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<mods:title id="A788010B4855797BF7E5FD5527EBBF04">A taxonomic revision and revalidation of Nycterilampus Montrouzier (Coleoptera: Elateridae, Agrypninae)</mods:title>
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<taxonomicName id="4C7015E3FD76FFC840FCD7CC8DA1CEB4" authority="Montrouzier" authorityName="Montrouzier" box="[151,491,151,177]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC840FCD7CC8D0DCEB4" bold="true" box="[151,327,151,177]" italics="true" pageId="1" pageNumber="48">Nycterilampus</emphasis>
Montrouzier
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, revalidated
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36A3DEBFD76FFC840FCD7BA8845CC46" pageId="1" pageNumber="48" type="reference_group">
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<taxonomicName id="4C7015E3FD76FFC840FCD7BA8E09CEFD" authority="Montrouzier, 1860: 258" authorityName="Montrouzier" authorityPageNumber="258" authorityYear="1860" box="[151,579,225,248]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC840FCD7BA8D79CEFD" box="[151,307,225,248]" italics="true" pageId="1" pageNumber="48">Nycterilampus</emphasis>
<treatmentCitation id="0AD14871FD76FFC84156D7BA8E09CEFD" author="Montrouzier" box="[317,579,225,248]" page="258" pageId="1" pageNumber="48" year="1860">
<bibRefCitation id="EFE11391FD76FFC84156D7BA8E47CEFD" author="Montrouzier" box="[317,525,225,248]" pageId="1" pageNumber="57" refString="Montrouzier, X. (1860) Essai sur la faune entomologique de la Nouvelle - Caledonie (Balade) et des iles des Pins, Art, Lifu, etc. Annales de la Societe Entomologique de France, serie 3, tome 8, 227 - 308." type="journal article" year="1860">Montrouzier, 1860</bibRefCitation>
: 258
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(original description);
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<bibRefCitation id="EFE11391FD76FFC84323D7BA8F91CEFD" author="Fauvel" box="[840,987,225,248]" pageId="1" pageNumber="57" refString="Fauvel, A. (1904) Faune analytique des coleopteres de la Nouvelle-Caledonie. Revue d ' Entomologie, 23, 132 - 133." type="journal article" year="1904">Fauvel, 1904</bibRefCitation>
: 132
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;
<treatmentCitation id="0AD14871FD76FFC8444ED7BA88BCCEFD" author="Hyslop" box="[1061,1270,225,248]" page="658" pageId="1" pageNumber="48" year="1921">
<bibRefCitation id="EFE11391FD76FFC8444ED7BA88F6CEFD" author="Hyslop" box="[1061,1212,225,248]" pageId="1" pageNumber="57" refString="Hyslop, J. A. (1921) Genotypes of the Elaterid beetles of the world. Proceedings of the United States National Museum, 58, 621 - 680." type="journal article" year="1921">Hyslop, 1921</bibRefCitation>
: 658
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. (
<typeStatus id="54CBD0C2FD76FFC84566D7BA8976CEFD" box="[1293,1340,225,248]" pageId="1" pageNumber="48">type</typeStatus>
species:
<taxonomicName id="4C7015E3FD76FFC840ACD65A8E06CF1D" authority="Montrouzier" authorityName="Montrouzier" box="[199,588,257,280]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="lifuanus">
<emphasis id="B904B272FD76FFC840ACD65A8DF5CF1D" box="[199,447,257,280]" italics="true" pageId="1" pageNumber="48">Nycterilampus lifuanus</emphasis>
Montrouzier
</taxonomicName>
, monotypic);
<treatmentCitation id="0AD14871FD76FFC8428BD65A8F9ACF1D" author="Schenkling" box="[736,976,257,280]" page="478" pageId="1" pageNumber="48" year="1927">
<bibRefCitation id="EFE11391FD76FFC8428BD65A8FD4CF1D" author="Schenkling" box="[736,926,257,280]" pageId="1" pageNumber="57" refString="Schenkling, S. (1927) Elateridae II. In: W. Junk &amp; Schenkling, S. (Eds.) Coleopterorum Catalogus. W. Junk, Berlin, pt. 88, n. 11, pp. 265 - 639." type="journal article" year="1927">Schenkling, 1927</bibRefCitation>
: 478
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(cat.).
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</paragraph>
<paragraph id="8BCF6E60FD76FFC840FCD67A8940CF3D" blockId="1.[151,1437,225,312]" box="[151,1290,289,312]" pageId="1" pageNumber="48">
<treatmentCitationGroup id="AB60494EFD76FFC840FCD67A8940CF3D" box="[151,1290,289,312]" pageId="1" pageNumber="48">
<taxonomicName id="4C7015E3FD76FFC840FCD67A8E51CF3D" authority="Fleutiaux, 1891: 391" authorityName="Fleutiaux" authorityPageNumber="391" authorityYear="1891" box="[151,539,289,312]" class="Insecta" family="Elateridae" genus="Nycterolampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC840FCD67A8D7DCF3D" box="[151,311,289,312]" italics="true" pageId="1" pageNumber="48">Nycterolampus</emphasis>
<treatmentCitation id="0AD14871FD76FFC84155D67A8E51CF3D" author="Fleutiaux" box="[318,539,289,312]" page="391" pageId="1" pageNumber="48" year="1891">
<bibRefCitation id="EFE11391FD76FFC84155D67A8DA3CF3D" author="Fleutiaux" box="[318,489,289,312]" pageId="1" pageNumber="57" refString="Fleutiaux, E. (1891) Elateridae de la Nouvelle Caledonie et dependences. Annales de la Societe Entomologique de France, 60, 387 - 397." type="journal article" year="1891">Fleutiaux, 1891</bibRefCitation>
: 391
</treatmentCitation>
</taxonomicName>
(misspelling);
<treatmentCitation id="0AD14871FD76FFC842D6D67A8FDECF3D" author="Schwarz" box="[701,916,289,312]" page="254" pageId="1" pageNumber="48" year="1906">
<bibRefCitation id="EFE11391FD76FFC842D6D67A8F15CF3D" author="Schwarz" box="[701,863,289,312]" pageId="1" pageNumber="57" refString="Schwarz, O. (1906) Coleoptera, Fam. Elateridae. In: Wytsman, P. (Ed.). Genera Insectorum, fasc. 46, Verteneuil &amp; Desmet, Bruxelles, pp. 310 - 315." type="book chapter" year="1906">Schwarz, 1906</bibRefCitation>
: 254
</treatmentCitation>
, 282;
<treatmentCitation id="0AD14871FD76FFC843BFD67A888FCF3D" author="Schenkling" box="[980,1221,289,312]" page="478" pageId="1" pageNumber="48" year="1927">
<bibRefCitation id="EFE11391FD76FFC843BFD67A88D9CF3D" author="Schenkling" box="[980,1171,289,312]" pageId="1" pageNumber="57" refString="Schenkling, S. (1927) Elateridae II. In: W. Junk &amp; Schenkling, S. (Eds.) Coleopterorum Catalogus. W. Junk, Berlin, pt. 88, n. 11, pp. 265 - 639." type="journal article" year="1927">Schenkling, 1927</bibRefCitation>
: 478
</treatmentCitation>
(cat.).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCF6E60FD76FFC840FCD63C8845CC46" blockId="1.[151,1438,359,1983]" pageId="1" pageNumber="48">
<emphasis id="B904B272FD76FFC840FCD63C8D1ACF84" bold="true" box="[151,336,359,385]" pageId="1" pageNumber="48">Redescription.</emphasis>
Male. Body about 3.9 times longer than wide, weakly convex. Vestiture consisting of fine, short, dense and semidecumbent yellowish setae, longer on metaventrite, covering the color of the integument. Head convex, with a small longitudinal carina at base; frontal carina complete. Antenna 11 or 12 segmented (
<figureCitation id="134B72E5FD76FFC84145D6878DE2CFF3" box="[302,424,476,502]" captionStart="FIGURES 2 9" captionStartId="4.[151,270,1940,1964]" captionTargetBox="[195,1371,193,1939]" captionTargetId="figure@4.[195,1400,171,1943]" captionTargetPageId="4" captionText="FIGURES 2 9. Nycterilampus velutinus Fleutiaux, 1891. 2, 3, antennae (male, female); 7, head and pronotum (female). N. lifuanus Montrouzier, 1860. 4, 5, antennae (male, female); 6, head; 8, head and pronotum (dorsal); 9 prothorax (ventral)." httpUri="https://zenodo.org/record/186018/files/figure.png" pageId="1" pageNumber="48">Figs. 2, 4</figureCitation>
) serrate or pectinate from antennomere IV. Mouthparts directed anteroventrally. Mandibles bidentate; maxillary palps with 4 articles, apical article securiform; labium with prementum and mentum well sclerotized, labial palps with 3 articles, last article securiform.
</paragraph>
</subSubSection>
<subSubSection id="C36A3DEBFD76FFC840AED50B8845CB51" pageId="1" pageNumber="48" type="description">
<paragraph id="8BCF6E60FD76FFC840AED50B88FDCCDB" blockId="1.[151,1438,359,1983]" pageId="1" pageNumber="48">
Prothorax (
<figureCitation id="134B72E5FD76FFC84121D50B8DF0CC6F" box="[330,442,592,618]" captionStart="FIGURES 2 9" captionStartId="4.[151,270,1940,1964]" captionTargetBox="[195,1371,193,1939]" captionTargetId="figure@4.[195,1400,171,1943]" captionTargetPageId="4" captionText="FIGURES 2 9. Nycterilampus velutinus Fleutiaux, 1891. 2, 3, antennae (male, female); 7, head and pronotum (female). N. lifuanus Montrouzier, 1860. 4, 5, antennae (male, female); 6, head; 8, head and pronotum (dorsal); 9 prothorax (ventral)." httpUri="https://zenodo.org/record/186018/files/figure.png" pageId="1" pageNumber="48">Figs. 79</figureCitation>
) parallel sided, lateral margins entirely carinate, hind angles produced and carinate, divergent, anterior angles small, rounded. Pronotum with a pair of glabrous and shiny median lateral spots. Prosternal lobe produced and with two lateral spines; prosternal process horizontal with preapical tooth. Scutellum (
<figureCitation id="134B72E5FD76FFC84176D59F8DE2CCDB" box="[285,424,708,734]" captionStart="FIGURES 10 18" captionStartId="5.[151,270,1942,1966]" captionTargetBox="[191,1377,200,1942]" captionTargetId="figure@5.[191,1378,197,1942]" captionTargetPageId="5" captionText="FIGURES 10 18. Nycterilampus lifuanus Montrouzier, 1860 (male). 10, 11, mesoscutum and scutellum (dorsal, lateral); 12, 13, mesoventrite (ventral, lateral); 14, 15, pterothorax (ventral, ventro lateral); 16, tarsal claw; 17, hind wing; 18, metanotum." httpUri="https://zenodo.org/record/186019/files/figure.png" pageId="1" pageNumber="48">Figs. 10, 11</figureCitation>
) pentagonal and abruptly elevated above the level of mesoscutum.
</paragraph>
<paragraph id="8BCF6E60FD76FFC840AED5B18FFFCD7D" blockId="1.[151,1438,359,1983]" pageId="1" pageNumber="48">
Mesoventrite (
<figureCitation id="134B72E5FD76FFC84113D5B18E41CD01" box="[376,523,746,772]" captionStart="FIGURES 10 18" captionStartId="5.[151,270,1942,1966]" captionTargetBox="[191,1377,200,1942]" captionTargetId="figure@5.[191,1378,197,1942]" captionTargetPageId="5" captionText="FIGURES 10 18. Nycterilampus lifuanus Montrouzier, 1860 (male). 10, 11, mesoscutum and scutellum (dorsal, lateral); 12, 13, mesoventrite (ventral, lateral); 14, 15, pterothorax (ventral, ventro lateral); 16, tarsal claw; 17, hind wing; 18, metanotum." httpUri="https://zenodo.org/record/186019/files/figure.png" pageId="1" pageNumber="48">Figs. 1213</figureCitation>
) with posterior area marginated and steeply declivous to anterior borders. Mesoventral cavity deep, parallel sided, borders narrowing abruptly between the mesocoxae, floor of cavity with an elongated, narrow and shiny median band, without golden setae; mesocoxal cavity open to both mesepimeron and mesepisternum; meso-metaventral suture distinct.
</paragraph>
<paragraph id="8BCF6E60FD76FFC840AED4DE8F1ECA46" blockId="1.[151,1438,359,1983]" pageId="1" pageNumber="48">
<collectingCountry id="F3672EF0FD76FFC840AED4DE8D48CD9A" box="[197,258,901,927]" name="India" pageId="1" pageNumber="48">Hind</collectingCountry>
wing (
<figureCitation id="134B72E5FD76FFC84132D4DE8DF8CD9A" box="[345,434,901,927]" captionStart="FIGURES 10 18" captionStartId="5.[151,270,1942,1966]" captionTargetBox="[191,1377,200,1942]" captionTargetId="figure@5.[191,1378,197,1942]" captionTargetPageId="5" captionText="FIGURES 10 18. Nycterilampus lifuanus Montrouzier, 1860 (male). 10, 11, mesoscutum and scutellum (dorsal, lateral); 12, 13, mesoventrite (ventral, lateral); 14, 15, pterothorax (ventral, ventro lateral); 16, tarsal claw; 17, hind wing; 18, metanotum." httpUri="https://zenodo.org/record/186019/files/figure.png" pageId="1" pageNumber="48">Fig. 17</figureCitation>
) about 2.5 times as long as wide; radial cell about 5 times longer than wide with proximal, posterior angle obtuse; r3 0.3 times total length of radial cell; r4 vestigial; CuA1 crossvein-like, between MP3+4 and CuA2; apical area bearing a triangular sclerotization posteriorly to the radial cell and two narrow, elongated, oblique and convergent sclerotizations at base; medial area with 5 free veins (MP3, MP4 + CuA1, CuA2, CuA3+4 + AA1+2 and AA3+4); wedge cell absent.
</paragraph>
<paragraph id="8BCF6E60FD76FFC840AED3098FF1CB02" blockId="1.[151,1438,359,1983]" pageId="1" pageNumber="48">
Tibia compressed laterally, apices with two small and subequal spurs and a row of spiniform setae on outer and inner margins; tarsomeres simple, ventrally pilose; apex of tarsomeres 13 bearing a row of lateroventrally spiniform setae; tarsomere 1 longer than 5, shorter than 24 combined; 2+3 shorter than 1; 2 longer than 3; 4 shorter than 3. Tarsal claw (
<figureCitation id="134B72E5FD76FFC842D9D39D8F40CAE5" box="[690,778,1222,1248]" captionStart="FIGURES 10 18" captionStartId="5.[151,270,1942,1966]" captionTargetBox="[191,1377,200,1942]" captionTargetId="figure@5.[191,1378,197,1942]" captionTargetPageId="5" captionText="FIGURES 10 18. Nycterilampus lifuanus Montrouzier, 1860 (male). 10, 11, mesoscutum and scutellum (dorsal, lateral); 12, 13, mesoventrite (ventral, lateral); 14, 15, pterothorax (ventral, ventro lateral); 16, tarsal claw; 17, hind wing; 18, metanotum." httpUri="https://zenodo.org/record/186019/files/figure.png" pageId="1" pageNumber="48">Fig. 16</figureCitation>
) simple bearing one long seta at base arising from the outer flat portion; empodium sclerotized with two strong short setae.
</paragraph>
<paragraph id="8BCF6E60FD76FFC840AED2488845CB51" blockId="1.[151,1438,359,1983]" pageId="1" pageNumber="48">Female. Longer than male and more convex. Antenna 11 or 12 segmented and serrate (Figs. 3,5). Baculi 3.74.0 times longer than coxites and bursa copulatrix without inner spines.</paragraph>
</subSubSection>
<subSubSection id="C36A3DEBFD76FFCA40AED23B8F4BCEDD" lastPageId="3" lastPageNumber="50" pageId="1" pageNumber="48" type="discussion">
<paragraph id="8BCF6E60FD76FFC840AED23B88B7CBA4" blockId="1.[151,1438,359,1983]" pageId="1" pageNumber="48">
<emphasis id="B904B272FD76FFC840AED23B8D1ACB7F" bold="true" box="[197,336,1376,1402]" pageId="1" pageNumber="48">Discussion.</emphasis>
<bibRefCitation id="EFE11391FD76FFC84133D23A8EDECB7E" box="[344,660,1377,1403]" pageId="1" pageNumber="57" refString="Van Zwaluwenburg, R. H. (1932) Check list of the Elateridae of Oceania. Bishop Museum Occasional Papers, 9, 3 - 28." type="journal article">Van Zwaluwenburg (1932)</bibRefCitation>
in a footnote mentioned that
<taxonomicName id="4C7015E3FD76FFC8439BD23A88D1CB7F" box="[1008,1179,1377,1402]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC8439BD23A88D1CB7F" box="[1008,1179,1377,1402]" italics="true" pageId="1" pageNumber="48">Nycterilampus</emphasis>
</taxonomicName>
should be a synonym of
<taxonomicName id="4C7015E3FD76FFC840DDD2D38D5FCBA4" box="[182,277,1416,1441]" class="Insecta" family="Elateridae" genus="Tetrigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC840DDD2D38D5FCBA4" box="[182,277,1416,1441]" italics="true" pageId="1" pageNumber="48">Tetrigus</emphasis>
</taxonomicName>
. In his paper on New Caledonian elaterids
<bibRefCitation id="EFE11391FD76FFC8437BD2DC8F9FCBA4" author="Fleutiaux" box="[784,981,1415,1441]" pageId="1" pageNumber="57" refString="Fleutiaux, E. (1947) Revision des Elaterides (Coleopteres) de L'Indo-Chine Francaise. Notes d ' Entomologie Chinoise, 11, 233 - 420." type="journal article" year="1947">Fleutiaux (1947)</bibRefCitation>
accepted that synonymy.
</paragraph>
<paragraph id="8BCF6E60FD76FFC840AED2F58D6FC8F8" blockId="1.[151,1438,359,1983]" pageId="1" pageNumber="48">
Comparing our specimens with the characters of
<taxonomicName id="4C7015E3FD76FFC8434AD2F58FF6CBC2" box="[801,956,1454,1480]" class="Insecta" family="Elateridae" genus="Tetrigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B904B272FD76FFC8434AD2F58F7ACBC2" box="[801,816,1454,1479]" italics="true" pageId="1" pageNumber="48">T</emphasis>
.
<emphasis id="B904B272FD76FFC8432BD2F58FF6CBC2" box="[832,956,1454,1479]" italics="true" pageId="1" pageNumber="48">parallelus</emphasis>
</taxonomicName>
in the paper by Casari-Chen (
<emphasis id="B904B272FD76FFC8455BD2F58972CBC2" box="[1328,1336,1454,1479]" italics="true" pageId="1" pageNumber="48">l</emphasis>
.
<emphasis id="B904B272FD76FFC84522D2F58915CBC2" box="[1353,1375,1454,1479]" italics="true" pageId="1" pageNumber="48">c.</emphasis>
), we noticed that
<taxonomicName id="4C7015E3FD76FFC8415BD28E8DABCBEB" box="[304,481,1493,1518]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC8415BD28E8DABCBEB" box="[304,481,1493,1518]" italics="true" pageId="1" pageNumber="48">Nycterilampus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C7015E3FD76FFC842EED28E8EA2CBEB" box="[645,744,1493,1518]" class="Insecta" family="Elateridae" genus="Tetrigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC842EED28E8EA2CBEB" box="[645,744,1493,1518]" italics="true" pageId="1" pageNumber="48">Tetrigus</emphasis>
</taxonomicName>
(sense strict) by: frons with complete carina; mandibles bidentate, anterior prosternal lobe with a blunt spine on either side (
<figureCitation id="134B72E5FD76FFC843C1D2A08FBBC810" box="[938,1009,1531,1557]" captionStart="FIGURES 2 9" captionStartId="4.[151,270,1940,1964]" captionTargetBox="[195,1371,193,1939]" captionTargetId="figure@4.[195,1400,171,1943]" captionTargetPageId="4" captionText="FIGURES 2 9. Nycterilampus velutinus Fleutiaux, 1891. 2, 3, antennae (male, female); 7, head and pronotum (female). N. lifuanus Montrouzier, 1860. 4, 5, antennae (male, female); 6, head; 8, head and pronotum (dorsal); 9 prothorax (ventral)." httpUri="https://zenodo.org/record/186018/files/figure.png" pageId="1" pageNumber="48">Fig. 9</figureCitation>
), tibiae with two small and subequal apical spurs; apices of parameres securiform (
<figureCitation id="134B72E5FD76FFC842C4D1798F77C839" box="[687,829,1570,1596]" captionStart="FIGURES 19 26" captionStartId="6.[151,271,1944,1968]" captionTargetBox="[211,1389,190,1919]" captionTargetId="figure@6.[203,1390,180,1924]" captionTargetPageId="6" captionText="FIGURES 19 26. Nycterilampus lifuanus Montrouzier, 1860 (male). 19, sternite IX; 20, tergites IX X; 21, sternite VIII; 22, tergite VIII; 23, abdomen (ventral); 24, 25, aedeagus (dorsal, ventral). N. velutinus Fleutiaux, 1891 (male). 26, aedeagus (ventral)." httpUri="https://zenodo.org/record/186020/files/figure.png" pageId="1" pageNumber="48">Figs. 24, 25</figureCitation>
); baculi 3.74.0 times longer than coxites (
<figureCitation id="134B72E5FD76FFC8455ED1798CF0C866" captionStart="FIGURES 27 30" captionStartId="7.[151,270,1579,1603]" captionTargetBox="[211,1420,201,1552]" captionTargetId="figure@7.[151,1436,194,1555]" captionTargetPageId="7" captionText="FIGURES 27 30. Nycterilampus lifuanus Montrouzier, 1860 (female). 27, tergite VIII; 28, sternite VIII; 29, ovipositor (ventral); 30, ovipositor and reproductive organs (dorsal)." httpUri="https://zenodo.org/record/186021/files/figure.png" pageId="1" pageNumber="48">Figs. 29, 30</figureCitation>
); bursa copulatrix without inner spines. In
<taxonomicName id="4C7015E3FD76FFC842A4D1128FF3C867" box="[719,953,1609,1634]" class="Insecta" family="Elateridae" genus="Tetrigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B904B272FD76FFC842A4D1128FF3C867" box="[719,953,1609,1634]" italics="true" pageId="1" pageNumber="48">Tetrigus parallelus</emphasis>
</taxonomicName>
, the mandibles are unidentate, anterior prosternal lobe without spines on either side, ventral parameres separated at base and baculi about 10.0 times longer than coxites. Although, our study does not include a cladistic analysis, these characters seem to be sufficient to indicate that
<taxonomicName id="4C7015E3FD76FFC841ACD1E68E39C8D3" box="[455,627,1725,1750]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC841ACD1E68E39C8D3" box="[455,627,1725,1750]" italics="true" pageId="1" pageNumber="48">Nycterilampus</emphasis>
</taxonomicName>
form a distinct monophyletic lineage and should be considered as a valid genus.
</paragraph>
<paragraph id="8BCF6E60FD76FFC840AED0518E10C9BA" blockId="1.[151,1438,359,1983]" pageId="1" pageNumber="48">
It is possible that other species of
<taxonomicName id="4C7015E3FD76FFC84232D0518EF2C926" box="[601,696,1802,1827]" class="Insecta" family="Elateridae" genus="Tetrigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC84232D0518EF2C926" box="[601,696,1802,1827]" italics="true" pageId="1" pageNumber="48">Tetrigus</emphasis>
</taxonomicName>
can be transferred to
<taxonomicName id="4C7015E3FD76FFC843D0D051882CC926" box="[955,1126,1802,1827]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC843D0D051882CC926" box="[955,1126,1802,1827]" italics="true" pageId="1" pageNumber="48">Nycterilampus</emphasis>
</taxonomicName>
, for instance,
<taxonomicName id="4C7015E3FD76FFC84564D0518D8DC94E" authority="Van Zwaluwenburg, 1933" authorityName="Van Zwaluwenburg" authorityYear="1933" class="Insecta" family="Elateridae" genus="Tetrigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="fleutiauxi">
<emphasis id="B904B272FD76FFC84564D0518957C926" box="[1295,1309,1802,1827]" italics="true" pageId="1" pageNumber="48">T</emphasis>
.
<emphasis id="B904B272FD76FFC84546D05189D6C926" box="[1325,1436,1802,1827]" italics="true" pageId="1" pageNumber="48">fleutiauxi</emphasis>
<bibRefCitation id="EFE11391FD76FFC840FCD06A8D8DC94E" author="Van" box="[151,455,1841,1867]" pageId="1" pageNumber="57" refString="Van Zwaluwenburg, R. H. (1933) New Elateridae (Col.) from Melanesia. Stylops, 2, 176 - 185." type="journal article" year="1933">Van Zwaluwenburg, 1933</bibRefCitation>
</taxonomicName>
from Ongea-ndriti, Ovalau, Kandavu,
<collectingCountry id="F3672EF0FD76FFC843E4D06A8FF1C94E" box="[911,955,1841,1867]" name="Fiji" pageId="1" pageNumber="48">Viti</collectingCountry>
Levu whose characters mentioned in the literature (
<bibRefCitation id="EFE11391FD76FFC84164D00C8DFEC974" author="Johnson" box="[271,436,1879,1905]" pageId="1" pageNumber="57" refString="Johnson, P. J. (2001) A new species of Cryptalaus from Fiji, with taxonomic and distributional notes and a key to the Hemirhipini of Eastern Melanesia and Polynesia (Coleoptera: Elateridae). Proceedings of the Hawaiian Entomological Society, 35, 1 - 12." type="journal article" year="2001">Johnson 2001</bibRefCitation>
) are appropriate with those of the genus
<taxonomicName id="4C7015E3FD76FFC843E6D0038872C974" box="[909,1080,1880,1905]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC843E6D0038872C974" box="[909,1080,1880,1905]" italics="true" pageId="1" pageNumber="48">Nycterilampus</emphasis>
</taxonomicName>
. However a complete revision of all
<taxonomicName id="4C7015E3FD76FFC840B0D0258D70C992" box="[219,314,1918,1943]" class="Insecta" family="Elateridae" genus="Tetrigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC840B0D0258D70C992" box="[219,314,1918,1943]" italics="true" pageId="1" pageNumber="48">Tetrigus</emphasis>
</taxonomicName>
species is necessary before that change is made. Therefore we restricted the present work to the two original species of
<taxonomicName id="4C7015E3FD76FFC841C3D0FE8E19C9BB" box="[424,595,1957,1982]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD76FFC841C3D0FE8E19C9BB" box="[424,595,1957,1982]" italics="true" pageId="1" pageNumber="48">Nycterilampus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF0F3EE8FD75FFCB40FCD0898F25C9EC" box="[151,879,2002,2026]" httpUri="https://zenodo.org/record/186017/files/figure.png" pageId="2" pageNumber="49" targetBox="[261,1350,196,1953]" targetPageId="2">
<paragraph id="8BCF6E60FD75FFCB40FCD0898F25C9EC" blockId="2.[151,879,2002,2026]" box="[151,879,2002,2026]" pageId="2" pageNumber="49">
<emphasis id="B904B272FD75FFCB40FCD0898D53C9EF" bold="true" box="[151,281,2002,2026]" pageId="2" pageNumber="49">FIGURE 1.</emphasis>
<taxonomicName id="4C7015E3FD75FFCB414BD0898E99C9EC" authority="Fleutiaux, 1891" authorityName="Fleutiaux" authorityYear="1891" box="[288,723,2002,2025]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B904B272FD75FFCB414BD0898E68C9EC" box="[288,546,2002,2025]" italics="true" pageId="2" pageNumber="49">Nycterilampus velutinus</emphasis>
Fleutiaux, 1891
</taxonomicName>
. Male habitus.
</paragraph>
</caption>
<paragraph id="8BCF6E60FD74FFCA40ADD7C38F4BCEDD" blockId="3.[151,1436,152,216]" pageId="3" pageNumber="50">
In order to better clarify the differences between
<taxonomicName id="4C7015E3FD74FFCA4374D7C3884ACEB4" box="[799,1024,152,177]" class="Insecta" family="Elateridae" genus="Tetrigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B904B272FD74FFCA4374D7C3884ACEB4" box="[799,1024,152,177]" italics="true" pageId="3" pageNumber="50">Tetrigus parallelus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7015E3FD74FFCA4454D7C388A0CEB4" box="[1087,1258,152,177]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD74FFCA4454D7C388A0CEB4" box="[1087,1258,152,177]" italics="true" pageId="3" pageNumber="50">Nycterilampus</emphasis>
</taxonomicName>
as well as the
<taxonomicName id="4C7015E3FD74FFCA40FCD7E48D08CEDD" box="[151,322,191,216]" class="Insecta" family="Elateridae" genus="Nycterilampus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B904B272FD74FFCA40FCD7E48D08CEDD" box="[151,322,191,216]" italics="true" pageId="3" pageNumber="50">Nycterilampus</emphasis>
</taxonomicName>
species we present the following key:
</paragraph>
</subSubSection>
</treatment>
</document>