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<document id="F0D821DBD4CF12294AD9F341ED2C4CA3" ID-CLB-Dataset="36839" ID-DOI="10.11646/zootaxa.3814.1.1" ID-GBIF-Dataset="493d9746-7061-41d2-8bff-0713bd07f56f" ID-ISSN="1175-5326" ID-Zenodo-Dep="286211" ID-ZooBank="A024EBCC-AD60-4104-AD4A-4E4843CE6983" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460432840378" checkinUser="plazi" docAuthor="Branch, William R., Bayliss, Julian &amp; Tolley, Krystal A." docDate="2014" docId="03F5304CFFC9FFE9FF21F9B643E4EDE9" docLanguage="en" docName="zt03814p036.pdf" docOrigin="Zootaxa 3814 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhampholeon (Rhinodigitum) bruessoworum Branch, Bayliss &amp; Tolley, 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="31" masterDocId="FFCC4834FFD2FFF7FFB6FFEA4265E875" masterDocTitle="Pygmy chameleons of the Rhampholeon platyceps compex (Squamata: Chamaeleonidae): Description of four new species from isolated sky islands of northern Mozambique" masterLastPageNumber="36" masterPageNumber="1" pageNumber="28" updateTime="1698331660971" updateUser="ExternalLinkService">
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<mods:title id="30220E061EF569A1DD277A07396EF3C1">Pygmy chameleons of the Rhampholeon platyceps compex (Squamata: Chamaeleonidae): Description of four new species from isolated sky islands of northern Mozambique</mods:title>
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<mods:namePart id="759A83B4D32760C4D6F7553A2D23CAD1">Branch, William R.</mods:namePart>
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<mods:namePart id="0001102C77E44C83D5F216FE48094D30">Bayliss, Julian</mods:namePart>
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<mods:namePart id="E7BB5D22F874E9666B6F14C0033E96FA">Tolley, Krystal A.</mods:namePart>
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<mods:title id="C8210363043694340DF03B32F84E192C">Zootaxa</mods:title>
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<mods:date id="2DD703DEF03201A085EC2D696906EEF4">2014</mods:date>
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<mods:number id="595DE8A8AD0BF911369D3741BC40D8EA">3814</mods:number>
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<treatment id="03F5304CFFC9FFE9FF21F9B643E4EDE9" ID-DOI="http://doi.org/10.5281/zenodo.6123219" ID-GBIF-Taxon="127668187" ID-Zenodo-Dep="6123219" LSID="urn:lsid:plazi:treatment:03F5304CFFC9FFE9FF21F9B643E4EDE9" httpUri="http://treatment.plazi.org/id/03F5304CFFC9FFE9FF21F9B643E4EDE9" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28">
<subSubSection id="C346D2D1FFC9FFECFF21F9B64177EEA9" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BE3815AFFC9FFECFF21F9B64147EE03" blockId="27.[151,802,1628,1687]" box="[151,802,1628,1654]" pageId="27" pageNumber="28">
<heading id="D0AB3636FFC9FFECFF21F9B64147EE03" bold="true" box="[151,802,1628,1654]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="B9285D48FFC9FFECFF21F9B64147EE03" bold="true" box="[151,802,1628,1654]" pageId="27" pageNumber="28">
<taxonomicName id="4C5CFAD9FFC9FFECFF21F9B640DAEE03" box="[151,703,1628,1654]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="bruessoworum" status="sp. nov." subGenus="Rhinodigitum">
<emphasis id="B9285D48FFC9FFECFF21F9B64327EE03" bold="true" box="[151,322,1628,1654]" italics="true" pageId="27" pageNumber="28">Rhampholeon</emphasis>
(
<emphasis id="B9285D48FFC9FFECFEE5F9B6439EEE03" bold="true" box="[339,507,1628,1654]" italics="true" pageId="27" pageNumber="28">Rhinodigitum</emphasis>
)
<emphasis id="B9285D48FFC9FFECFDBAF9B640DAEE03" bold="true" box="[524,703,1628,1654]" italics="true" pageId="27" pageNumber="28">bruessoworum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BE033FFC9FFECFD72F9B64147EE03" box="[708,802,1628,1654]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE3815AFFC9FFECFF21F9954060EEE2" blockId="27.[151,802,1628,1687]" box="[151,517,1662,1687]" pageId="27" pageNumber="28">Mount Inago Pygmy Chameleon</paragraph>
<paragraph id="8BE3815AFFC9FFECFF21F92C4177EEA9" blockId="27.[151,786,1734,1757]" box="[151,786,1734,1757]" pageId="27" pageNumber="28">
<emphasis id="B9285D48FFC9FFECFF21F92C4374EEA9" bold="true" box="[151,273,1734,1756]" pageId="27" pageNumber="28">Synonymy:</emphasis>
<taxonomicName id="4C5CFAD9FFC9FFECFEA1F92C43C1EEA9" box="[279,420,1734,1756]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis id="B9285D48FFC9FFECFEA1F92C43C1EEA9" box="[279,420,1734,1756]" italics="true" pageId="27" pageNumber="28">Rhampholeon</emphasis>
</taxonomicName>
sp. Bayliss
<emphasis id="B9285D48FFC9FFECFDA8F922402EEEA9" box="[542,587,1734,1757]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. 2010, p17. fig. 13.
</paragraph>
</subSubSection>
<subSubSection id="C346D2D1FFC9FFECFF21F8E34115EF1F" pageId="27" pageNumber="28" type="etymology">
<paragraph id="8BE3815AFFC9FFECFF21F8E34115EF1F" blockId="27.[151,1437,1801,2006]" pageId="27" pageNumber="28">
<emphasis id="B9285D48FFC9FFECFF21F8E34373EF57" bold="true" box="[151,278,1801,1826]" pageId="27" pageNumber="28">Etymology</emphasis>
. The specific epithet honours the contributions of the brothers Carl and Darren Bruessow to the protection of wildlife in southern
<collectingCountry id="F34BC1CAFFC9FFECFDA6F8C7400DEF33" box="[528,616,1837,1862]" name="Malawi" pageId="27" pageNumber="28">Malawi</collectingCountry>
, particularly via the Mount Mulanje Conservation Trust.
<emphasis id="B9285D48FFC9FFECFF71F8BB4376EF1F" bold="true" box="[199,275,1873,1898]" pageId="27" pageNumber="28">
<typeStatus id="54E73FF8FFC9FFECFF71F8BB436AEF1F" box="[199,271,1873,1898]" pageId="27" pageNumber="28">Types</typeStatus>
.
</emphasis>
The
<typeStatus id="54E73FF8FFC9FFECFEF0F8B84313EF1F" box="[326,374,1874,1898]" pageId="27" pageNumber="28">type</typeStatus>
series comprises three specimens, including:
</paragraph>
</subSubSection>
<subSubSection id="C346D2D1FFC9FFEAFF71F89F47F9E88D" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BE3815AFFC9FFECFF71F89F4380EFA3" blockId="27.[151,1437,1801,2006]" pageId="27" pageNumber="28">
<emphasis id="B9285D48FFC9FFECFF71F89F434BEFFB" box="[199,302,1909,1934]" italics="true" pageId="27" pageNumber="28">
<typeStatus id="54E73FF8FFC9FFECFF71F89F434BEFFB" box="[199,302,1909,1934]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
</emphasis>
.- An adult female (
<collectionCode id="ED4D199FFFC9FFECFDA2F89C402AEFFB" box="[532,591,1910,1934]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="27" pageNumber="28">PEM</collectionCode>
R20375;
<figureCitation id="13679DDFFFC9FFECFD73F89F416CEFFB" box="[709,777,1909,1934]" captionStart="FIGURE 9" captionStartId="28.[151,250,1953,1975]" captionTargetBox="[177,1407,214,1913]" captionTargetId="figure@28.[164,1423,193,1932]" captionTargetPageId="28" captionText="FIGURE 9. Rhampholeon bruessoworum sp. nov.: A—Holotype; B—Allotype; C—Habitat on Mt. Inago (J. Bayliss)." httpUri="https://zenodo.org/record/286220/files/figure.png" pageId="27" pageNumber="28">Fig. 9</figureCitation>
A) collected by J. Bayliss,
<date id="FFE2A79AFFC9FFECFBF0F89F477EEFFB" box="[1094,1307,1909,1934]" pageId="27" pageNumber="28" value="2009-09-05">5 September 2009</date>
, in a small patch of wet forest at the base of a granite inselberg of Mt. Inago, Zambézia Province,
<collectingCountry id="F34BC1CAFFC9FFECFBCEF8734769EFC7" box="[1144,1292,1945,1970]" name="Mozambique" pageId="27" pageNumber="28">Mozambique</collectingCountry>
(15˚04'51”S 37˚23'37”E, ca
<quantity id="4CA42CBFFFC9FFECFEF2F85443FFEFA0" box="[324,410,1982,2006]" metricMagnitude="3" metricUnit="m" metricValue="1.478" pageId="27" pageNumber="28" unit="m" value="1478.0">1478 m</quantity>
a.s.l.).
</paragraph>
<caption id="DF23D1D2FFCEFFEBFF21F84B4734EFC2" box="[151,1361,1953,1975]" httpUri="https://zenodo.org/record/286220/files/figure.png" pageId="28" pageNumber="29" targetBox="[177,1407,214,1913]" targetPageId="28">
<paragraph id="8BE3815AFFCEFFEBFF21F84B4734EFC2" blockId="28.[151,1361,1953,1975]" box="[151,1361,1953,1975]" pageId="28" pageNumber="29">
<emphasis id="B9285D48FFCEFFEBFF21F84B4377EFC2" bold="true" box="[151,274,1953,1975]" pageId="28" pageNumber="29">FIGURE 9.</emphasis>
<taxonomicName id="4C5CFAD9FFCEFFEBFEAFF84B4024EFC2" box="[281,577,1953,1975]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="bruessoworum" status="sp. nov.">
<emphasis id="B9285D48FFCEFFEBFEAFF84B4024EFC2" box="[281,577,1953,1975]" italics="true" pageId="28" pageNumber="29">Rhampholeon bruessoworum</emphasis>
</taxonomicName>
<emphasis id="B9285D48FFCEFFEBFDF1F84B40FDEFC2" bold="true" box="[583,664,1953,1975]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A21BE033FFCEFFEBFDF1F84B40FDEFC2" box="[583,664,1953,1975]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A—Holotype; B—Allotype; C—Habitat on Mt. Inago (J. Bayliss).
</paragraph>
</caption>
<paragraph id="8BE3815AFFCFFFEAFF71FF7D46E7E8C5" blockId="29.[199,1436,151,248]" box="[199,1154,151,176]" pageId="29" pageNumber="30">
<emphasis id="B9285D48FFCFFFEAFF71FF7D4341E8C5" box="[199,292,151,176]" italics="true" pageId="29" pageNumber="30">Allotype</emphasis>
. An adult male (
<collectionCode id="ED4D199FFFCFFFEAFE69FF72407FE8C5" box="[479,538,152,176]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="29" pageNumber="30">PEM</collectionCode>
R20376,
<figureCitation id="13679DDFFFCFFFEAFD3EFF7D40AEE8C5" box="[648,715,151,176]" captionStart="FIGURE 9" captionStartId="28.[151,250,1953,1975]" captionTargetBox="[177,1407,214,1913]" captionTargetId="figure@28.[164,1423,193,1932]" captionTargetPageId="28" captionText="FIGURE 9. Rhampholeon bruessoworum sp. nov.: A—Holotype; B—Allotype; C—Habitat on Mt. Inago (J. Bayliss)." httpUri="https://zenodo.org/record/286220/files/figure.png" pageId="29" pageNumber="30">Fig. 9</figureCitation>
B), same collecting details as
<typeStatus id="54E73FF8FFCFFFEAFBAEFF7D461BE8C5" box="[1048,1150,151,176]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BE3815AFFCFFFEAFF71FF57465AE8A0" blockId="29.[199,1436,151,248]" box="[199,1087,188,213]" pageId="29" pageNumber="30">
<emphasis id="B9285D48FFCFFFEAFF71FF57434BE8A1" box="[199,302,189,212]" italics="true" pageId="29" pageNumber="30">
<typeStatus id="54E73FF8FFCFFFEAFF71FF57434BE8A1" box="[199,302,189,212]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
</emphasis>
. An adult female (
<collectionCode id="ED4D199FFFCFFFEAFE48FF56405CE8A1" box="[510,569,188,212]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="29" pageNumber="30">PEM</collectionCode>
R20374), same collecting details as
<typeStatus id="54E73FF8FFCFFFEAFC63FF56465EE8A0" box="[981,1083,188,213]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BE3815AFFCFFFEAFF71FF3547F9E88D" blockId="29.[199,1436,151,248]" box="[199,1436,223,248]" pageId="29" pageNumber="30">
<emphasis id="B9285D48FFCFFFEAFF71FF35435EE88D" bold="true" box="[199,315,223,248]" pageId="29" pageNumber="30">Meristics.</emphasis>
Measurements of the
<typeStatus id="54E73FF8FFCFFFEAFD99FF0A403AE88D" box="[559,607,224,248]" pageId="29" pageNumber="30">type</typeStatus>
series of
<taxonomicName id="4C5CFAD9FFCFFFEAFD7EFF354669E88D" box="[712,1036,223,248]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="bruessoworum" status="sp. nov.">
<emphasis id="B9285D48FFCFFFEAFD7EFF354669E88D" box="[712,1036,223,248]" italics="true" pageId="29" pageNumber="30">Rhampholeon bruessoworum</emphasis>
</taxonomicName>
<emphasis id="B9285D48FFCFFFEAFBA5FF35460EE88D" bold="true" box="[1043,1131,223,248]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A21BE033FFCFFFEAFBA5FF35460EE88D" box="[1043,1131,223,248]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are summarized in
<tableCitation id="C6DEB4E1FFCFFFEAFAF2FF3547FDE88D" box="[1348,1432,223,248]" captionStart="TABLE 9" captionStartId="29.[151,235,332,355]" captionTargetBox="[159,1269,414,922]" captionTargetPageId="29" captionText="TABLE 9. Measurements (mm) for the type series of Rhampholeon bruessoworum sp. nov. from Mount Inago, Mozambique (PEM: Port Elizabeth Museum)." httpUri="http://table.plazi.org/id/DF23D1D2FFCFFFEAFF21FEA640E5E9F6" pageId="29" pageNumber="30" tableUuid="DF23D1D2FFCFFFEAFF21FEA640E5E9F6">Table 9</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="DF23D1D2FFCFFFEAFF21FEA640E5E9F6" ID-Table-UUID="DF23D1D2FFCFFFEAFF21FEA640E5E9F6" httpUri="http://table.plazi.org/id/DF23D1D2FFCFFFEAFF21FEA640E5E9F6" pageId="29" pageNumber="30" targetBox="[159,1269,414,922]" targetIsTable="true" targetPageId="29">
<paragraph id="8BE3815AFFCFFFEAFF21FEA640E5E9F6" blockId="29.[151,1436,331,387]" pageId="29" pageNumber="30">
<emphasis id="B9285D48FFCFFFEAFF21FEA6436BE916" bold="true" box="[151,270,332,355]" pageId="29" pageNumber="30">TABLE 9.</emphasis>
Measurements (mm) for the type series of
<taxonomicName id="4C5CFAD9FFCFFFEAFCA2FEA14629E916" box="[788,1100,331,355]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="bruessoworum" status="sp. nov.">
<emphasis id="B9285D48FFCFFFEAFCA2FEA14629E916" box="[788,1100,331,355]" italics="true" pageId="29" pageNumber="30">Rhampholeon bruessoworum</emphasis>
</taxonomicName>
<emphasis id="B9285D48FFCFFFEAFBEDFEA646D3E916" bold="true" box="[1115,1206,332,355]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A21BE033FFCFFFEAFBEDFEA646D3E916" box="[1115,1206,332,355]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Mount Inago, Mozambique (PEM: Port Elizabeth Museum).
</paragraph>
</caption>
<paragraph id="8BE3815AFFCFFFEAFF29FE0C46F2EBEF" pageId="29" pageNumber="30">
<table id="F95C73FAFFCF0008FF29FE744690EBEF" box="[159,1269,414,922]" gridcols="4" gridrows="5" pageId="29" pageNumber="30">
<tr id="356C8318FFCF0008FF29FE744690E989" box="[159,1269,414,508]" gridrow="0" pageId="29" pageNumber="30">
<th id="76BDEA64FFCF0008FF29FE744336E989" box="[159,339,414,508]" gridcol="0" gridrow="0" pageId="29" pageNumber="30">Sex</th>
<th id="76BDEA64FFCF0008FD31FE744174E989" box="[647,785,414,508]" gridcol="1" gridrow="0" pageId="29" pageNumber="30">PEM R20375 Holotype F</th>
<th id="76BDEA64FFCF0008FCC9FE74466FE989" box="[895,1034,414,508]" gridcol="2" gridrow="0" pageId="29" pageNumber="30">PEM R20376 Allotype M</th>
<th id="76BDEA64FFCF0008FBDDFE744690E989" box="[1131,1269,414,508]" gridcol="3" gridrow="0" pageId="29" pageNumber="30">PEM R20374 Paratype F</th>
</tr>
<tr id="356C8318FFCF0008FF29FDFA4690EA0D" box="[159,1269,528,632]" gridrow="1" pageId="29" pageNumber="30">
<th id="76BDEA64FFCF0008FF29FDFA4336EA0D" box="[159,339,528,632]" gridcol="0" gridrow="1" pageId="29" pageNumber="30">Snout-vent length Tail length Total length</th>
<td id="76BDEA64FFCF0008FD31FDFA4174EA0D" box="[647,785,528,632]" gridcol="1" gridrow="1" pageId="29" pageNumber="30">47.3 14.8 62.1</td>
<td id="76BDEA64FFCF0008FCC9FDFA466FEA0D" box="[895,1034,528,632]" gridcol="2" gridrow="1" pageId="29" pageNumber="30">39.0 17.1 56.1</td>
<td id="76BDEA64FFCF0008FBDDFDFA4690EA0D" box="[1131,1269,528,632]" gridcol="3" gridrow="1" pageId="29" pageNumber="30">47.5 15.3 62.7</td>
</tr>
<tr id="356C8318FFCF0008FF29FD664690EA81" box="[159,1269,652,756]" gridrow="2" pageId="29" pageNumber="30">
<th id="76BDEA64FFCF0008FF29FD664336EA81" box="[159,339,652,756]" gridcol="0" gridrow="2" pageId="29" pageNumber="30">Head length Head width Orbit diameter</th>
<td id="76BDEA64FFCF0008FD31FD664174EA81" box="[647,785,652,756]" gridcol="1" gridrow="2" pageId="29" pageNumber="30">15.0 9.2 4.4</td>
<td id="76BDEA64FFCF0008FCC9FD66466FEA81" box="[895,1034,652,756]" gridcol="2" gridrow="2" pageId="29" pageNumber="30">14.5 7.9 4.1</td>
<td id="76BDEA64FFCF0008FBDDFD664690EA81" box="[1131,1269,652,756]" gridcol="3" gridrow="2" pageId="29" pageNumber="30">15.9 9.7 4.9</td>
</tr>
<tr id="356C8318FFCF0008FF29FCE24690EB05" box="[159,1269,776,880]" gridrow="3" pageId="29" pageNumber="30">
<th id="76BDEA64FFCF0008FF29FCE24336EB05" box="[159,339,776,880]" gridcol="0" gridrow="3" pageId="29" pageNumber="30">Snout length Inter-orbital Rostral process</th>
<td id="76BDEA64FFCF0008FD31FCE24174EB05" box="[647,785,776,880]" gridcol="1" gridrow="3" pageId="29" pageNumber="30">5.6 6.3 1.6</td>
<td id="76BDEA64FFCF0008FCC9FCE2466FEB05" box="[895,1034,776,880]" gridcol="2" gridrow="3" pageId="29" pageNumber="30">4.8 6.5 2.1</td>
<td id="76BDEA64FFCF0008FBDDFCE24690EB05" box="[1131,1269,776,880]" gridcol="3" gridrow="3" pageId="29" pageNumber="30">5.6 6.8 1.6</td>
</tr>
<tr id="356C8318FFCF0008FF29FC6E4690EBEF" box="[159,1269,900,922]" gridrow="4" pageId="29" pageNumber="30">
<th id="76BDEA64FFCF0008FF29FC6E4336EBEF" box="[159,339,900,922]" gridcol="0" gridrow="4" pageId="29" pageNumber="30">Casque to Eye</th>
<td id="76BDEA64FFCF0008FD31FC6E4174EBEF" box="[647,785,900,922]" gridcol="1" gridrow="4" pageId="29" pageNumber="30">10.8</td>
<td id="76BDEA64FFCF0008FCC9FC6E466FEBEF" box="[895,1034,900,922]" gridcol="2" gridrow="4" pageId="29" pageNumber="30">10.4</td>
<td id="76BDEA64FFCF0008FBDDFC6E4690EBEF" box="[1131,1269,900,922]" gridcol="3" gridrow="4" pageId="29" pageNumber="30">11.8</td>
</tr>
</table>
</paragraph>
<subSubSection id="C346D2D1FFCFFFEAFF71FC3442BFED2A" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8BE3815AFFCFFFEAFF71FC3442BFED2A" blockId="29.[151,1437,990,2023]" pageId="29" pageNumber="30">
<emphasis id="B9285D48FFCFFFEAFF71FC344324EB82" bold="true" box="[199,321,990,1015]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
The Mt. Inago Pygmy Chameleon is referable to
<taxonomicName id="4C5CFAD9FFCFFFEAFC1EFC344621EB82" box="[936,1092,990,1015]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
<emphasis id="B9285D48FFCFFFEAFC1EFC344621EB82" box="[936,1092,990,1015]" italics="true" pageId="29" pageNumber="30">Rhampholeon</emphasis>
</taxonomicName>
(subgenus
<emphasis id="B9285D48FFCFFFEAFB67FC34470CEB82" box="[1233,1385,990,1015]" italics="true" pageId="29" pageNumber="30">Rhinodigitum</emphasis>
) by possessing an unpigmented parietal peritoneum, claws that are strongly bicuspid, smooth plantar surfaces, and a rostral process. It can be distinguished from most other species in
<taxonomicName id="4C5CFAD9FFCFFFEAFC25FBCC4686EC4A" authority="Rhinodigitum" authorityName="Rhinodigitum" box="[915,1251,1062,1087]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
<emphasis id="B9285D48FFCFFFEAFC25FBCC464AEC4A" box="[915,1071,1062,1087]" italics="true" pageId="29" pageNumber="30">Rhampholeon</emphasis>
(
<emphasis id="B9285D48FFCFFFEAFBF7FBCC46BFEC4A" box="[1089,1242,1062,1087]" italics="true" pageId="29" pageNumber="30">Rhinodigitum</emphasis>
)
</taxonomicName>
by having deep inguinal (absent or indistinct in
<emphasis id="B9285D48FFCFFFEAFE4CFBA04644EC16" box="[506,1057,1098,1123]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="4C5CFAD9FFCFFFEAFE4CFBA040C6EC16" box="[506,675,1098,1123]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="boulengeri">Rh. boulengeri</taxonomicName>
,
<taxonomicName id="4C5CFAD9FFCFFFEAFD18FBA0413DEC16" box="[686,856,1098,1123]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="nchisiensis">Rh. nchisiensis</taxonomicName>
,
<taxonomicName id="4C5CFAD9FFCFFFEAFCD5FBA04644EC16" box="[867,1057,1098,1123]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="uluguruensis">Rh. uluguruensis</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C5CFAD9FFCFFFEAFBEBFBA046BDEC16" box="[1117,1240,1098,1123]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="moyeri">
<emphasis id="B9285D48FFCFFFEAFBEBFBA046BDEC16" box="[1117,1240,1098,1123]" italics="true" pageId="29" pageNumber="30">Rh. moyeri</emphasis>
</taxonomicName>
) and axillary pits (also absent in
<taxonomicName id="4C5CFAD9FFCFFFEAFEF7FB84438FECF2" box="[321,490,1134,1159]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="nchisiensis">
<emphasis id="B9285D48FFCFFFEAFEF7FB84438FECF2" box="[321,490,1134,1159]" italics="true" pageId="29" pageNumber="30">Rh. nchisiensis</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="4C5CFAD9FFCFFFEAFD1EFB84415AECF2" box="[680,831,1134,1159]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="platyceps">
<emphasis id="B9285D48FFCFFFEAFD1EFB84415AECF2" box="[680,831,1134,1159]" italics="true" pageId="29" pageNumber="30">Rh. platyceps</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5CFAD9FFCFFFEAFCC1FB844671ECF3" box="[887,1044,1134,1159]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="maspictus" status="sp. nov.">
<emphasis id="B9285D48FFCFFFEAFCC1FB844671ECF3" box="[887,1044,1134,1159]" italics="true" pageId="29" pageNumber="30">Rh. maspictus</emphasis>
</taxonomicName>
<emphasis id="B9285D48FFCFFFEAFBAAFB844613ECF2" bold="true" box="[1052,1142,1134,1159]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A21BE033FFCFFFEAFBAAFB844613ECF2" box="[1052,1142,1134,1159]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in its small size (&lt;
<quantity id="4CA42CBFFFCFFFEAFAFAFB8447F9ECF3" box="[1356,1436,1134,1159]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="30" unit="mm" value="50.0">50 mm</quantity>
SVL), relatively large rostral process in males, and weakly developed crenulations along the dorsal crest. It differs from
<taxonomicName id="4C5CFAD9FFCFFFEAFF6AFB5C43D6ECBA" box="[220,435,1206,1231]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="chapmanorum">
<emphasis id="B9285D48FFCFFFEAFF6AFB5C43D6ECBA" box="[220,435,1206,1231]" italics="true" pageId="29" pageNumber="30">Rh. chapmanorum</emphasis>
</taxonomicName>
in having a relatively large rostral process in males (small in both sexes in
<taxonomicName id="4C5CFAD9FFCFFFEAFAC0FB5C435CEC86" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="chapmanorum">
<emphasis id="B9285D48FFCFFFEAFAC0FB5C435CEC86" italics="true" pageId="29" pageNumber="30">Rh. chapmanorum</emphasis>
</taxonomicName>
), and from all other members of the
<taxonomicName id="4C5CFAD9FFCFFFEAFD68FB304113EC86" box="[734,886,1242,1267]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="platyceps">
<emphasis id="B9285D48FFCFFFEAFD68FB304113EC86" box="[734,886,1242,1267]" italics="true" pageId="29" pageNumber="30">Rh. platyceps</emphasis>
</taxonomicName>
complex in
<collectingCountry id="F34BC1CAFFCFFFEAFBB3FB3046FCEC86" box="[1029,1177,1242,1267]" name="Mozambique" pageId="29" pageNumber="30">Mozambique</collectingCountry>
(i.e.
<taxonomicName id="4C5CFAD9FFCFFFEAFB62FB304717EC86" box="[1236,1394,1242,1267]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="maspictus" status="sp. nov.">
<emphasis id="B9285D48FFCFFFEAFB62FB304717EC86" box="[1236,1394,1242,1267]" italics="true" pageId="29" pageNumber="30">Rh. maspictus</emphasis>
</taxonomicName>
<emphasis id="B9285D48FFCFFFEAFACCFB3042ADED62" bold="true" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A21BE033FFCFFFEAFACCFB3042ADED62" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C5CFAD9FFCFFFEAFF61FB1443E8ED62" box="[215,397,1278,1303]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="nebulauctor" status="sp. nov.">
<emphasis id="B9285D48FFCFFFEAFF61FB1443E8ED62" box="[215,397,1278,1303]" italics="true" pageId="29" pageNumber="30">Rh. nebulauctor</emphasis>
</taxonomicName>
<emphasis id="B9285D48FFCFFFEAFE23FB144394ED62" bold="true" box="[405,497,1278,1303]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A21BE033FFCFFFEAFE23FB144394ED62" box="[405,497,1278,1303]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C5CFAD9FFCFFFEAFD9CFB1440CEED62" box="[554,683,1278,1303]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="tilburyi" status="sp. nov.">
<emphasis id="B9285D48FFCFFFEAFD9CFB1440CEED62" box="[554,683,1278,1303]" italics="true" pageId="29" pageNumber="30">Rh. tilburyi</emphasis>
</taxonomicName>
<emphasis id="B9285D48FFCFFFEAFD05FB14416BED62" bold="true" box="[691,782,1278,1303]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A21BE033FFCFFFEAFD05FB14416BED62" box="[691,782,1278,1303]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) in having a relatively longer tail in both sexes. From all other
<taxonomicName id="4C5CFAD9FFCFFFEAFF6CFAC84313ED4E" box="[218,374,1314,1339]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
<emphasis id="B9285D48FFCFFFEAFF6CFAC84313ED4E" box="[218,374,1314,1339]" italics="true" pageId="29" pageNumber="30">Rhampholeon</emphasis>
</taxonomicName>
it is also genetically well differentiated, and all chameleons examined form a monophyletic clade.
</paragraph>
</subSubSection>
<subSubSection id="C346D2D1FFCFFFE9FF71FA80401EEC09" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph id="8BE3815AFFCFFFEAFF71FA8046E4EDF6" blockId="29.[151,1437,990,2023]" box="[199,1153,1386,1411]" pageId="29" pageNumber="30">
<emphasis id="B9285D48FFCFFFEAFF71FA804386EDF6" bold="true" box="[199,483,1386,1411]" pageId="29" pageNumber="30">
Description of
<typeStatus id="54E73FF8FFCFFFEAFEC0FA804386EDF6" box="[374,483,1386,1411]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
. Adult female, viscera exposed by a single ventral incision.
</paragraph>
<paragraph id="8BE3815AFFCFFFEAFF71FA644344EF46" blockId="29.[151,1437,990,2023]" pageId="29" pageNumber="30">
Head: Dorsum of head flattened, with no upward flexure of the snout; casque flat, edged with weakly-defined lateral crests that are mainly restricted to the posterior region of the casque; temporal crest weakly-developed, comprising a single, interrupted row of large tubercles; parietal crest almost absent, composed of a few enlarged tubercles in the mid-line; supraorbital ridges reduced to a few scattered enlarged scales but with a very small multiscaled process forming a soft horn at each end of the inter-orbital ridge that passes across the crown and is composed of 10 small granular tubercles, and that demarcate the posterior edge of a slight frontal depression; inferior orbital rim with 4 (right) and 5 (left) enlarged tubercles; snout bordered on each side by moderately developed rostral crests, that fuse together at the tip of the snout which is adorned with a very small, flattened rostral process (
<quantity id="4CA42CBFFFCFFFEAFEFFF94443C4EEB2" box="[329,417,1710,1735]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="29" pageNumber="30" unit="mm" value="1.6">1.6 mm</quantity>
) which underneath is only slightly free from the rostral, and is four small tubercular scales long and four tubercular scales wide at its base; nares opening posterio-ventrally; no gular or mental appendages; scales on throat homogenous, more conical but smaller than those on crown of head and subequal in size to those on the belly.
</paragraph>
<paragraph id="8BE3815AFFCFFFE9FF71F8D44385E88D" blockId="29.[151,1437,990,2023]" lastBlockId="30.[151,1437,151,1437]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">Body: Dorsal crest very weakly developed, reduced to 9 crenulations of enlarged, but not obviously spinose scales; crenulations most strongly developed over mid-body, reduced in size over on neck and which are present on the tail in more reduced form; deep axillary and inguinal pits are present; flank scalation heterogeneous, composed of small, stellate granules with few scattered, enlarged spinose tubercles, the largest at the shoulder and in two clusters; chest, belly and lower surface of tail smooth; limb scalation more tubercular, with a few enlarged, spinose tubercles on the forearms; claws strongly bicuspid; accessory planter spines on the soles of the fore and hind feet are present, but reduced to very small, soft, spinose scales at the base of the claws; tail flattened laterally, flexing downward on the distal third.</paragraph>
<paragraph id="8BE3815AFFCCFFE9FF71FEEE43A8E9FD" blockId="30.[151,1437,151,1437]" pageId="30" pageNumber="31">
Colour in life (based on two images of
<typeStatus id="54E73FF8FFCCFFE9FD20FEEE409EE968" box="[662,763,260,285]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
, JB;
<figureCitation id="13679DDFFFCCFFE9FC8EFEEE411BE969" box="[824,894,260,285]" captionStart="FIGURE 9" captionStartId="28.[151,250,1953,1975]" captionTargetBox="[177,1407,214,1913]" captionTargetId="figure@28.[164,1423,193,1932]" captionTargetPageId="28" captionText="FIGURE 9. Rhampholeon bruessoworum sp. nov.: A—Holotype; B—Allotype; C—Habitat on Mt. Inago (J. Bayliss)." httpUri="https://zenodo.org/record/286220/files/figure.png" pageId="30" pageNumber="31">Fig. 9</figureCitation>
A): Mid-body mottled brown with two narrow, oblique purple-brown lateral stripes; dorsal surface of fore-body, neck, top of head and upper surfaces of limbs darker brown; Throat lightly mottled cream extending onto chin and labials, which have a light yellow flush; belly and base of tail pale brown.
</paragraph>
<paragraph id="8BE3815AFFCCFFE9FF71FE7E4197E9A5" blockId="30.[151,1437,151,1437]" pageId="30" pageNumber="31">Colour in preservative: Body mottled brown with two narrow oblique reddish-brown bars on mid-flanks; lower surface of neck, belly, base of tail, soles of feet, and lower limbs pale brown.</paragraph>
<paragraph id="8BE3815AFFCCFFE9FF71FE364007EA6D" blockId="30.[151,1437,151,1437]" pageId="30" pageNumber="31">
<emphasis id="B9285D48FFCCFFE9FF71FE3643BFE980" bold="true" box="[199,474,476,501]" pageId="30" pageNumber="31">Description of Allotype</emphasis>
(as for
<typeStatus id="54E73FF8FFCCFFE9FD99FE3640F1E980" box="[559,660,476,501]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
, unless noted): Adult male, incision along base of tail with hemipenal muscle removed; hemipenes not everted.
</paragraph>
<paragraph id="8BE3815AFFCCFFE9FF71FDCE434CEAF0" blockId="30.[151,1437,151,1437]" pageId="30" pageNumber="31">Head: Parietal crest almost absent, composed of a four enlarged tubercles in the mid-line; five enlarged tubercles on both sides of the inferior orbital rim; rostral process six small tubercular scales long and four tubercles wide at base.</paragraph>
<paragraph id="8BE3815AFFCCFFE9FF71FD6543BAEA85" blockId="30.[151,1437,151,1437]" pageId="30" pageNumber="31">
Body: Dorsal crest very weakly developed, reduced to nine crenulations of enlarged but not obviously spinose scales, subequal in development to those of the female
<typeStatus id="54E73FF8FFCCFFE9FC8BFD5E41C6EAB8" box="[829,931,692,717]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
; tail relatively long (30.5% SVL), with a prominent hemipeneal bulge.
</paragraph>
<paragraph id="8BE3815AFFCCFFE9FF71FD164283EB28" blockId="30.[151,1437,151,1437]" pageId="30" pageNumber="31">
Colour in life (based on two images of allotype;
<figureCitation id="13679DDFFFCCFFE9FD59FD164156EB61" box="[751,819,764,789]" captionStart="FIGURE 9" captionStartId="28.[151,250,1953,1975]" captionTargetBox="[177,1407,214,1913]" captionTargetId="figure@28.[164,1423,193,1932]" captionTargetPageId="28" captionText="FIGURE 9. Rhampholeon bruessoworum sp. nov.: A—Holotype; B—Allotype; C—Habitat on Mt. Inago (J. Bayliss)." httpUri="https://zenodo.org/record/286220/files/figure.png" pageId="30" pageNumber="31">Fig. 9</figureCitation>
B): Mid-body mottled brown with small orange brown blotches along dorsal crest, separated by 23 crenulations; throat, belly and lower surfaces of limbs and tail pale brown.
</paragraph>
<paragraph id="8BE3815AFFCCFFE9FF71FC8D43DDEBD0" blockId="30.[151,1437,151,1437]" pageId="30" pageNumber="31">Colour in preservative: After preservation body pale brown with vestiges of two narrow oblique stripes on the flanks; pale brown below.</paragraph>
<paragraph id="8BE3815AFFCCFFE9FF71FC454173EC65" blockId="30.[151,1437,151,1437]" pageId="30" pageNumber="31">
<emphasis id="B9285D48FFCCFFE9FF71FC4543CDEBBD" bold="true" box="[199,424,943,968]" pageId="30" pageNumber="31">
<typeStatus id="54E73FF8FFCCFFE9FF71FC454351EBBD" box="[199,308,943,968]" pageId="30" pageNumber="31" type="paratype">Paratype</typeStatus>
variation
</emphasis>
(as for
<typeStatus id="54E73FF8FFCCFFE9FE48FC454006EBBD" box="[510,611,943,968]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
, unless noted): Adult female with a large ventral incision. The soft horn on the supraorbital ridge is very small and hardly protrudes; only four enlarged tubercles on inferior orbital rim; plantar spines very reduced to small, soft, spinose scales.
</paragraph>
<paragraph id="8BE3815AFFCCFFE9FF71FBF6401EEC09" blockId="30.[151,1437,151,1437]" pageId="30" pageNumber="31">
<emphasis id="B9285D48FFCCFFE9FF71FBF6429BEC40" bold="true" box="[199,254,1052,1077]" pageId="30" pageNumber="31">Size.</emphasis>
Presumably a small species, as all three specimens in the
<typeStatus id="54E73FF8FFCCFFE9FC2DFBF741AEEC40" box="[923,971,1053,1077]" pageId="30" pageNumber="31">type</typeStatus>
series were sexually mature, with turgid testes or developing ova. Largest male - PEM R20376 (allotype) 39.0 + 17.1 =
<quantity id="4CA42CBFFFCCFFE9FB86FBD546FCEC22" box="[1072,1177,1087,1112]" metricMagnitude="-2" metricUnit="m" metricValue="5.61" pageId="30" pageNumber="31" unit="mm" value="56.1">56.1 mm</quantity>
; largest female—PEM R20374 (
<typeStatus id="54E73FF8FFCCFFE9FF48FB8E4301EC08" box="[254,356,1124,1149]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
) 47.5 + 15.3 =
<quantity id="4CA42CBFFFCCFFE9FDA6FB8E4012EC09" box="[528,631,1124,1149]" metricMagnitude="-2" metricUnit="m" metricValue="6.27" pageId="30" pageNumber="31" unit="mm" value="62.7">62.7 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C346D2D1FFCCFFE9FF71FB6D43E4EDE9" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BE3815AFFCCFFE9FF71FB6D4741ECD5" blockId="30.[151,1437,151,1437]" box="[199,1316,1159,1185]" pageId="30" pageNumber="31">
<emphasis id="B9285D48FFCCFFE9FF71FB6D4305ECD5" bold="true" box="[199,352,1159,1184]" pageId="30" pageNumber="31">Distribution.</emphasis>
Restricted to the
<typeStatus id="54E73FF8FFCCFFE9FD93FB634030ECD4" box="[549,597,1161,1185]" pageId="30" pageNumber="31">type</typeStatus>
locality; Mt. Inago, Zambézia Province, northern
<collectingCountry id="F34BC1CAFFCCFFE9FB3FFB6D4744ECD5" box="[1161,1313,1159,1184]" name="Mozambique" pageId="30" pageNumber="31">Mozambique</collectingCountry>
.
</paragraph>
<paragraph id="8BE3815AFFCCFFE9FF71FB4643E4EDE9" blockId="30.[151,1437,151,1437]" pageId="30" pageNumber="31">
<emphasis id="B9285D48FFCCFFE9FF71FB46434FECB0" bold="true" box="[199,298,1196,1221]" pageId="30" pageNumber="31">Habitat.</emphasis>
The Inago Massif shows habitat zonation, and is surrounded by
<taxonomicName id="4C5CFAD9FFCCFFE9FBBCFB4646FBECB0" box="[1034,1182,1196,1221]" class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" order="Fabales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="B9285D48FFCCFFE9FBBCFB4646FBECB0" box="[1034,1182,1196,1221]" italics="true" pageId="30" pageNumber="31">Brachystegia</emphasis>
</taxonomicName>
woodland at the base. The
<typeStatus id="54E73FF8FFCCFFE9FF7FFB3B429CEC9C" box="[201,249,1233,1257]" pageId="30" pageNumber="31">type</typeStatus>
series were collected at night in mid-altitude (~
<quantity id="4CA42CBFFFCCFFE9FCA6FB25410DEC9D" box="[784,872,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="31" unit="m" value="1500.0">1500 m</quantity>
) evergreen forest at the base of a granite inselberg (
<figureCitation id="13679DDFFFCCFFE9FF29FB1E4280ED79" box="[159,229,1268,1293]" captionStart="FIGURE 9" captionStartId="28.[151,250,1953,1975]" captionTargetBox="[177,1407,214,1913]" captionTargetId="figure@28.[164,1423,193,1932]" captionTargetPageId="28" captionText="FIGURE 9. Rhampholeon bruessoworum sp. nov.: A—Holotype; B—Allotype; C—Habitat on Mt. Inago (J. Bayliss)." httpUri="https://zenodo.org/record/286220/files/figure.png" pageId="30" pageNumber="31">Fig. 9</figureCitation>
C). This forest
<typeStatus id="54E73FF8FFCCFFE9FE21FB1F43A2ED78" box="[407,455,1269,1293]" pageId="30" pageNumber="31">type</typeStatus>
comprised relatively large trees between
<quantity id="4CA42CBFFFCCFFE9FC1CFB1E4669ED79" box="[938,1036,1268,1292]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="30" pageNumber="31" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">20-30 m</quantity>
high, with the upper canopy layer composed of species such as
<emphasis id="B9285D48FFCCFFE9FE6AFAF341A5ED45" box="[476,960,1303,1328]" italics="true" pageId="30" pageNumber="31">
<taxonomicName id="4C5CFAD9FFCCFFE9FE6AFAF3415CED45" authority="Schefflera" authorityName="Schefflera" box="[476,825,1303,1328]" class="Magnoliopsida" family="Putranjivaceae" genus="Drypetes" kingdom="Plantae" order="Malpighiales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="natalensis">Drypetes natalensis, Schefflera</taxonomicName>
umbellifera
</emphasis>
and
<taxonomicName id="4C5CFAD9FFCCFFE9FC40FAF34683ED45" box="[1014,1254,1303,1328]" class="Aves" family="Vangidae" genus="Newtonia" kingdom="Animalia" order="Passeriformes" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="buchananii">
<emphasis id="B9285D48FFCCFFE9FC40FAF34683ED45" box="[1014,1254,1303,1328]" italics="true" pageId="30" pageNumber="31">Newtonia buchananii</emphasis>
</taxonomicName>
, whilst the midcanopy layer (&lt;
<quantity id="4CA42CBFFFCCFFE9FED7FAD643C7ED21" box="[353,418,1340,1364]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="31" unit="m" value="20.0">20 m</quantity>
) was dominated by
<emphasis id="B9285D48FFCCFFE9FD2BFAD6475CED20" box="[669,1337,1340,1365]" italics="true" pageId="30" pageNumber="31">
<taxonomicName id="4C5CFAD9FFCCFFE9FD2BFAD6461DED20" authority="Myrianthus" authorityName="Myrianthus" box="[669,1144,1340,1365]" class="Magnoliopsida" family="Sapotaceae" genus="Chrysophyllum" kingdom="Plantae" order="Ericales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="gorungosum">Chrysophyllum gorungosum, Myrianthus</taxonomicName>
holstii, bridelia
</emphasis>
sp. and
<taxonomicName id="4C5CFAD9FFCCFFE9FF21FA8A429EED02" box="[151,251,1376,1399]" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="B9285D48FFCCFFE9FF21FA8A429EED02" box="[151,251,1376,1399]" italics="true" pageId="30" pageNumber="31">Garcinia</emphasis>
</taxonomicName>
sp. These remnant forest patches are greatly reduced in size (&lt;10 ha), fragmented, and highly threatened (Bayliss
<emphasis id="B9285D48FFCCFFE9FF4CFA6F434EEDE8" box="[250,299,1412,1437]" italics="true" pageId="30" pageNumber="31">et al</emphasis>
. 2010).
</paragraph>
</subSubSection>
</treatment>
</document>