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<mods:title id="C4C481A1842CDBEB103C01DDF5A39807">Early-Middle Ordovician cephalopods from Ny Friesland, Spitsbergen - a pelagic fauna with Laurentian affinities</mods:title>
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<mods:namePart id="2BC009EB00DF52D5D5820B8AF716714F">Kröger, Björn</mods:namePart>
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<treatment id="5F4487ACFF9FFF9AFD9E7CB6FEA17BBA" ID-DOI="http://doi.org/10.5281/zenodo.5793548" ID-GBIF-Taxon="191693650" ID-Zenodo-Dep="5793548" LSID="urn:lsid:plazi:treatment:5F4487ACFF9FFF9AFD9E7CB6FEA17BBA" httpUri="http://treatment.plazi.org/id/5F4487ACFF9FFF9AFD9E7CB6FEA17BBA" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
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<heading id="8C1A81D6FF9FFF9DFD9E7CB6FC3279D8" box="[584,1004,930,957]" centered="true" fontSize="11" level="2" pageId="51" pageNumber="52" reason="2">
Genus
<taxonomicName id="10ED4D39FF9FFF9DFD4C7CB6FC3279D8" authority="Hyatt, 1900" authorityName="Hyatt" authorityYear="1900" box="[666,1004,930,957]" class="Cephalopoda" family="Protocycloceratidae" genus="Protocycloceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetrabranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="genus">
<emphasis id="E599EAA8FF9FFF9DFD4C7CB6FC8479D9" bold="true" box="[666,858,930,957]" italics="true" pageId="51" pageNumber="52">Protocycloceras</emphasis>
Hyatt, 1900
</taxonomicName>
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<subSubSection id="9FF76531FF9FFF9DFF6B7CF0FBAB7E2B" pageId="51" pageNumber="52" type="type_taxon">
<paragraph id="D75236BAFF9FFF9DFF6B7CF0FE867999" blockId="51.[189,344,995,1022]" box="[189,344,995,1022]" pageId="51" pageNumber="52">
<heading id="8C1A81D6FF9FFF9DFF6B7CF0FE867999" bold="true" box="[189,344,995,1022]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="E599EAA8FF9FFF9DFF6B7CF0FE867999" bold="true" box="[189,344,995,1022]" pageId="51" pageNumber="52">
<typeStatus id="08568818FF9FFF9DFF6B7CF0FF24799A" box="[189,250,996,1022]" pageId="51" pageNumber="52">Type</typeStatus>
species
</emphasis>
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<paragraph id="D75236BAFF9FFF9DFF6B7B06FBAB7E2B" blockId="51.[189,1398,1042,1103]" pageId="51" pageNumber="52">
<taxonomicName id="10ED4D39FF9FFF9DFF6B7B06FDAD7E49" authority="Billings, 1859" authorityName="Billings" authorityYear="1859" box="[189,627,1042,1069]" class="Liliopsida" family="Orchidaceae" genus="Orthoceras" kingdom="Plantae" order="Asparagales" pageId="51" pageNumber="52" phylum="Tracheophyta" rank="species" species="lamarcki">
<emphasis id="E599EAA8FF9FFF9DFF6B7B06FE697E48" box="[189,439,1042,1068]" italics="true" pageId="51" pageNumber="52">Orthoceras lamarcki</emphasis>
<bibRefCitation id="B37C4B4BFF9FFF9DFE127B06FDAD7E49" author="Billings E." box="[452,627,1042,1069]" pageId="51" pageNumber="52" pagination="345 - 367" refId="ref45339" refString="Billings E. 1859. Fossils of the calciferous sandrock, including those of a deposit of White Limestone at Mingan, supposed to belong to the formation. Canadian Naturalist and Geologist, Proceedings of the Society of Natural History Montreal 4: 345 - 367." type="journal article" year="1859">Billings, 1859</bibRefCitation>
</taxonomicName>
, from the Fort Cassin Formation (Floian Stage), Township of Godmanchester, Huntingdon County, southwest
<collectingRegion id="1529F858FF9FFF9DFD207B21FC8D7E2B" box="[758,851,1077,1103]" country="Canada" name="Quebec" pageId="51" pageNumber="52">Quebec</collectingRegion>
; by original designation.
</paragraph>
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<subSubSection id="9FF76531FF9FFF9DFF6B7B63FBA77F0B" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="D75236BAFF9FFF9DFF6B7B63FEE87EF5" blockId="51.[189,310,1143,1169]" box="[189,310,1143,1169]" pageId="51" pageNumber="52">
<heading id="8C1A81D6FF9FFF9DFF6B7B63FEE87EF5" bold="true" box="[189,310,1143,1169]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="E599EAA8FF9FFF9DFF6B7B63FEE87EF5" bold="true" box="[189,310,1143,1169]" pageId="51" pageNumber="52">Diagnosis</emphasis>
</heading>
</paragraph>
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Nearly straight, annulated longicones with circular or subcircular cross section and straight sutures; camerae narrow, about 67 over a length similar to conch cross section; siphuncle located between center and ventral conch margin; siphuncular segments concave; septal necks short, loxochoanitic to orthochoanitic; connecting ring comparatively thick; siphuncular deposits wedge out adorally at steep angle to ventral side; diaphragms in apical portions of siphuncle; epi-and hyposeptal cameral deposits more strongly developed on ventral side (adopted from
<bibRefCitation id="B37C4B4BFF9FFF9DFC9A7A42FBB37F0B" author="Kroger B. &amp; Landing E." box="[844,1133,1365,1392]" pageId="51" pageNumber="52" pagination="664 - 693" refId="ref48326" refString="Kroger B. &amp; Landing E. 2009. Cephalopods and paleoenvironments of the Fort Cassin Formation (Upper Lower Ordovician), eastern New York and adjacent Vermont. Journal of Paleontology 83: 664 - 693. https: // doi. org / 10.1666 / 08 - 181.1" type="journal article" year="2009">Kröger &amp; Landing 2009</bibRefCitation>
).
</paragraph>
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<caption id="83926632FF9FFF9DFF6B7894FBB57D85" ID-DOI="http://doi.org/10.5281/zenodo.5793526" ID-Zenodo-Dep="5793526" httpUri="https://zenodo.org/record/5793526/files/figure.png" pageId="51" pageNumber="52" startId="51.[189,232,1920,1946]" targetBox="[206,1395,1569,1878]" targetPageId="51">
<paragraph id="D75236BAFF9FFF9DFF6B7894FBB57D85" blockId="51.[188,1398,1920,2017]" pageId="51" pageNumber="52">
<emphasis id="E599EAA8FF9FFF9DFF6B7894FEC47DFE" bold="true" box="[189,282,1920,1946]" pageId="51" pageNumber="52">Fig. 34.</emphasis>
Diagrams of relative conch width, relative chamber length, and relative siphuncular diameter of
<taxonomicName id="10ED4D39FF9FFF9DFF6A78B0FDCB7DD9" authority="Kröger &amp; Pohle, 2021" authorityName="Kröger &amp; Pohle" authorityYear="2021" box="[188,533,1955,1981]" class="Cephalopoda" family="Troedssonellidae" genus="Buttsoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="buldrebreenensis" status="sp. nov.">
<emphasis id="E599EAA8FF9FFF9DFF6A78B0FDCB7DD9" box="[188,533,1955,1981]" italics="true" pageId="51" pageNumber="52">Buttsoceras buldrebreenensis</emphasis>
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<taxonomicNameLabel id="FEAA57D3FF9FFF9DFDCB78B0FDA47DDA" box="[541,634,1956,1982]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
, from the Olenidsletta Member, Floian, Ordovician, Profilstranda section, Ny Friesland, Spitsbergen. See Supp. file 1 for details of measurements.
</paragraph>
</caption>
<subSubSection id="9FF76531FF98FF9AFF6B7E1DFEA17BBA" pageId="52" pageNumber="53" type="discussion">
<paragraph id="D75236BAFF98FF9AFF6B7E1DFEF17B47" blockId="52.[189,303,265,291]" box="[189,303,265,291]" pageId="52" pageNumber="53">
<heading id="8C1A81D6FF98FF9AFF6B7E1DFEF17B47" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="E599EAA8FF98FF9AFF6B7E1DFEF17B47" bold="true" box="[189,303,265,291]" pageId="52" pageNumber="53">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="D75236BAFF98FF9AFF6B7E23FEA17BBA" blockId="52.[189,1400,311,478]" pageId="52" pageNumber="53">
The genus
<taxonomicName id="10ED4D39FF98FF9AFE957E23FDDC7B35" authorityName="Hyatt" authorityYear="1900" box="[323,514,311,337]" class="Cephalopoda" family="Protocycloceratidae" genus="Protocycloceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetrabranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="E599EAA8FF98FF9AFE957E23FDDC7B35" box="[323,514,311,337]" italics="true" pageId="52" pageNumber="53">Protocycloceras</emphasis>
</taxonomicName>
is
<typeStatus id="08568818FF98FF9AFDFF7E2CFD827B36" box="[553,604,312,338]" pageId="52" pageNumber="53">type</typeStatus>
genus for the
<taxonomicName id="10ED4D39FF98FF9AFCDB7E23FB127B36" authority="Kobayashi, 1935" authorityName="Kobayashi" authorityYear="1935" box="[781,1228,311,338]" class="Cephalopoda" family="Protocycloceratidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetrabranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="family">Protocycloceratidae Kobayashi, 1935</taxonomicName>
. Based on the cladistic analysis (below) this family is abandoned. The genus clusters consistently with other members of the Troedsonellida, such as
<taxonomicName id="10ED4D39FF98FF9AFDF17E6AFD6C7BF3" authorityName="Ulrich &amp; Foerste" authorityYear="1936" box="[551,690,382,407]" class="Cephalopoda" family="Troedssonellidae" genus="Buttsoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="E599EAA8FF98FF9AFDF17E6AFD6C7BF3" box="[551,690,382,407]" italics="true" pageId="52" pageNumber="53">Buttsoceras</emphasis>
</taxonomicName>
(see below). The close relationship between the two genera is also suggested by the combination of diaphragms and endosiphuncular linings, which is unknown in any other genus.
</paragraph>
</subSubSection>
</treatment>
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