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<document id="6505AC365C244FC32533A2FF648D2814" ID-CLB-Dataset="27972" ID-DOI="10.11646/zootaxa.4521.1.3" ID-GBIF-Dataset="496b64db-ed0b-43a2-bb95-8e75fcf30a7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2609668" ID-ZooBank="12685FBF-7507-48D0-9753-D747B6ECC778" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553623393857" checkinUser="plazi" docAuthor="Cruz-Gómez, Christopher &amp; Bastida-Zavala, J. Rolando" docDate="2018" docId="039187C9361DFFA26199F906D26B9385" docLanguage="en" docName="zootaxa.4521.1.3.pdf" docOrigin="Zootaxa 4521 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bhawania goodei Webster 1884" docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="FFA8FFB13619FFA7610EFFECD75E9618" masterDocTitle="Chrysopetalids (Phyllodocida: Chrysopetalidae) from the southern Mexican Pacific, including a new species" masterLastPageNumber="88" masterPageNumber="61" pageNumber="65" updateTime="1698713678056" updateUser="ExternalLinkService">
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<mods:title id="9A566E22BDE02E94A8E21765053589AA">Chrysopetalids (Phyllodocida: Chrysopetalidae) from the southern Mexican Pacific, including a new species</mods:title>
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<mods:namePart id="2307F51FD5024624D6861B418314FE1F">Cruz-Gómez, Christopher</mods:namePart>
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<treatment id="039187C9361DFFA26199F906D26B9385" ID-DOI="http://doi.org/10.5281/zenodo.5950273" ID-GBIF-Taxon="154498659" ID-Zenodo-Dep="5950273" LSID="urn:lsid:plazi:treatment:039187C9361DFFA26199F906D26B9385" httpUri="http://treatment.plazi.org/id/039187C9361DFFA26199F906D26B9385" lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65">
<subSubSection id="C3226554361DFFA36199F906D518911D" box="[151,582,1770,1797]" pageId="4" pageNumber="65" type="nomenclature">
<paragraph id="8B8736DF361DFFA36199F906D518911D" blockId="4.[151,582,1770,1830]" box="[151,582,1770,1797]" pageId="4" pageNumber="65">
<heading id="D0CF81B3361DFFA36199F906D518911D" bold="true" box="[151,582,1770,1797]" fontSize="11" level="1" pageId="4" pageNumber="65" reason="1">
<taxonomicName id="4C384D5C361DFFA36199F906D518911D" ID-CoL="LQ6J" authority="Webster, 1884" authorityName="Webster" authorityYear="1884" box="[151,582,1770,1797]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="65" phylum="Annelida" rank="species" species="goodei">
<emphasis id="B94CEACD361DFFA36199F906D518911D" bold="true" box="[151,582,1770,1797]" pageId="4" pageNumber="65">
<emphasis id="B94CEACD361DFFA36199F906D64D911C" bold="true" box="[151,275,1770,1796]" italics="true" pageId="4" pageNumber="65">Bhawania</emphasis>
cf.
<emphasis id="B94CEACD361DFFA36031F906D6D1911C" bold="true" box="[319,399,1770,1796]" italics="true" pageId="4" pageNumber="65">goodei</emphasis>
<bibRefCitation id="EFA94B2E361DFFA36098F907D518911D" author="Webster, H. E." box="[406,582,1771,1797]" pageId="4" pageNumber="65" pagination="305 - 327" refId="ref15636" refString="Webster, H. E. (1884) Annelida from Bermuda, collected by G. Brown Goode. Bulletin United States National Museum, 25, 305 - 327. https: // doi. org / 10.5479 / si. 03629236.25.305" type="journal article" year="1884">Webster, 1884</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3226554361DFFA36199F8E1D4919173" pageId="4" pageNumber="65" type="description">
<paragraph id="8B8736DF361DFFA36199F8E1D683913E" blockId="4.[151,582,1770,1830]" box="[151,477,1805,1830]" pageId="4" pageNumber="65">
(
<figureCitation id="13032A5A361DFFA36190F8E1D659913E" box="[158,263,1805,1830]" captionStart="FIGURE 3" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[178,1409,200,1931]" captionTargetId="figure@8.[170,1423,189,1942]" captionTargetPageId="8" captionText="FIGURE 3. Bhawania cf. goodei. A, parapodium middle segment; BE, neurochaetal types; F, lateral paleae; G, subunit 1 palea; H, meddle main palea; IK, median paleae group. Scale bars: 30 µm (BCDE), 100 µm (FGHIJK), 200 µm (A)." httpUri="https://zenodo.org/record/2609674/files/figure.png" pageId="4" pageNumber="65">Figures 3</figureCitation>
AK, 8, 14E, 15B)
</paragraph>
<paragraph id="8B8736DF361DFFA36199F8BAD4919173" blockId="4.[151,975,1877,1899]" box="[151,975,1877,1899]" pageId="4" pageNumber="65">
<materialsCitation id="3B503C82361DFFA36199F8BAD4959173" ID-GBIF-Occurrence="2028520809" box="[151,971,1877,1899]" country="Bermuda" location="Western Atlantic" pageId="4" pageNumber="65" specimenCount="1" typeStatus="holotype">
<typeStatus id="5483887D361DFFA36199F8BAD7979173" box="[151,201,1878,1899]" pageId="4" pageNumber="65">Type</typeStatus>
locality of the nominal species:
<collectingCountry id="F32F764F361DFFA3631CF8B9D52F9173" box="[530,625,1877,1899]" name="Bermuda" pageId="4" pageNumber="65">Bermuda</collectingCountry>
,
<location id="8EE76004361DFFA36375F8BAD4789173" LSID="urn:lsid:plazi:treatment:039187C9361DFFA26199F906D26B9385:8EE76004361DFFA36375F8BAD4789173" box="[635,806,1877,1899]" country="Bermuda" name="Western Atlantic" pageId="4" pageNumber="65">Western Atlantic</location>
(
<bibRefCitation id="EFA94B2E361DFFA3623CF8B9D49A9173" author="Webster, H. E." box="[818,964,1877,1899]" pageId="4" pageNumber="65" pagination="305 - 327" refId="ref15636" refString="Webster, H. E. (1884) Annelida from Bermuda, collected by G. Brown Goode. Bulletin United States National Museum, 25, 305 - 327. https: // doi. org / 10.5479 / si. 03629236.25.305" type="journal article" year="1884">Webster 1884</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226554361DFFA36199F874D42891CD" pageId="4" pageNumber="65" type="materials_examined">
<paragraph id="8B8736DF361DFFA36199F874D42891CD" blockId="4.[151,1436,1944,2005]" pageId="4" pageNumber="65">
<emphasis id="B94CEACD361DFFA36199F874D6B591A9" bold="true" box="[151,491,1944,1969]" pageId="4" pageNumber="65">
Material examined.
<materialsCitation id="3B503C82361DFFA36089F874D6B591A9" ID-GBIF-Occurrence="2028520800" box="[391,491,1944,1969]" collectingDate="2008-04-03" collectorName="J. Jarquin" latitude="15.966389" location="Chacahua Lagoon" longLatPrecision="21" longitude="-97.67889" pageId="4" pageNumber="65" specimenCount="1" stateProvince="Oaxaca">
<collectingRegion id="49FCF83D361DFFA36089F874D6BB91A9" box="[391,485,1944,1969]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="65">Oaxaca</collectingRegion>
:
</materialsCitation>
</emphasis>
UMAR-Poly 855 (mouth of
<location id="8EE76004361DFFA3623EF874D4A391A9" LSID="urn:lsid:plazi:treatment:039187C9361DFFA26199F906D26B9385:8EE76004361DFFA3623EF874D4A391A9" box="[816,1021,1944,1969]" latitude="15.966389" longLatPrecision="21" longitude="-97.67889" name="Chacahua Lagoon" pageId="4" pageNumber="65" stateProvince="Oaxaca">Chacahua Lagoon</location>
,
<geoCoordinate id="EE0C5018361DFFA36506F874D3C891A9" box="[1032,1174,1944,1969]" direction="north" orientation="latitude" pageId="4" pageNumber="65" precision="15" value="15.966389">15°5759N</geoCoordinate>
,
<geoCoordinate id="EE0C5018361DFFA365AFF874D26D91A8" box="[1185,1331,1944,1968]" direction="west" orientation="longitude" pageId="4" pageNumber="65" precision="15" value="-97.67889">97°4044W</geoCoordinate>
, on tubes of sabellariids, 0.5 m,
<date id="FF86101F361DFFA3609EF850D57791CC" box="[400,553,1980,2005]" pageId="4" pageNumber="65" value="2008-04-03">
<collectingDate id="EFC2E9F7361DFFA3609EF850D57791CC" box="[400,553,1980,2005]" pageId="4" pageNumber="65" value="2008-04-03">April 3, 2008</collectingDate>
</date>
, coll.
<collectorName id="26CD5309361DFFA36363F851D58391CC" box="[621,733,1980,2004]" pageId="4" pageNumber="65">J. Jarquín</collectorName>
),
<specimenCount id="9D3EFD56361DFFA363E1F850D42F91CD" box="[751,881,1980,2005]" pageId="4" pageNumber="65" type="generic">1 specimen</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226554361CFFA261C9FF7BD5B89545" pageId="5" pageNumber="66" type="description">
<paragraph id="8B8736DF361CFFA261C9FF7BD6239790" blockId="5.[151,1437,151,1437]" pageId="5" pageNumber="66">
<emphasis id="B94CEACD361CFFA261C9FF7BD60796A8" bold="true" box="[199,345,151,176]" pageId="5" pageNumber="66">Description.</emphasis>
Based on one incomplete specimen, 6.8 mm long, 1.2 mm wide, with only 48 anterior segments. Body rectangular and wide, first segments slightly tapered. Body pale yellow to brown. Paleae fans yellow to golden, imbricated over dorsum, covering it totally. Prostomium rounded, completely retracted among the first five anterior segments. Two lateral antennae; a median antenna inserted anterior to the prostomium. Two pairs of reddish eyes; second pair slightly larger than first pair. Nuchal fold not observed. Two cylindrical palps, only visible in ventral view. Mouth fold semicircular, inserted between second and third segment. Eversible proboscis, stylets not observed.
</paragraph>
<paragraph id="8B8736DF361CFFA261C9FE78D6259520" blockId="5.[151,1437,151,1437]" pageId="5" pageNumber="66">
Parapodium from the 15th segment. Notochaetae separated in three main groups and one subgroup (
<figureCitation id="13032A5A361CFFA2643AFE78D2D197B4" box="[1332,1423,404,429]" captionStart="FIGURE 3" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[178,1409,200,1931]" captionTargetId="figure@8.[170,1423,189,1942]" captionTargetPageId="8" captionText="FIGURE 3. Bhawania cf. goodei. A, parapodium middle segment; BE, neurochaetal types; F, lateral paleae; G, subunit 1 palea; H, meddle main palea; IK, median paleae group. Scale bars: 30 µm (BCDE), 100 µm (FGHIJK), 200 µm (A)." httpUri="https://zenodo.org/record/2609674/files/figure.png" pageId="5" pageNumber="66">Fig. 3A</figureCitation>
). Lateral group inserted over the notoaciculum; comprising 810 slender and feather-like paleae with 57 ribs and 0 1 raised rib (
<figureCitation id="13032A5A361CFFA26027FE30D6DE97EC" box="[297,384,476,501]" captionStart="FIGURE 3" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[178,1409,200,1931]" captionTargetId="figure@8.[170,1423,189,1942]" captionTargetPageId="8" captionText="FIGURE 3. Bhawania cf. goodei. A, parapodium middle segment; BE, neurochaetal types; F, lateral paleae; G, subunit 1 palea; H, meddle main palea; IK, median paleae group. Scale bars: 30 µm (BCDE), 100 µm (FGHIJK), 200 µm (A)." httpUri="https://zenodo.org/record/2609674/files/figure.png" pageId="5" pageNumber="66">Fig. 3F</figureCitation>
). As subunit 1, one symmetrical palea with eight ribs (
<figureCitation id="13032A5A361CFFA262FAFE30D30E97EC" box="[1012,1104,476,501]" captionStart="FIGURE 3" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[178,1409,200,1931]" captionTargetId="figure@8.[170,1423,189,1942]" captionTargetPageId="8" captionText="FIGURE 3. Bhawania cf. goodei. A, parapodium middle segment; BE, neurochaetal types; F, lateral paleae; G, subunit 1 palea; H, meddle main palea; IK, median paleae group. Scale bars: 30 µm (BCDE), 100 µm (FGHIJK), 200 µm (A)." httpUri="https://zenodo.org/record/2609674/files/figure.png" pageId="5" pageNumber="66">Fig. 3G</figureCitation>
). Main group, comprising 15 symmetrical paleae with 1315 ribs and three raised ribs (
<figureCitation id="13032A5A361CFFA26229FE13D4DC9400" box="[807,898,511,536]" captionStart="FIGURE 3" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[178,1409,200,1931]" captionTargetId="figure@8.[170,1423,189,1942]" captionTargetPageId="8" captionText="FIGURE 3. Bhawania cf. goodei. A, parapodium middle segment; BE, neurochaetal types; F, lateral paleae; G, subunit 1 palea; H, meddle main palea; IK, median paleae group. Scale bars: 30 µm (BCDE), 100 µm (FGHIJK), 200 µm (A)." httpUri="https://zenodo.org/record/2609674/files/figure.png" pageId="5" pageNumber="66">Fig. 3H</figureCitation>
). Median group, comprising three, symmetrical paleae; distal paleae with 18 ribs and three raised ribs, next distal palea smaller than main paleae with 12 ribs and five raised ribs; the most proximal and smallest with eight ribs, sometimes with 12 raised ribs (
<figureCitation id="13032A5A361CFFA26417FDABD2019478" box="[1305,1375,583,608]" captionStart="FIGURE 3" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[178,1409,200,1931]" captionTargetId="figure@8.[170,1423,189,1942]" captionTargetPageId="8" captionText="FIGURE 3. Bhawania cf. goodei. A, parapodium middle segment; BE, neurochaetal types; F, lateral paleae; G, subunit 1 palea; H, meddle main palea; IK, median paleae group. Scale bars: 30 µm (BCDE), 100 µm (FGHIJK), 200 µm (A)." httpUri="https://zenodo.org/record/2609674/files/figure.png" pageId="5" pageNumber="66">Fig. 3</figureCitation>
IK). Neuropodium conical and larger than notopodium. Neurochaetae composed of four groups (
<figureCitation id="13032A5A361CFFA26407FD80D207949C" box="[1289,1369,620,645]" captionStart="FIGURE 3" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[178,1409,200,1931]" captionTargetId="figure@8.[170,1423,189,1942]" captionTargetPageId="8" captionText="FIGURE 3. Bhawania cf. goodei. A, parapodium middle segment; BE, neurochaetal types; F, lateral paleae; G, subunit 1 palea; H, meddle main palea; IK, median paleae group. Scale bars: 30 µm (BCDE), 100 µm (FGHIJK), 200 µm (A)." httpUri="https://zenodo.org/record/2609674/files/figure.png" pageId="5" pageNumber="66">Figs 3</figureCitation>
BE). Neurochaetiger with 13 superior spinigers with a long and curved slim blade, with a faintly serration basally. Midsuperior group 45 falcigers with medium length blades (1/3 of spiniger blade length) and serration basally. Midinferior group with 57 falcigers with medium blades (1/3 of spiniger blade length), serration basally. Inferior group of 24 falcigers with short and smooth blades (2/7 of spiniger blade length) with fine serration basally. Pygidium unknown.
</paragraph>
<paragraph id="8B8736DF361CFFA261C9FCA8D5B89545" blockId="5.[151,1437,151,1437]" box="[199,742,836,861]" pageId="5" pageNumber="66">
<emphasis id="B94CEACD361CFFA261C9FCA8D6749545" bold="true" box="[199,298,836,861]" pageId="5" pageNumber="66">Habitat.</emphasis>
Intertidal (0.5 m). On sabellariid tubes.
</paragraph>
</subSubSection>
<subSubSection id="C3226554361CFFA261C9FC8BD2779598" box="[199,1321,871,896]" pageId="5" pageNumber="66" type="distribution">
<paragraph id="8B8736DF361CFFA261C9FC8BD2779598" blockId="5.[151,1437,151,1437]" box="[199,1321,871,896]" pageId="5" pageNumber="66">
<emphasis id="B94CEACD361CFFA261C9FC8BD63E9598" bold="true" box="[199,352,871,896]" pageId="5" pageNumber="66">Distribution.</emphasis>
Southern Mexican Pacific, only known from Chacahua Bay,
<collectingRegion id="49FCF83D361CFFA2651EFC84D3399598" box="[1040,1127,872,896]" country="Mexico" name="Oaxaca" pageId="5" pageNumber="66">Oaxaca</collectingRegion>
,
<collectingCountry id="F32F764F361CFFA2657CFC8BD3999598" box="[1138,1223,871,896]" name="Mexico" pageId="5" pageNumber="66">Mexico</collectingCountry>
(
<figureCitation id="13032A5A361CFFA265D9FC8BD2459598" box="[1239,1307,871,896]" captionStart="FIGURE 8" captionStartId="14.[151,250,1055,1077]" captionTargetBox="[286,1302,197,1033]" captionTargetId="figure@14.[280,1310,193,1038]" captionTargetPageId="14" captionText="FIGURE 8. Historical and new records of Arichlidon watsonae n. sp., Bhawania cf. goodei, Chrysopetalum elegantoides, C. maculata and C. occidentale in the Tropical Eastern Pacific. Empty symbols indicate previous records." httpUri="https://zenodo.org/record/2609684/files/figure.png" pageId="5" pageNumber="66">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226554361CFFA261C9FC60D26B9385" pageId="5" pageNumber="66" type="discussion">
<paragraph id="8B8736DF361CFFA261C9FC60D45592F0" blockId="5.[151,1437,151,1437]" pageId="5" pageNumber="66">
<emphasis id="B94CEACD361CFFA261C9FC60D66595BD" bold="true" box="[199,315,908,933]" pageId="5" pageNumber="66">Remarks.</emphasis>
The genus
<taxonomicName id="4C384D5C361CFFA260CEFC60D56C95BC" authorityName="Schmarda" authorityYear="1861" box="[448,562,908,932]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="genus">
<emphasis id="B94CEACD361CFFA260CEFC60D56C95BC" box="[448,562,908,932]" italics="true" pageId="5" pageNumber="66">Bhawania</emphasis>
</taxonomicName>
is poorly known in the Pacific coast of
<collectingCountry id="F32F764F361CFFA262F6FC60D31395BD" box="[1016,1101,908,933]" name="Mexico" pageId="5" pageNumber="66">Mexico</collectingCountry>
with only two species of this genus recorded from the region:
<taxonomicName id="4C384D5C361CFFA2630CFC5DD53395D0" authorityName="Webster" authorityYear="1884" box="[514,621,944,968]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="goodei">
<emphasis id="B94CEACD361CFFA2630CFC5DD53395D0" box="[514,621,944,968]" italics="true" pageId="5" pageNumber="66">B. goodei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C384D5C361CFFA263ADFC5DD4F295D0" authority="(Gravier 1908)" baseAuthorityName="Gravier" baseAuthorityYear="1908" box="[675,940,943,968]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="riveti">
<emphasis id="B94CEACD361CFFA263ADFC5DD5A295DF" box="[675,764,944,968]" italics="true" pageId="5" pageNumber="66">B. riveti</emphasis>
(
<bibRefCitation id="EFA94B2E361CFFA26204FC43D4FA95D0" author="Gravier, M. C." box="[778,932,943,968]" pageId="5" pageNumber="66" pagination="40 - 44" refId="ref13655" refString="Gravier, M. C. (1908) Sur les annelides polychetes rapportes par M. Le Dr. Rivet de Payta (Perou). Bulletin du Museum National d'Histoire Naturelle, 14, 40 - 44. https: // doi. org / 10.5962 / bhl. part. 5649" type="journal article" year="1908">Gravier 1908</bibRefCitation>
)
</taxonomicName>
. Specially,
<taxonomicName id="4C384D5C361CFFA26525FC5DD3CB95D0" authorityName="Webster" authorityYear="1884" box="[1067,1173,944,968]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="goodei">
<emphasis id="B94CEACD361CFFA26525FC5DD3CB95D0" box="[1067,1173,944,968]" italics="true" pageId="5" pageNumber="66">B. goodei</emphasis>
</taxonomicName>
currently is reported as having a wide distribution which includes Asia, Africa and both American coasts (
<bibRefCitation id="EFA94B2E361CFFA2654BFC38D3B895F4" author="Webster, H. E." box="[1093,1254,980,1005]" pageId="5" pageNumber="66" pagination="305 - 327" refId="ref15636" refString="Webster, H. E. (1884) Annelida from Bermuda, collected by G. Brown Goode. Bulletin United States National Museum, 25, 305 - 327. https: // doi. org / 10.5479 / si. 03629236.25.305" type="journal article" year="1884">Webster 1884</bibRefCitation>
;
<bibRefCitation id="EFA94B2E361CFFA265FDFC38D2C995F4" author="Augener, H." box="[1267,1431,980,1004]" pageId="5" pageNumber="66" pagination="67 - 625" refId="ref13025" refString="Augener, H. (1918) Polychaeta. Beitrage zur Kenntnis des Meeresfauna West-Afrikas. Fol. 2. Lief. 2. Herausgegeben von W. Michaelsen, Hamburg, 559 pp. [pp. 67 - 625]" type="book chapter" year="1918">Augener 1918</bibRefCitation>
;
<bibRefCitation id="EFA94B2E361CFFA26199FC14D6749208" author="Monro, C. C. A." box="[151,298,1016,1040]" pageId="5" pageNumber="66" pagination="1 - 96" refId="ref14583" refString="Monro, C. C. A. (1933) The Polychaeta Errantia collected by Dr C. Crossland at Colon, in the Panama Region, and the Galapagos Islands during the expedition of the S. Y. St. George. Proceedings of the Zoological Society of London, 1933, 1 - 96. https: // doi. org / 10.1111 / j. 1096 - 3642.1933. tb 01578. x" type="journal article" year="1933">Monro 1933</bibRefCitation>
;
<bibRefCitation id="EFA94B2E361CFFA26035FC1BD52E9208" author="Imajima, M. &amp; Hartman, O." box="[315,624,1015,1040]" pageId="5" pageNumber="66" pagination="1 - 237" refId="ref14100" refString="Imajima, M. &amp; Hartman, O. (1964) The polychaetous annelids of Japan. Part I. Allan Hancock Fundation Publications, Occasional paper, 26, 1 - 237." type="journal article" year="1964">Imajima &amp; Hartman 1964</bibRefCitation>
).
<taxonomicName id="4C384D5C361CFFA26386FC14D40C9208" authorityName="Webster" authorityYear="1884" box="[648,850,1016,1040]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="goodei">
<emphasis id="B94CEACD361CFFA26386FC14D40C9208" box="[648,850,1016,1040]" italics="true" pageId="5" pageNumber="66">Bhawania goodei</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFA94B2E361CFFA2653BFC1BD3B89208" author="Webster, H. E." box="[1077,1254,1015,1040]" pageId="5" pageNumber="66" pagination="305 - 327" refId="ref15636" refString="Webster, H. E. (1884) Annelida from Bermuda, collected by G. Brown Goode. Bulletin United States National Museum, 25, 305 - 327. https: // doi. org / 10.5479 / si. 03629236.25.305" type="journal article" year="1884">Webster (1884)</bibRefCitation>
from
<collectingCountry id="F32F764F361CFFA2643BFC1BD2C29208" box="[1333,1436,1015,1040]" name="Bermuda" pageId="5" pageNumber="66">Bermuda</collectingCountry>
Island; later,
<bibRefCitation id="EFA94B2E361CFFA2603CFBF0D682922D" author="Perkins, T. H." box="[306,476,1052,1077]" pageId="5" pageNumber="66" pagination="856 - 915" refId="ref14825" refString="Perkins, T. H. (1985) Chrysopetalum, Bhawania and two new genera of Chrysopetalidae (Polychaeta) from Florida. Proceedings of the Biological Society of Washington, 98, 856 - 915." type="journal article" year="1985">Perkins (1985)</bibRefCitation>
redescribed the species with specimens from Florida. This species has also been recorded by
<bibRefCitation id="EFA94B2E361CFFA26022FBD3D44C9240" author="Kudenov, J. D." box="[300,786,1087,1112]" pageId="5" pageNumber="66" pagination="65 - 91" refId="ref14238" refString="Kudenov, J. D. (1975) Errant polychaetes from the Gulf of California. Journal of Natural History, 9, 65 - 91. https: // doi. org / 10.1080 / 00222937500770061" type="journal article" year="1975">Kudenov (1975, Puerto Peñasco, Sonora)</bibRefCitation>
,
<bibRefCitation id="EFA94B2E361CFFA2622FFBD3D3A19240" author="Fauchald, K." box="[801,1279,1087,1112]" pageId="5" pageNumber="66" pagination="1 - 81" refId="ref13517" refString="Fauchald, K. (1977) Polychaetes from intertidal areas in Panama, with a review of previous shallow-water records. Smithsonian Contributions to Zoology, 221, 1 - 81. https: // doi. org / 10.5479 / si. 00810282.221" type="journal article" year="1977">
Fauchald (1977, Taboga Island,
<collectingCountry id="F32F764F361CFFA26593FBACD3A69240" box="[1181,1272,1088,1112]" name="Panama" pageId="5" pageNumber="66">Panama</collectingCountry>
)
</bibRefCitation>
and Bastida-
<bibRefCitation id="EFA94B2E361CFFA26199FB88D5219264" author="Bastida-Zavala, J. R." box="[151,639,1124,1149]" pageId="5" pageNumber="66" pagination="11 - 39" refId="ref13067" refString="Bastida-Zavala, J. R. (1993) Taxonomia y composicion biogeografica de los poliquetos (Annelida: Polychaeta) de la bahia de La Paz, B. C. S., Mexico. Revista de Investigacion Cientifica, UABCS, 4 (1), 11 - 39." type="journal article" year="1993">Zavala (1993, La Paz, Baja California Sur)</bibRefCitation>
. These records need to be thorougly checked as seems unlikely that
<taxonomicName id="4C384D5C361CFFA2648AFB89D7BD92B8" authorityName="Webster" authorityYear="1884" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="goodei">
<emphasis id="B94CEACD361CFFA2648AFB89D7BD92B8" italics="true" pageId="5" pageNumber="66">B. goodei</emphasis>
</taxonomicName>
has traversed the
<collectingCountry id="F32F764F361CFFA260BEFB65D55792B8" box="[432,521,1161,1184]" name="Panama" pageId="5" pageNumber="66">Panama</collectingCountry>
Canal to the Pacific coast of America. The specimen revised here agree with the redescription by
<bibRefCitation id="EFA94B2E361CFFA2605AFB40D6A792DD" author="Perkins, T. H." box="[340,505,1196,1221]" pageId="5" pageNumber="66" pagination="856 - 915" refId="ref14825" refString="Perkins, T. H. (1985) Chrysopetalum, Bhawania and two new genera of Chrysopetalidae (Polychaeta) from Florida. Proceedings of the Biological Society of Washington, 98, 856 - 915." type="journal article" year="1985">Perkins (1985)</bibRefCitation>
except for the presence of the nuchal fold (
<bibRefCitation id="EFA94B2E361CFFA262ECFB40D3B692DC" author="Perkins, T. H." box="[994,1256,1196,1221]" pageId="5" pageNumber="66" pagination="856 - 915" refId="ref14825" refString="Perkins, T. H. (1985) Chrysopetalum, Bhawania and two new genera of Chrysopetalidae (Polychaeta) from Florida. Proceedings of the Biological Society of Washington, 98, 856 - 915." type="journal article" year="1985">Perkins 1985: fig. 21A</bibRefCitation>
). This character was not observed since the single specimen is damaged.
</paragraph>
<paragraph id="8B8736DF361CFFA261C9FB18D26B9385" blockId="5.[151,1437,151,1437]" pageId="5" pageNumber="66">
<bibRefCitation id="EFA94B2E361CFFA261C9FB18D62A9314" author="Gravier, M. C." box="[199,372,1268,1293]" pageId="5" pageNumber="66" pagination="40 - 44" refId="ref13655" refString="Gravier, M. C. (1908) Sur les annelides polychetes rapportes par M. Le Dr. Rivet de Payta (Perou). Bulletin du Museum National d'Histoire Naturelle, 14, 40 - 44. https: // doi. org / 10.5962 / bhl. part. 5649" type="journal article" year="1908">Gravier (1908)</bibRefCitation>
described briefly
<taxonomicName id="4C384D5C361CFFA2635AFB18D4509314" baseAuthorityName="Gravier" baseAuthorityYear="1908" box="[596,782,1268,1292]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="riveti">
<emphasis id="B94CEACD361CFFA2635AFB18D4509314" box="[596,782,1268,1292]" italics="true" pageId="5" pageNumber="66">Bhawania riveti</emphasis>
</taxonomicName>
based on one incomplete specimen from Payta,
<collectingCountry id="F32F764F361CFFA2646CFB18D2C79314" box="[1378,1433,1268,1292]" name="Peru" pageId="5" pageNumber="66">Perú</collectingCountry>
.
<taxonomicName id="4C384D5C361CFFA26199FAF4D6DC9328" authorityName="Webster" authorityYear="1884" box="[151,386,1303,1328]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="goodei">
<emphasis id="B94CEACD361CFFA26199FAF4D6579328" box="[151,265,1304,1328]" italics="true" pageId="5" pageNumber="66">Bhawania</emphasis>
cf.
<emphasis id="B94CEACD361CFFA26038FAF4D6DC9328" box="[310,386,1304,1328]" italics="true" pageId="5" pageNumber="66">goodei</emphasis>
</taxonomicName>
differs from Graviers in some features: notoaciculum in
<taxonomicName id="4C384D5C361CFFA2651AFAF5D32E9337" baseAuthorityName="Gravier" baseAuthorityYear="1908" box="[1044,1136,1304,1328]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="riveti">
<emphasis id="B94CEACD361CFFA2651AFAF5D32E9337" box="[1044,1136,1304,1328]" italics="true" pageId="5" pageNumber="66">B. riveti</emphasis>
</taxonomicName>
is wide and long (
<bibRefCitation id="EFA94B2E361CFFA2644BFAFBD626934D" author="Gravier, M. C." pageId="5" pageNumber="66" pagination="93 - 126" refId="ref13709" refString="Gravier, M. C. (1910) Annelides polychetes recueillis a Payta (Perou) par M. le Dr. Rivet, Membre de la Mission geodesique de l'Equateur. In: France. Ministere de l'instruction publique, Mission du service geographique de l'Armee pour la mesure d'un arc de meridien equatorial en Amerique du Sud sous le controle scientifique de l'Academie des Sciences 1899 - 1906. Tome 9 (Zoologie). Fascicule 3 (Mollusques, Annelides, Oligochetes). Gauthier-Villars, Paris, pp. 93 - 126. https: // doi. org / 10.5962 / bhl. title. 980" type="book chapter" year="1910">Gravier 1910: pl. VII, fig.31</bibRefCitation>
) while
<taxonomicName id="4C384D5C361CFFA260C2FAD1D507934C" authorityName="Webster" authorityYear="1884" box="[460,601,1340,1365]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="goodei">
<emphasis id="B94CEACD361CFFA260C2FAD1D6BA934C" box="[460,484,1341,1364]" italics="true" pageId="5" pageNumber="66">B.</emphasis>
cf.
<emphasis id="B94CEACD361CFFA26303FAD0D507934C" box="[525,601,1340,1364]" italics="true" pageId="5" pageNumber="66">goodei</emphasis>
</taxonomicName>
is smaller and slimmer (
<figureCitation id="13032A5A361CFFA26262FAD0D498934C" box="[876,966,1340,1365]" captionStart="FIGURE 3" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[178,1409,200,1931]" captionTargetId="figure@8.[170,1423,189,1942]" captionTargetPageId="8" captionText="FIGURE 3. Bhawania cf. goodei. A, parapodium middle segment; BE, neurochaetal types; F, lateral paleae; G, subunit 1 palea; H, meddle main palea; IK, median paleae group. Scale bars: 30 µm (BCDE), 100 µm (FGHIJK), 200 µm (A)." httpUri="https://zenodo.org/record/2609674/files/figure.png" pageId="5" pageNumber="66">Fig. 3A</figureCitation>
); regarding paleae fan,
<bibRefCitation id="EFA94B2E361CFFA265C5FAD0D22C934C" author="Gravier, M. C." box="[1227,1394,1340,1365]" pageId="5" pageNumber="66" pagination="40 - 44" refId="ref13655" refString="Gravier, M. C. (1908) Sur les annelides polychetes rapportes par M. Le Dr. Rivet de Payta (Perou). Bulletin du Museum National d'Histoire Naturelle, 14, 40 - 44. https: // doi. org / 10.5962 / bhl. part. 5649" type="journal article" year="1908">Gravier (1908)</bibRefCitation>
did not mention if his species presented internal raised ribs in the paleae, character present in
<taxonomicName id="4C384D5C361CFFA265AEFA8DD26D9360" authorityName="Webster" authorityYear="1884" box="[1184,1331,1375,1400]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="goodei">
<emphasis id="B94CEACD361CFFA265AEFA8DD3E69360" box="[1184,1208,1377,1400]" italics="true" pageId="5" pageNumber="66">B.</emphasis>
cf.
<emphasis id="B94CEACD361CFFA265E9FA8CD26D9360" box="[1255,1331,1376,1400]" italics="true" pageId="5" pageNumber="66">goodei</emphasis>
</taxonomicName>
; also the length of the paleae in
<taxonomicName id="4C384D5C361CFFA26098FA69D6AE9384" baseAuthorityName="Gravier" baseAuthorityYear="1908" box="[406,496,1413,1436]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="riveti">
<emphasis id="B94CEACD361CFFA26098FA69D6AE9384" box="[406,496,1413,1436]" italics="true" pageId="5" pageNumber="66">B. riveti</emphasis>
</taxonomicName>
is about 1/3 of body width, while in
<taxonomicName id="4C384D5C361CFFA26280FA69D3439384" authorityName="Webster" authorityYear="1884" box="[910,1053,1412,1437]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="66" phylum="Annelida" rank="species" species="goodei">
<emphasis id="B94CEACD361CFFA26280FA69D4C19384" box="[910,927,1413,1436]" italics="true" pageId="5" pageNumber="66">B</emphasis>
. cf.
<emphasis id="B94CEACD361CFFA262DFFA68D3439384" box="[977,1053,1412,1436]" italics="true" pageId="5" pageNumber="66">goodei</emphasis>
</taxonomicName>
is about 1/2 body width.
</paragraph>
</subSubSection>
</treatment>
</document>