class="Insecta" family="Heliodinidae" genus="Philocoristis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB9BB2B5E6A617465569FCE633C" box="[568,718,1801,1827]" italics="true" pageId="6" pageNumber="123">Philocoristis</emphasis>
Meyrick, 1927
</taxonomicName>
. However, all these genera lack CuP in the forewing.
</paragraph>
</subSubSection>
<subSubSection id="C3166520BB2A5E6B63F1635699966588" pageId="7" pageNumber="124" type="etymology">
<paragraph id="8BB336ABBB2A5E6B63F163569C44653C" blockId="7.[189,324,265,291]" box="[189,324,265,291]" pageId="7" pageNumber="124">
<heading id="D0FB81C7BB2A5E6B63F163569C44653C" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="7" pageNumber="124" reason="3">
<emphasis id="B978EAB9BB2A5E6B63F163569C44653C" bold="true" box="[189,324,265,291]" pageId="7" pageNumber="124">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336ABBB2A5E6B63F1636899966588" blockId="7.[189,1398,311,408]" pageId="7" pageNumber="124">
The genus name is derived from the Greek ʻ
<emphasis id="B978EAB9BB2A5E6B619563689E1F654E" box="[729,799,311,337]" italics="true" pageId="7" pageNumber="124">helios</emphasis>
ʼ (sun), the root of the family name
<taxonomicName id="4C0C4D28BB2A5E6B679463689873654E" authorityName="Heinemann" authorityYear="1877" box="[1240,1395,311,337]" class="Insecta" family="Heliodinidae" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="family">Heliodinidae</taxonomicName>
, and Andes, in reference to the Andes Range, among whose amazing landscapes the
<typeStatus id="54B78809BB2A5E6B678D630499F4656A" box="[1217,1268,347,373]" pageId="7" pageNumber="124">type</typeStatus>
species of
<taxonomicName id="4C0C4D28BB2A5E6B63F163229C596588" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[189,345,381,407]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B978EAB9BB2A5E6B63F163229C596588" box="[189,345,381,407]" italics="true" pageId="7" pageNumber="124">Helioandesia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB2A5E6B622D63219CCC6587" box="[353,460,382,408]" pageId="7" pageNumber="124" rank="genus">gen. nov.</taxonomicNameLabel>
was discovered. The name is considered feminine in gender.
</paragraph>
</subSubSection>
<subSubSection id="C3166520BB2A5E6763F163E09F876559" lastPageId="11" lastPageNumber="128" pageId="7" pageNumber="124" type="description">
<paragraph id="8BB336ABBB2A5E6B63F163E09C4E65C6" blockId="7.[189,334,447,473]" box="[189,334,447,473]" pageId="7" pageNumber="124">
<heading id="D0FB81C7BB2A5E6B63F163E09C4E65C6" bold="true" box="[189,334,447,473]" fontSize="11" level="3" pageId="7" pageNumber="124" reason="3">
<emphasis id="B978EAB9BB2A5E6B63F163E09C4E65C6" bold="true" box="[189,334,447,473]" pageId="7" pageNumber="124">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336ABBB2A5E6B63A663B19C2A6617" blockId="7.[188,1398,494,660]" box="[234,298,494,520]" pageId="7" pageNumber="124">
<emphasis id="B978EAB9BB2A5E6B63A663B19C2A6617" bold="true" box="[234,298,494,520]" pageId="7" pageNumber="124">Male</emphasis>
</paragraph>
<paragraph id="8BB336ABBB2A5E6B63F1604E9C86668B" blockId="7.[188,1398,494,660]" pageId="7" pageNumber="124">
HEAD. Frons and vertex with appressed scales; narrow, elongated scales behind compound eyes. Antenna filiform, about 0.7 times as long as forewing, basal five segments of flagellum slightly broadened (
<figureCitation id="13372A2EBB2A5E6B638A60089C24666E" box="[198,292,599,625]" captionStart="Fig" captionStartId="7.[189,232,1928,1954]" captionTargetBox="[189,1398,757,1889]" captionTargetId="figure-163@7.[189,1398,757,1889]" captionTargetPageId="7" captionText="Fig. 3. Helioandesia tarregai gen. et sp. nov., ♂, holotype (IDEA-LEPI-2020-015). A. Dorsal view. B. Head, lateral. C. Basal flagellomeres of right antenna. D. Three bulging metallic spots on the basal half of forewing. E. Hair pencil from base of subcosta, ventral. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452227" httpUri="https://zenodo.org/record/4452227/files/figure.png" pageId="7" pageNumber="124">Fig. 3C</figureCitation>
) due to narrow superimposed scales on dorsal side. Labial palpus (
<figureCitation id="13372A2EBB2A5E6B6725600899C7666E" box="[1129,1223,599,625]" captionStart="Fig" captionStartId="7.[189,232,1928,1954]" captionTargetBox="[189,1398,757,1889]" captionTargetId="figure-163@7.[189,1398,757,1889]" captionTargetPageId="7" captionText="Fig. 3. Helioandesia tarregai gen. et sp. nov., ♂, holotype (IDEA-LEPI-2020-015). A. Dorsal view. B. Head, lateral. C. Basal flagellomeres of right antenna. D. Three bulging metallic spots on the basal half of forewing. E. Hair pencil from base of subcosta, ventral. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452227" httpUri="https://zenodo.org/record/4452227/files/figure.png" pageId="7" pageNumber="124">Fig. 3B</figureCitation>
) porrect, with appressed scales.
</paragraph>
<caption id="DF736623BB2A5E6B63F165D7996E63F6" ID-DOI="http://doi.org/10.5281/zenodo.4452227" ID-Zenodo-Dep="4452227" httpUri="https://zenodo.org/record/4452227/files/figure.png" pageId="7" pageNumber="124" startId="7.[189,232,1928,1954]" targetBox="[189,1398,757,1889]" targetPageId="7">
<paragraph id="8BB336ABBB2A5E6B63F165D7996E63F6" blockId="7.[189,1399,1928,2025]" pageId="7" pageNumber="124">
<emphasis id="B978EAB9BB2A5E6B63F165D79C0D63BD" bold="true" box="[189,269,1928,1954]" pageId="7" pageNumber="124">Fig. 3.</emphasis>
<taxonomicName id="4C0C4D28BB2A5E6B625465D79F1E63BD" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[280,542,1928,1954]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB2A5E6B625465D79F1E63BD" box="[280,542,1928,1954]" italics="true" pageId="7" pageNumber="124">Helioandesia tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB2A5E6B616465D79FE463BD" box="[552,740,1928,1954]" pageId="7" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, ♂, holotype (IDEA-LEPI-2020-015).
<emphasis id="B978EAB9BB2A5E6B67F565D799CE63BD" bold="true" box="[1209,1230,1928,1954]" pageId="7" pageNumber="124">A</emphasis>
. Dorsal view.
<emphasis id="B978EAB9BB2A5E6B63F165F49DD163DA" bold="true" box="[189,209,1963,1989]" pageId="7" pageNumber="124">B</emphasis>
. Head, lateral.
<emphasis id="B978EAB9BB2A5E6B62C465F49C9C63DA" bold="true" box="[392,412,1963,1989]" pageId="7" pageNumber="124">C</emphasis>
. Basal flagellomeres of right antenna.
<emphasis id="B978EAB9BB2A5E6B602565F49E7E63DA" bold="true" box="[873,894,1963,1989]" pageId="7" pageNumber="124">D</emphasis>
. Three bulging metallic spots on the basal half of forewing.
<emphasis id="B978EAB9BB2A5E6B62C665919C9E63F7" bold="true" box="[394,414,1998,2024]" pageId="7" pageNumber="124">E</emphasis>
. Hair pencil from base of subcosta, ventral. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BB336ABBB255E6463F163569E7665CC" blockId="8.[188,1399,265,467]" pageId="8" pageNumber="125">
THORAX. Covered with appressed scales. Tibial spurs 0-2-4; metatibia with inner spur of proximal pair about twice as long as outer one. Forewing lanceolate (
<figureCitation id="13372A2EBB255E64602C63739EAA6559" box="[864,938,300,326]" captionStart="Fig" captionStartId="8.[188,231,1963,1989]" captionTargetBox="[190,1397,1272,1927]" captionTargetId="figure-364@8.[190,1397,1272,1927]" captionTargetPageId="8" captionText="Fig. 4. Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024), wing venation showing frenulum with two bristles of unequal length. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452229" httpUri="https://zenodo.org/record/4452229/files/figure.png" pageId="8" pageNumber="125">Fig. 4</figureCitation>
), with strongly bulging metallic spots (
<figureCitation id="13372A2EBB255E64638A63109C4C6575" box="[198,332,335,362]" captionStart="Fig" captionStartId="7.[189,232,1928,1954]" captionTargetBox="[189,1398,757,1889]" captionTargetId="figure-163@7.[189,1398,757,1889]" captionTargetPageId="7" captionText="Fig. 3. Helioandesia tarregai gen. et sp. nov., ♂, holotype (IDEA-LEPI-2020-015). A. Dorsal view. B. Head, lateral. C. Basal flagellomeres of right antenna. D. Three bulging metallic spots on the basal half of forewing. E. Hair pencil from base of subcosta, ventral. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452227" httpUri="https://zenodo.org/record/4452227/files/figure.png" pageId="8" pageNumber="125">Fig. 3A, D</figureCitation>
); hair pencil (
<figureCitation id="13372A2EBB255E6462BF63109F4D6576" box="[499,589,335,361]" captionStart="Fig" captionStartId="7.[189,232,1928,1954]" captionTargetBox="[189,1398,757,1889]" captionTargetId="figure-163@7.[189,1398,757,1889]" captionTargetPageId="7" captionText="Fig. 3. Helioandesia tarregai gen. et sp. nov., ♂, holotype (IDEA-LEPI-2020-015). A. Dorsal view. B. Head, lateral. C. Basal flagellomeres of right antenna. D. Three bulging metallic spots on the basal half of forewing. E. Hair pencil from base of subcosta, ventral. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452227" httpUri="https://zenodo.org/record/4452227/files/figure.png" pageId="8" pageNumber="125">Fig. 3E</figureCitation>
) from base of subcosta ventrally; 12 veins; five R, all separate; M1 and M2 separate; CuA1 and CuA2 separate, proximal part of latter lost; CuP present as tubular vein near margin; 1A+2A present. Hindwing (
<figureCitation id="13372A2EBB255E64612663C99FB365AF" box="[618,691,406,432]" captionStart="Fig" captionStartId="8.[188,231,1963,1989]" captionTargetBox="[190,1397,1272,1927]" captionTargetId="figure-364@8.[190,1397,1272,1927]" captionTargetPageId="8" captionText="Fig. 4. Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024), wing venation showing frenulum with two bristles of unequal length. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452229" httpUri="https://zenodo.org/record/4452229/files/figure.png" pageId="8" pageNumber="125">Fig. 4</figureCitation>
) lanceolate, six veins; Sc+R1 and Rs separate; M1 and M2 stalked; CuA1 and CuA2 stalked; frenulum a single bristle.
</paragraph>
<paragraph id="8BB336ABBB255E6463F163A49E5166BE" blockId="8.[189,1399,506,674]" pageId="8" pageNumber="125">
ABDOMEN. Tergum VIII (
<figureCitation id="13372A2EBB255E6462AB63A59F44660A" box="[487,580,506,533]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[327,1261,319,1818]" captionTargetId="figure-11@9.[326,1261,319,1818]" captionTargetPageId="9" captionText="Fig. 5. Male genitalia and eighth segment of Helioandesia tarregai gen. et sp. nov., ♂, paratype (IDEA- LEPI-2020-021). A. Male genitalia, ventral view, phallus removed. B. Phallus, lateral view. C. Tergum and pleural lobes of eighth segment. D. Socius, closed arrow in A. E. Small spine-like cornuti on the vesica, open arrow in A. F. Medial part of the cornutus. G. Coremata between pleural lobes of segment VIII and genitalia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452231" httpUri="https://zenodo.org/record/4452231/files/figure.png" pageId="8" pageNumber="125">Fig. 5C</figureCitation>
) triangular; anterior margin slightly concave, antero-lateral vertices as small processes slightly extending anteriorly beneath tergum VII; lateral margin widely convex; posterior margin about ¼ of anterior margin, with wide rounded notch in middle. Pleural lobe (
<figureCitation id="13372A2EBB255E646658601E986D6643" box="[1300,1389,577,604]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[327,1261,319,1818]" captionTargetId="figure-11@9.[326,1261,319,1818]" captionTargetPageId="9" captionText="Fig. 5. Male genitalia and eighth segment of Helioandesia tarregai gen. et sp. nov., ♂, paratype (IDEA- LEPI-2020-021). A. Male genitalia, ventral view, phallus removed. B. Phallus, lateral view. C. Tergum and pleural lobes of eighth segment. D. Socius, closed arrow in A. E. Small spine-like cornuti on the vesica, open arrow in A. F. Medial part of the cornutus. G. Coremata between pleural lobes of segment VIII and genitalia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452231" httpUri="https://zenodo.org/record/4452231/files/figure.png" pageId="8" pageNumber="125">Fig. 5C</figureCitation>
) triangular, with slightly sclerotized stripe near posterior margin. Sternum VIII not differentiated. Coremata (
<figureCitation id="13372A2EBB255E64627360D89C9B66BD" box="[319,411,647,674]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[327,1261,319,1818]" captionTargetId="figure-11@9.[326,1261,319,1818]" captionTargetPageId="9" captionText="Fig. 5. Male genitalia and eighth segment of Helioandesia tarregai gen. et sp. nov., ♂, paratype (IDEA- LEPI-2020-021). A. Male genitalia, ventral view, phallus removed. B. Phallus, lateral view. C. Tergum and pleural lobes of eighth segment. D. Socius, closed arrow in A. E. Small spine-like cornuti on the vesica, open arrow in A. F. Medial part of the cornutus. G. Coremata between pleural lobes of segment VIII and genitalia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452231" httpUri="https://zenodo.org/record/4452231/files/figure.png" pageId="8" pageNumber="125">Fig. 5G</figureCitation>
) between pleural lobes and genitalia.
</paragraph>
<paragraph id="8BB336ABBB255E6463F160919CF767C6" blockId="8.[188,1401,712,986]" pageId="8" pageNumber="125">
GENITALIA (
<figureCitation id="13372A2EBB255E64627060969C8666FC" box="[316,390,712,739]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[327,1261,319,1818]" captionTargetId="figure-11@9.[326,1261,319,1818]" captionTargetPageId="9" captionText="Fig. 5. Male genitalia and eighth segment of Helioandesia tarregai gen. et sp. nov., ♂, paratype (IDEA- LEPI-2020-021). A. Male genitalia, ventral view, phallus removed. B. Phallus, lateral view. C. Tergum and pleural lobes of eighth segment. D. Socius, closed arrow in A. E. Small spine-like cornuti on the vesica, open arrow in A. F. Medial part of the cornutus. G. Coremata between pleural lobes of segment VIII and genitalia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452231" httpUri="https://zenodo.org/record/4452231/files/figure.png" pageId="8" pageNumber="125">Fig. 5</figureCitation>
). Tegumen posteriorly expanded, somewhat conical, narrowing posteriorly. Gnathos absent. Socius (
<figureCitation id="13372A2EBB255E64623360B39CDE6718" box="[383,478,748,775]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[327,1261,319,1818]" captionTargetId="figure-11@9.[326,1261,319,1818]" captionTargetPageId="9" captionText="Fig. 5. Male genitalia and eighth segment of Helioandesia tarregai gen. et sp. nov., ♂, paratype (IDEA- LEPI-2020-021). A. Male genitalia, ventral view, phallus removed. B. Phallus, lateral view. C. Tergum and pleural lobes of eighth segment. D. Socius, closed arrow in A. E. Small spine-like cornuti on the vesica, open arrow in A. F. Medial part of the cornutus. G. Coremata between pleural lobes of segment VIII and genitalia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452231" httpUri="https://zenodo.org/record/4452231/files/figure.png" pageId="8" pageNumber="125">Fig. 5D</figureCitation>
) narrow, conical, with hair-like setae, apex coincides with that of tegumen. Saccus narrow, rod-shaped, about twice as long as tegumen, slightly broadened near base, distal end slightly swollen. Valva simple, with a few hair-like setae, costal margin broadly concave, ventral margin broadly convex, apex rounded. Phallus cylindrical, elongated, slightly longer than tegumen + saccus, slightly curved ventrally; vesica with prominent, long spine-like cornutus (
<figureCitation id="13372A2EBB255E64677A612699B1678C" box="[1078,1201,888,915]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[327,1261,319,1818]" captionTargetId="figure-11@9.[326,1261,319,1818]" captionTargetPageId="9" captionText="Fig. 5. Male genitalia and eighth segment of Helioandesia tarregai gen. et sp. nov., ♂, paratype (IDEA- LEPI-2020-021). A. Male genitalia, ventral view, phallus removed. B. Phallus, lateral view. C. Tergum and pleural lobes of eighth segment. D. Socius, closed arrow in A. E. Small spine-like cornuti on the vesica, open arrow in A. F. Medial part of the cornutus. G. Coremata between pleural lobes of segment VIII and genitalia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452231" httpUri="https://zenodo.org/record/4452231/files/figure.png" pageId="8" pageNumber="125">Fig. 5B, F</figureCitation>
) slightly smaller than half of phallus, and group of small spine-like cornuti (
<figureCitation id="13372A2EBB255E6460CA61C39EE267A8" box="[902,994,924,951]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[327,1261,319,1818]" captionTargetId="figure-11@9.[326,1261,319,1818]" captionTargetPageId="9" captionText="Fig. 5. Male genitalia and eighth segment of Helioandesia tarregai gen. et sp. nov., ♂, paratype (IDEA- LEPI-2020-021). A. Male genitalia, ventral view, phallus removed. B. Phallus, lateral view. C. Tergum and pleural lobes of eighth segment. D. Socius, closed arrow in A. E. Small spine-like cornuti on the vesica, open arrow in A. F. Medial part of the cornutus. G. Coremata between pleural lobes of segment VIII and genitalia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452231" httpUri="https://zenodo.org/record/4452231/files/figure.png" pageId="8" pageNumber="125">Fig. 5E</figureCitation>
); insertion of ductus ejaculatorius near basal third of phallus.
</paragraph>
<paragraph id="8BB336ABBB255E6463A6665E9C456004" blockId="8.[187,1399,1025,1121]" box="[234,325,1025,1051]" pageId="8" pageNumber="125">
<emphasis id="B978EAB9BB255E6463A6665E9C456004" bold="true" box="[234,325,1025,1051]" pageId="8" pageNumber="125">Female</emphasis>
</paragraph>
<paragraph id="8BB336ABBB255E6463F1667B9F78607E" blockId="8.[187,1399,1025,1121]" pageId="8" pageNumber="125">
Mainly similar to male, but hindwing with two asymmetrical bristles on frenulum (
<figureCitation id="13372A2EBB255E64673D667B99B76021" box="[1137,1207,1060,1086]" captionStart="Fig" captionStartId="8.[188,231,1963,1989]" captionTargetBox="[190,1397,1272,1927]" captionTargetId="figure-364@8.[190,1397,1272,1927]" captionTargetPageId="8" captionText="Fig. 4. Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024), wing venation showing frenulum with two bristles of unequal length. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452229" httpUri="https://zenodo.org/record/4452229/files/figure.png" pageId="8" pageNumber="125">Fig. 4</figureCitation>
), and abdominal segment VIII not modified as in male.
</paragraph>
<paragraph id="8BB336ABBB255E6763F166D19F876559" blockId="8.[189,1399,1160,1222]" lastBlockId="11.[189,1398,265,327]" lastPageId="11" lastPageNumber="128" pageId="8" pageNumber="125">
GENITALIA (
<figureCitation id="13372A2EBB255E64627666D79C8260BC" box="[314,386,1160,1187]" captionStart="Fig" captionStartId="10.[191,234,1927,1953]" captionTargetBox="[189,1398,559,1861]" captionTargetId="figure-92@10.[189,1398,559,1861]" captionTargetPageId="10" captionText="Fig. 6. Female genitalia of Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024) A. Female genitalia, ventral view. B. Papillae analis. C. Ornamentation of the proximal part of the signum. D. Ornamentation of the distal part of the signum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452233" httpUri="https://zenodo.org/record/4452233/files/figure.png" pageId="8" pageNumber="125">Fig. 6</figureCitation>
). Papillae anales (
<figureCitation id="13372A2EBB255E64611B66D79FB160BC" box="[599,689,1160,1187]" captionStart="Fig" captionStartId="10.[191,234,1927,1953]" captionTargetBox="[189,1398,559,1861]" captionTargetId="figure-92@10.[189,1398,559,1861]" captionTargetPageId="10" captionText="Fig. 6. Female genitalia of Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024) A. Female genitalia, ventral view. B. Papillae analis. C. Ornamentation of the proximal part of the signum. D. Ornamentation of the distal part of the signum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452233" httpUri="https://zenodo.org/record/4452233/files/figure.png" pageId="8" pageNumber="125">Fig. 6B</figureCitation>
) narrow, elongated, slightly sclerotized, with hair-like setae. Posterior apophysis rod-like, about twice as long as papillae anales. Anterior apophysis rod-like, slightly shorter than posterior apophysis. Transverse bridge as rectangular transversal band, anterior vertices fused with posterior end of anterior apophyses. Ostium bursae membranous. Ductus bursae narrow, mainly membranous; proximal fourth widened, with distal half sclerotized; slightly widened and sclerotized near corpus bursae. Ductus seminalis arises on distal fourth of ductus bursae. Corpus bursae mainly membranous, lenticular; signum on left side, narrow, elongated, minutely sculptured, proximal (
<figureCitation id="13372A2EBB275E66638B63E69C2365CB" box="[199,291,441,468]" captionStart="Fig" captionStartId="10.[191,234,1927,1953]" captionTargetBox="[189,1398,559,1861]" captionTargetId="figure-92@10.[189,1398,559,1861]" captionTargetPageId="10" captionText="Fig. 6. Female genitalia of Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024) A. Female genitalia, ventral view. B. Papillae analis. C. Ornamentation of the proximal part of the signum. D. Ornamentation of the distal part of the signum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452233" httpUri="https://zenodo.org/record/4452233/files/figure.png" pageId="10" pageNumber="127">Fig. 6C</figureCitation>
) portion coarser than distal (
<figureCitation id="13372A2EBB275E66613163E69FDA65CB" box="[637,730,441,468]" captionStart="Fig" captionStartId="10.[191,234,1927,1953]" captionTargetBox="[189,1398,559,1861]" captionTargetId="figure-92@10.[189,1398,559,1861]" captionTargetPageId="10" captionText="Fig. 6. Female genitalia of Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024) A. Female genitalia, ventral view. B. Papillae analis. C. Ornamentation of the proximal part of the signum. D. Ornamentation of the distal part of the signum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452233" httpUri="https://zenodo.org/record/4452233/files/figure.png" pageId="10" pageNumber="127">Fig. 6D</figureCitation>
) portion, slightly invaginate and broadened subapically, length about ¾ that of corpus bursae. Appendix bursae spherical, membranous, on the middle of corpus bursae, on the opposite side of signum.
</paragraph>
</subSubSection>
</treatment>
</document>
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