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<document id="3F5D0049F0B6E04AB2E990CEB882A1D3" ID-DOI="10.11646/zootaxa.3641.3.1" ID-GBIF-Dataset="0934da3b-4a76-425c-b481-32fee1b6654a" ID-ISSN="1175-5326" ID-Zenodo-Dep="223165" ID-ZooBank="8E36BE56-61EA-4FB9-BEF8-BCDA043EA6DE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460595293200" checkinUser="plazi" docAuthor="Yamamoto, Shûhei &amp; Maruyama, Munetoshi" docDate="2013" docId="03D94D1BFFD7FFFB53C4FF41C08F8F47" docLanguage="en" docName="zt03641p222.pdf" docOrigin="Zootaxa 3641 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aleochara (Coprochara) verna Say 1833" docType="treatment" docVersion="7" lastPageNumber="213" masterDocId="FFE03563FFDEFFF75353FF9FC5128D18" masterDocTitle="Revision of the Subgenus Coprochara Mulsant &amp; Rey of the Genus Aleochara Gravenhorst from Japan (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="222" masterPageNumber="201" pageNumber="210" updateTime="1698326675319" updateUser="plazi">
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<mods:title id="69B9CCBE1D58C8CB07BF17B96A654696">Revision of the Subgenus Coprochara Mulsant &amp; Rey of the Genus Aleochara Gravenhorst from Japan (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="BBE62626C0D13EBC7A988E376795F545">Yamamoto, Shûhei</mods:namePart>
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<mods:namePart id="A7B9358386055ACF8863796DE90D8550">Maruyama, Munetoshi</mods:namePart>
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<treatment id="03D94D1BFFD7FFFB53C4FF41C08F8F47" ID-DOI="http://doi.org/10.5281/zenodo.6149269" ID-GBIF-Taxon="119566534" ID-Zenodo-Dep="6149269" LSID="urn:lsid:plazi:treatment:03D94D1BFFD7FFFB53C4FF41C08F8F47" httpUri="http://treatment.plazi.org/id/03D94D1BFFD7FFFB53C4FF41C08F8F47" lastPageId="12" lastPageNumber="213" pageId="9" pageNumber="210">
<subSubSection id="C36AAF86FFD7FFFE53C4FF41C1C08C84" pageId="9" pageNumber="210" type="nomenclature">
<paragraph id="8BCFFC0DFFD7FFFE53C4FF41C7958DE0" blockId="9.[151,647,222,281]" box="[151,647,222,248]" pageId="9" pageNumber="210">
<heading id="D0874B61FFD7FFFE53C4FF41C7958DE0" bold="true" box="[151,647,222,248]" fontSize="11" level="1" pageId="9" pageNumber="210" reason="1">
<taxonomicName id="4C70878EFFD7FFFE53C4FF41C7958DE0" ID-CoL="BMYL" authority="Say, 1833" authorityName="Say" authorityYear="1833" box="[151,647,222,248]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="verna" subGenus="Coprochara">
<emphasis id="B904201FFFD7FFFE53C4FF41C7958DE0" bold="true" box="[151,647,222,248]" pageId="9" pageNumber="210">
<emphasis id="B904201FFFD7FFFE53C4FF41C4068DE0" bold="true" box="[151,276,222,248]" italics="true" pageId="9" pageNumber="210">Aleochara</emphasis>
(
<emphasis id="B904201FFFD7FFFE5276FF41C4A48DE0" bold="true" box="[293,438,222,248]" italics="true" pageId="9" pageNumber="210">Coprochara</emphasis>
)
<emphasis id="B904201FFFD7FFFE5294FF41C7198DE0" bold="true" box="[455,523,222,248]" italics="true" pageId="9" pageNumber="210">verna</emphasis>
Say, 1833
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FE9EC4D18C01" blockId="9.[151,647,222,281]" box="[151,451,257,281]" pageId="9" pageNumber="210">
<figureCitation id="134BE088FFD7FFFE53C4FE9EC5F18C01" box="[151,227,257,281]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[278,1306,191,1972]" captionTargetId="figure@2.[278,1308,161,1975]" captionTargetPageId="2" captionText="FIGURES 1 4. Japanese species of Aleochara (Coprochara) in habitus. 1. Aleochara (Coprochara) verna; 2. A. (C.) binotata; 3 4. A. (C.) squalithorax: gray type (Fig. 3) and yellow type (Fig. 4)." httpUri="https://zenodo.org/record/223166/files/figure.png" pageId="9" pageNumber="210">Figs. 1</figureCitation>
,
<figureCitation id="134BE088FFD7FFFE53A2FE9EC40F8C01" box="[241,285,257,281]" captionStart="FIGURES 5 13" captionStartId="3.[151,264,1073,1096]" captionTargetBox="[163,1424,194,1036]" captionTargetId="figure@3.[151,1436,171,1052]" captionTargetPageId="3" captionText="FIGURES 5 13. Japanese species of Aleochara (Coprochara). 5 7. Aleochara (Coprochara) verna; 8 10. A. (C.) binotata; 11 13. A. (C.) squalithorax. (5, 8, 11: elytra; 6, 9, 12: intercoxal process of metaventrite; 7, 10, 13: mesoventrite and metaventrite)." httpUri="https://zenodo.org/record/223167/files/figure.png" pageId="9" pageNumber="210">5, 7</figureCitation>
,
<figureCitation id="134BE088FFD7FFFE527AFE9EC4638C01" box="[297,369,257,281]" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="9" pageNumber="210">14, 21</figureCitation>
,
<figureCitation id="134BE088FFD7FFFE522EFE9EC48E8C01" box="[381,412,257,281]" captionStart="FIGURES 42 43" captionStartId="17.[151,264,1257,1280]" captionTargetBox="[157,1430,811,1230]" captionTargetId="figure@17.[151,1436,807,1235]" captionTargetPageId="17" captionText="FIGURES 42 43. Habitats of the subgenus Coprochara of the genus Aleochara in Japan. 93. the summit of Sobo-san (located in boundaries between Ôita and Miyazaki Prefectures; only observed; A. (C.) verna: arrow); 43. Watari (Fukutsu, Fukuoka Prefecture; A. (C.) squalithorax: arrow)." httpUri="https://zenodo.org/record/223172/files/figure.png" pageId="9" pageNumber="210">42</figureCitation>
,
<figureCitation id="134BE088FFD7FFFE52F4FE9EC4D18C01" box="[423,451,257,281]" captionStart="FIGURES 44 45" captionStartId="18.[151,264,1417,1440]" captionTargetBox="[172,1408,196,1380]" captionTargetId="figure@18.[151,1436,171,1396]" captionTargetPageId="18" captionText="FIGURES 44 45. Collecting localities of Aleochara (Coprochara) spp. based on specimens examined in Japan. 44. Aleochara (Coprochara) verna (Indicated by red circle) and A. (C.) binotata (Indicated by yellow square); 45. A. (C.) squalithorax (Indicated by blue triangle)." httpUri="https://zenodo.org/record/223173/files/figure.png" pageId="9" pageNumber="210">44</figureCitation>
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FED6C1C08C84" blockId="9.[151,1437,329,781]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53C4FED6C4C78C47" box="[151,469,329,351]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="verna" subGenus="Coprochara">
<emphasis id="B904201FFFD7FFFE53C4FED6C4128C47" box="[151,256,329,351]" italics="true" pageId="9" pageNumber="210">Aleochara</emphasis>
(
<emphasis id="B904201FFFD7FFFE5243FED6C4998C47" box="[272,395,329,351]" italics="true" pageId="9" pageNumber="210">Coprochara</emphasis>
)
<emphasis id="B904201FFFD7FFFE52C8FED5C4C78C47" box="[411,469,330,351]" italics="true" pageId="9" pageNumber="210">verna</emphasis>
</taxonomicName>
; Say, 1833: 58 (original description); Klimaszewski, 1984: 22; Lohse, 1986: 95; Lohse, 1989: 239; Welch, 1997: 3, 14; Maus, 1998: 87, 96; Maus
<emphasis id="B904201FFFD7FFFE51B6FEF6C6018C66" box="[741,787,360,382]" italics="true" pageId="9" pageNumber="210">et al</emphasis>
., 1998: 241; Maus &amp; Ashe, 1998 (online); Naomi
<emphasis id="B904201FFFD7FFFE5673FEF6C05C8C66" box="[1312,1358,360,382]" italics="true" pageId="9" pageNumber="210">et al</emphasis>
., 2000: 107; Maus
<emphasis id="B904201FFFD7FFFE526AFE17C4768C85" box="[313,356,391,413]" italics="true" pageId="9" pageNumber="210">et al</emphasis>
., 2001: 205; Smetana, 2004: 356; Gouix &amp; Klimaszewski, 2007: 25; Park
<emphasis id="B904201FFFD7FFFE570AFE17C1948C85" box="[1113,1158,391,413]" italics="true" pageId="9" pageNumber="210">et al</emphasis>
., 2011.
</paragraph>
</subSubSection>
<subSubSection id="C36AAF86FFD7FFFE53C4FE3AC6F88E14" pageId="9" pageNumber="210" type="description">
<paragraph id="8BCFFC0DFFD7FFFE53C4FE3AC7348CA2" blockId="9.[151,1437,329,781]" box="[151,550,421,443]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53C4FE3AC7308CA2" authority="Sachse, 1852: 117" authorityName="Sachse" authorityPageNumber="117" authorityYear="1852" box="[151,546,421,443]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="languida">
<emphasis id="B904201FFFD7FFFE53C4FE3AC4728CA3" box="[151,352,421,443]" italics="true" pageId="9" pageNumber="210">Aleochara languida</emphasis>
Sachse, 1852: 117
</taxonomicName>
.
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FE5BC7668CC1" blockId="9.[151,1437,329,781]" box="[151,628,451,474]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53C4FE5BC7628CC1" authority="Mulsant &amp; Rey, 1874: 159" authorityName="Mulsant &amp; Rey" authorityPageNumber="159" authorityYear="1874" box="[151,624,451,474]" class="Insecta" family="Staphylinidae" genus="Baryodma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="pauxilla">
<emphasis id="B904201FFFD7FFFE53C4FE5BC44B8CC2" box="[151,345,452,474]" italics="true" pageId="9" pageNumber="210">Baryodma pauxilla</emphasis>
Mulsant &amp; Rey, 1874: 159
</taxonomicName>
.
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FE7DC71D8CE0" blockId="9.[151,1437,329,781]" box="[151,527,482,504]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53C4FE7DC7198CE0" authority="Sharp, 1883: 148" authorityName="Sharp" authorityPageNumber="148" authorityYear="1883" box="[151,523,482,504]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="B904201FFFD7FFFE53C4FE7DC4468CE0" box="[151,340,482,504]" italics="true" pageId="9" pageNumber="210">Aleochara alticola</emphasis>
Sharp, 1883: 148
</taxonomicName>
.
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FD9EC79F8F0F" blockId="9.[151,1437,329,781]" box="[151,653,513,535]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53C4FD9EC7178F0F" authority="Casey, 1906: 161" authorityName="Casey" authorityPageNumber="161" authorityYear="1906" box="[151,517,513,535]" class="Insecta" family="Staphylinidae" genus="Baryodma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="B904201FFFD7FFFE53C4FD9EC45F8F0F" box="[151,333,513,535]" italics="true" pageId="9" pageNumber="210">Baryodma minuta</emphasis>
Casey, 1906: 161
</taxonomicName>
. [homonym].
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FDBFC4E78F2E" blockId="9.[151,1437,329,781]" box="[151,501,544,566]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53C4FDBFC4E38F2E" authority="Casey, 1911: 6" authorityName="Casey" authorityPageNumber="6" authorityYear="1911" box="[151,497,544,566]" class="Insecta" family="Staphylinidae" genus="Baryodma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B904201FFFD7FFFE53C4FDBFC4468F2E" box="[151,340,544,566]" italics="true" pageId="9" pageNumber="210">Baryodma pumilio</emphasis>
Casey, 1911: 6
</taxonomicName>
.
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FDA0C4EA8F4C" blockId="9.[151,1437,329,781]" box="[151,504,574,597]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53C4FDA0C4E68F4C" authority="Casey, 1911: 6" authorityName="Casey" authorityPageNumber="6" authorityYear="1911" box="[151,500,574,597]" class="Insecta" family="Staphylinidae" genus="Baryodma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="tolerata">
<emphasis id="B904201FFFD7FFFE53C4FDA0C4448F4D" box="[151,342,575,597]" italics="true" pageId="9" pageNumber="210">Baryodma tolerata</emphasis>
Casey, 1911: 6
</taxonomicName>
.
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FDC2C7EF8F6B" blockId="9.[151,1437,329,781]" box="[151,765,605,627]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53C4FDC2C7308F6B" authority="Muona, 1977: 16" authorityName="Muona" authorityPageNumber="16" authorityYear="1977" box="[151,546,605,627]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="tecumsehi">
<emphasis id="B904201FFFD7FFFE53C4FDC2C4798F6B" box="[151,363,605,627]" italics="true" pageId="9" pageNumber="210">Aleochara tecumsehi</emphasis>
Muona, 1977: 16
</taxonomicName>
. [replacement name].
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FDE3C7F88F89" blockId="9.[151,1437,329,781]" box="[151,746,635,658]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53C4FDE3C71C8F89" authority="Likovsky, 1984: 8" authorityName="Likovsky" authorityPageNumber="8" authorityYear="1984" box="[151,526,635,658]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="tanumi">
<emphasis id="B904201FFFD7FFFE53C4FDE3C45F8F8A" box="[151,333,636,658]" italics="true" pageId="9" pageNumber="210">Aleochara tanumi</emphasis>
Likovský, 1984: 8
</taxonomicName>
. [replacement name].
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FD05C7F68FA8" blockId="9.[151,1437,329,781]" box="[151,740,666,688]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53C4FD05C71A8FA8" authority="Likovsky, 1984: 8" authorityName="Likovsky" authorityPageNumber="8" authorityYear="1984" box="[151,520,666,688]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="cedari">
<emphasis id="B904201FFFD7FFFE53C4FD05C4558FA8" box="[151,327,666,688]" italics="true" pageId="9" pageNumber="210">Aleochara cedari</emphasis>
Likovský, 1984: 8
</taxonomicName>
. [replacement name].
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FD25C74F8FF6" blockId="9.[151,1437,329,781]" pageId="9" pageNumber="210">
<taxonomicName id="4C70878EFFD7FFFE53F1FD26C4968FD7" box="[162,388,697,719]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="bipustulata">
<emphasis id="B904201FFFD7FFFE53F1FD26C4968FD7" box="[162,388,697,719]" italics="true" pageId="9" pageNumber="210">Aleochara bipustulata</emphasis>
</taxonomicName>
”; Sharp, 1888: 282 (as
<taxonomicName id="4C70878EFFD7FFFE5126FD26C6B48FD7" authority="Gravenhorst" authorityName="Gravenhorst" box="[629,934,697,719]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="B904201FFFD7FFFE5126FD26C6328FD7" box="[629,800,697,719]" italics="true" pageId="9" pageNumber="210">Aleochara nitida</emphasis>
Gravenhorst
</taxonomicName>
, a synonym of
<taxonomicName id="4C70878EFFD7FFFE5712FD25C1DD8FD7" box="[1089,1231,697,719]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="bipustulata">
<emphasis id="B904201FFFD7FFFE5712FD25C1DD8FD7" box="[1089,1231,697,719]" italics="true" pageId="9" pageNumber="210">A. bipustulata</emphasis>
</taxonomicName>
); Adachi, 1957: 34; Naomi, 1989: 280; Smetana, 2004: 355.
</paragraph>
<paragraph id="8BCFFC0DFFD7FFFE53C4FD68C6F88E14" blockId="9.[151,1437,329,781]" box="[151,1002,758,781]" pageId="9" pageNumber="210">See Klimaszewski (1984) and Gouix &amp; Klimaszewski (2007) for further references.</paragraph>
</subSubSection>
<subSubSection id="C36AAF86FFD7FFFC53C4FCA6C0868C7A" lastPageId="11" lastPageNumber="212" pageId="9" pageNumber="210" type="materials_examined">
<paragraph id="8BCFFC0DFFD7FFFD53C4FCA6C1D18917" blockId="9.[151,1437,825,2037]" lastBlockId="10.[151,1437,151,2006]" lastPageId="10" lastPageNumber="211" pageId="9" pageNumber="210">
<emphasis id="B904201FFFD7FFFE53C4FCA6C4998E4A" bold="true" box="[151,395,825,850]" pageId="9" pageNumber="210">Materials examined.</emphasis>
<collectingCountry id="F367BC9DFFD7FFFE52C1FCA5C4F98E4A" box="[402,491,826,850]" name="Japan" pageId="9" pageNumber="210">JAPAN</collectingCountry>
: [
<emphasis id="B904201FFFD7FFFE52ACFCA6C7678E4A" bold="true" box="[511,629,825,850]" pageId="9" pageNumber="210">Hokkaidô</emphasis>
]: 1 3, Memuro-dake (Penkenûshi-dake), Shimizu-chô, Kamikawa-gun,
<date id="FFCEDACDFFD7FFFE53C4FCC2C4208E6D" box="[151,306,861,885]" pageId="9" pageNumber="210" value="1995-07-15">15. VII. 1995</date>
, K. Haga leg. (cHaga); 1 3, 1 Ƥ, Shari-dake, Shari-machi, Shari-gun,
<date id="FFCEDACDFFD7FFFE5702FCC2C1FE8E6D" box="[1105,1260,861,885]" pageId="9" pageNumber="210" value="1996-07-23">23. VII. 1996</date>
, K. Mizota leg. (KUM). [
<emphasis id="B904201FFFD7FFFE5250FC1EC4728E82" bold="true" box="[259,352,897,922]" pageId="9" pageNumber="210">Honshû</emphasis>
]: 1 3, Matsugabô-dam, Yamakami, Sôma-shi, Fukushima-ken,
<date id="FFCEDACDFFD7FFFE5761FC1EC1DF8E81" box="[1074,1229,897,921]" pageId="9" pageNumber="210" value="2006-12-20">20. XII. 2006</date>
, S. Saitô leg. (leaf litter in forest; cWat); 1 Ƥ, Azuma-yama, Inawashiro-machi, Yama-gun, Fukushima-ken,
<date id="FFCEDACDFFD7FFFE57FCFC3AC04E8EA5" box="[1199,1372,933,957]" pageId="9" pageNumber="210" value="1996-08-19">19. VIII. 1996</date>
, S.-I. Naomi leg. (KUM); 2 3, Ômomo, Minamiaizu-machi, Aizu-gun, Fukushima-ken,
<date id="FFCEDACDFFD7FFFE571BFC56C1C58EF9" box="[1096,1239,969,993]" pageId="9" pageNumber="210" value="1990-06-23">23. VI. 1990</date>
, T. Watanabe leg. (cWat); 1 3, 2 sex?, Maeshirane-san, Chûgushi, Nikkô-shi, Tochigi-ken,
<date id="FFCEDACDFFD7FFFE5082FC72C171891D" box="[977,1123,1005,1029]" pageId="9" pageNumber="210" value="1990-06-23">23. VI. 1990</date>
, T. Watanabe leg. (cWat); 4 sex?, Yumoto, Nikkô-shi, Tochigi-ken,
<date id="FFCEDACDFFD7FFFE511DFB8EC7CC8931" box="[590,734,1041,1065]" pageId="9" pageNumber="210" value="1982-06-29">29. VI. 1982</date>
, S.-I. Naomi leg. (KUM); 1 sex?, same data, but
<date id="FFCEDACDFFD7FFFE57ADFB8EC0858931" box="[1278,1431,1041,1065]" pageId="9" pageNumber="210" value="1994-07-10">10. VII. 1994</date>
; 2 sex?, Chûzenji, Nikkô-shi, Tochigi-ken,
<date id="FFCEDACDFFD7FFFE512EFBAAC62E8955" box="[637,828,1077,1101]" pageId="9" pageNumber="210" value="1982-06-28" valueMax="1982-06-30" valueMin="1982-06-28">2830. VI. 1982</date>
, S.-I. Naomi leg. (KUM); 1 Ƥ, Akagi-san (Komagatake peak), Fujimi-mura, Seta-gun, Gunma-ken,
<date id="FFCEDACDFFD7FFFE518EFBC6C6978969" box="[733,901,1113,1137]" pageId="9" pageNumber="210" value="1996-07-20">20. VII. 1996</date>
, S. Nomura leg. (KUM); 1 Ƥ, Hôshi-onsen hotspring, Minakami-machi, Tone-gun, Gunma-ken,
<date id="FFCEDACDFFD7FFFE5055FBE2C6B7898D" box="[774,933,1149,1173]" pageId="9" pageNumber="210" value="1997-06-26">26. VI. 1997</date>
, S.-I. Naomi leg. (KUM); 2 3, 6 ƤƤ, Akaguna-yama, Kamihino, Fujioka-shi, Gunma-ken,
<date id="FFCEDACDFFD7FFFE51A5FB3EC69B89A1" box="[758,905,1185,1209]" pageId="9" pageNumber="210" value="2001-09-13">13. IX. 2001</date>
, T. Watanabe leg. (cWat); 1 3, same data, but
<date id="FFCEDACDFFD7FFFE53C4FB5AC43189C5" box="[151,291,1221,1245]" pageId="9" pageNumber="210" value="2002-05-15">15. V. 2002</date>
; 1 3, Watarase-yûsuichi retarding basin, Fujioka-shi, Tochigi-ken,
<date id="FFCEDACDFFD7FFFE5767FB5AC1DB89C5" box="[1076,1225,1221,1245]" pageId="9" pageNumber="210" value="2002-04-13">13. IV. 2002</date>
, T. Watanabe leg. (cWat); 3 3, 3 ƤƤ, Matsuzawa-tôge pass (alt.
<quantity id="4C8851E8FFD7FFFE51E8FB76C61E8819" box="[699,780,1257,1281]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="9" pageNumber="210" unit="m" value="1150.0">1150m</quantity>
), Haruna-san, Takasaki-shi, Gunma-ken,
<date id="FFCEDACDFFD7FFFE578DFB76C0688819" box="[1246,1402,1257,1281]" pageId="9" pageNumber="210" value="1996-07-27">27. VII. 1996</date>
, S. Nomura leg. (KUM); 3 3, Nidoage-tôge pass, Kawaura, Kurabuchi-machi, Takasaki-shi, Gunma-ken,
<date id="FFCEDACDFFD7FFFE5618FA92C5C08851" pageId="9" pageNumber="210" value="2002-09-11">11. IX. 2002</date>
, T. Watanabe leg. (cWat); 6 sex?, Kanaibuchi-machi, Takasaki-shi, Gunma-ken,
<date id="FFCEDACDFFD7FFFE573DFAAEC0108851" box="[1134,1282,1329,1353]" pageId="9" pageNumber="210" value="2005-03-27">27. III. 2005</date>
, T. Watanabe leg. (cWat); 1 3, 7 sex?, Wagamine-machi, Takasaki-shi, Gunma-ken,
<date id="FFCEDACDFFD7FFFE50E1FACAC1508875" box="[946,1090,1365,1389]" pageId="9" pageNumber="210" value="2005-03-27">27. III. 2005</date>
, T. Watanabe leg. (cWat); 1 3, Kirizumi, Sakamoto, Matsuida-machi, Annaka-shi, Gunma-ken,
<date id="FFCEDACDFFD7FFFE5028FAE6C1378889" box="[891,1061,1401,1425]" pageId="9" pageNumber="210" value="1962-08-19">19. VIII. 1962</date>
,
<collectingCountry id="F367BC9DFFD7FFFE5761FAE5C1548889" box="[1074,1094,1402,1425]" name="Italy" pageId="9" pageNumber="210">Y</collectingCountry>
. Shibata leg. (cShi); 3 3, 3 ƤƤ, Kamioshidari, Higashimatsuyama-shi, Saitama-ken,
<date id="FFCEDACDFFD7FFFE5073FA02C6B188AD" box="[800,931,1437,1461]" pageId="9" pageNumber="210" value="1998-11-01">1. XI. 1998</date>
, K. Toyoda leg. (KUM); 1 3, Tenmoku-zan, Chichibu-shi, Saitama-ken,
<date id="FFCEDACDFFD7FFFE52B2FA5EC79688C1" box="[481,644,1473,1497]" pageId="9" pageNumber="210" value="1998-06-12">12. VI. 1998</date>
, K. Toyoda leg. (KUM); 3 ƤƤ, Hirogawarasawa, Nakatsugawa, Chichibu-shi, Saitama-ken,
<date id="FFCEDACDFFD7FFFE5282FA7AC74A88E5" box="[465,600,1509,1533]" pageId="9" pageNumber="210" value="1999-10-11">11. X. 1999</date>
, K. Toyoda leg. (KUM); 1 Ƥ, Tone-gawa, Sakae-machi, Inba-gun, Chibaken,
<date id="FFCEDACDFFD7FFFE539DF996C4578B39" box="[206,325,1545,1569]" pageId="9" pageNumber="210" value="1991-05-05">5. V. 1991</date>
, T. Takeda leg. (KUM); 1 Ƥ, Amatsu-kaigan, Uchiura, Kamogawa-shi, Chiba-ken,
<date id="FFCEDACDFFD7FFFE57A6F996C06B8B39" box="[1269,1401,1545,1569]" pageId="9" pageNumber="210" value="1990-11-03">3. XI. 1990</date>
, T. Takeda leg. (KUM); 1 Ƥ, Heisaura, Sakai, Tateyama-shi, Chiba-ken,
<date id="FFCEDACDFFD7FFFE50E4F9B2C1298B5D" box="[951,1083,1581,1605]" pageId="9" pageNumber="210" value="1997-04-01">1. IV. 1997</date>
, M. Maruyama leg. (KUM); 1 sex?, river mouth of Obitsu-gawa, Kuroto, Kisarazu-shi, Chiba-ken,
<date id="FFCEDACDFFD7FFFE50F1F9CEC13E8B71" box="[930,1068,1617,1641]" pageId="9" pageNumber="210" value="1998-10-25">25. X. 1998</date>
, K. Toyoda leg. (KUM); 1 sex?, Takao-san, Hachiôji-shi, Tôkyô-to,
<date id="FFCEDACDFFD7FFFE516BF9EAC7C38B95" box="[568,721,1653,1677]" pageId="9" pageNumber="210" value="1962-04-16">16. IV. 1962</date>
,
<collectingCountry id="F367BC9DFFD7FFFE51B1F9E9C7E48B95" box="[738,758,1654,1677]" name="Italy" pageId="9" pageNumber="210">Y</collectingCountry>
. Shibata leg. (cShi); 1 Ƥ, Arasaki, Nagai, Yokosuka-shi, Kanagawa-ken,
<date id="FFCEDACDFFD7FFFE521CF906C4F18BA9" box="[335,483,1689,1713]" pageId="9" pageNumber="210" value="1963-11-17">17. XI. 1963</date>
,
<collectingCountry id="F367BC9DFFD7FFFE52A2F905C7178BA9" box="[497,517,1690,1713]" name="Italy" pageId="9" pageNumber="210">Y</collectingCountry>
. Shibata leg. (cShi); 1 sex?, Hayama-machi, Miura-gun, Kanagawa-ken,
<date id="FFCEDACDFFD7FFFE561EF906C5C08BCD" pageId="9" pageNumber="210" value="1953-03-27">27. III. 1953</date>
, R. Kiryû leg. (SCM); 1 sex?, Kamimizo, Chuô-ku, Sagamihara-shi, Kanagawa-ken,
<date id="FFCEDACDFFD7FFFE57F0F922C0398BCD" box="[1187,1323,1725,1749]" pageId="9" pageNumber="210" value="1954-05-23">23. V. 1954</date>
, R. Kiryû leg. (SCM); 1 3, Shônan-kaigan, Kanagawa-ken,
<date id="FFCEDACDFFD7FFFE51B5F97EC66F8BE1" box="[742,893,1761,1785]" pageId="9" pageNumber="210" value="1976-07-02">2. VII. 1976</date>
, Niizato leg. (without collectors given name; KUM); 1 Ƥ, Mitsuishi (Manazuru-misaki), Manazuru-machi, Ashigarashimo-gun, Kanagawa-ken,
<date id="FFCEDACDFFD7FFFE57A8F89AC08A8A05" box="[1275,1432,1797,1821]" pageId="9" pageNumber="210" value="1989-07-15">15. VII. 1989</date>
, T. Watanabe leg. (under rubbish on the beach; cWat); 1 Ƥ, Susugaya (Tanzawayamafudakake), Kiyokawa-mura, Aikô-gun, Kanagawa-ken,
<date id="FFCEDACDFFD7FFFE5284F8D2C77B8A7D" box="[471,617,1869,1893]" pageId="9" pageNumber="210" value="1993-11-06">6. XI. 1993</date>
, T. Watanabe leg. (cWat); 1 3, 1 sex?, Tônosawa, Hakone-machi, Ashigarashimo-gun, kanagawa-ken,
<date id="FFCEDACDFFD7FFFE5167F8EEC7AC8A91" box="[564,702,1905,1929]" pageId="9" pageNumber="210" value="1971-10-24">24. X. 1971</date>
,
<collectingCountry id="F367BC9DFFD7FFFE5198F8EDC7CD8A91" box="[715,735,1906,1929]" name="Italy" pageId="9" pageNumber="210">Y</collectingCountry>
. Hirano leg. (KUM); 1 Ƥ, Usui-tôge, Karuizawa, Karuizawamachi, Kitasaku-gun, Nagano-ken,
<date id="FFCEDACDFFD7FFFE5163F80AC7C08AB5" box="[560,722,1941,1965]" pageId="9" pageNumber="210" value="1999-07-31">31. VII. 1999</date>
, S.-I. Naomi leg. (KUM); 13 sex?, Kamihikawa, Daibosatsu, Kôshû-shi, Yamanashi-ken,
<date id="FFCEDACDFFD7FFFE5281F826C7708AC9" box="[466,610,1977,2001]" pageId="9" pageNumber="210" value="1986-06-14">14. VI. 1986</date>
, T. Watanabe leg. (cWat); 4 3, 1 Ƥ, near Tsuchimuro-gawa, Daibosatsu, Yamanashi-ken,
<date id="FFCEDACDFFD7FFFE5201F842C7108AED" box="[338,514,2013,2037]" pageId="9" pageNumber="210" value="1982-10-16" valueMax="1982-10-17" valueMin="1982-10-16">1617. X. 1982</date>
,
<collectingCountry id="F367BC9DFFD7FFFE515EF841C7338AED" box="[525,545,2014,2037]" name="Italy" pageId="9" pageNumber="210">Y</collectingCountry>
. Shibata leg. (cShi); 1 sex?, Hôoh-goya lodge, Minamiarupusu-shi, Yamanashiken,
<date id="FFCEDACDFFD4FFFD539DFF08C47B8DB7" box="[206,361,151,175]" pageId="10" pageNumber="211" value="1967-07-26">26. VII. 1967</date>
,
<collectingCountry id="F367BC9DFFD4FFFD5226FF07C49B8DB7" box="[373,393,152,175]" name="Italy" pageId="10" pageNumber="211">Y</collectingCountry>
. Shibata leg. (cShi); 1 Ƥ, same locality,
<date id="FFCEDACDFFD4FFFD5003FF08C6F38DB7" box="[848,993,151,175]" pageId="10" pageNumber="211" value="1990-06-11">11. VI. 1990</date>
, K. Hosoda leg. (cIto); 1 Ƥ, same data, but
<date id="FFCEDACDFFD4FFFD5396FF23C4428DCC" box="[197,336,188,212]" pageId="10" pageNumber="211" value="1993-06-02">2. VI. 1993</date>
; 1 sex?, Hirogawara (Shirane-san), Minamiarupusu-shi, Yamanashi-ken,
<date id="FFCEDACDFFD4FFFD57CCFF23C04F8DCC" box="[1183,1373,188,212]" pageId="10" pageNumber="211" value="1982-07-09" valueMax="1982-07-13" valueMin="1982-07-09">913. VII. 1982</date>
, S.-I. Naomi leg. (KUM); 1 3, 2 sex?, Yanbushi-tôge pass, Hayakawa-chô, Minamikoma-gun, Yamanashi-ken,
<date id="FFCEDACDFFD4FFFD5603FF40C5C08C04" pageId="10" pageNumber="211" value="2002-07-04">4. VII. 2002</date>
, A. Sugimura leg. (KUM); 1 Ƥ, 1 sex?, river mouth of Yodo-gawa, Ôsaka-shi, Ôsaka-fu,
<date id="FFCEDACDFFD4FFFD57B0FE9BC0648C04" box="[1251,1398,260,284]" pageId="10" pageNumber="211" value="1956-03-17">17. III. 1956</date>
,
<collectingCountry id="F367BC9DFFD4FFFD56D7FE9AC08A8C04" box="[1412,1432,261,284]" name="Italy" pageId="10" pageNumber="211">Y</collectingCountry>
. Hayashi leg. (cHay). [
<emphasis id="B904201FFFD4FFFD52C0FEB8C4E58C58" bold="true" box="[403,503,295,320]" pageId="10" pageNumber="211">Shikoku</emphasis>
]: 1 3, 2 ƤƤ, 1 sex?, Ôtaki-san, Shionoe-chô, Takamatsu-shi, Kagawa-ken,
<date id="FFCEDACDFFD4FFFD5609FEB8C5C08C7C" pageId="10" pageNumber="211" value="2008-06-08">8. VI. 2008</date>
, H. Fujimoto leg. (KUM); 2 3, Tsurugi-san, Tokushima-ken,
<date id="FFCEDACDFFD4FFFD50E2FED3C1628C7C" box="[945,1136,332,356]" pageId="10" pageNumber="211" value="1981-06-19" valueMax="1981-06-20" valueMin="1981-06-19">1920. VI. 1981</date>
, S.-I. Naomi leg. (KUM). [
<emphasis id="B904201FFFD4FFFD53F3FEF0C5EF8C90" bold="true" box="[160,253,367,392]" pageId="10" pageNumber="211">Kyûshû</emphasis>
]:
<date id="FFCEDACDFFD4FFFD5248FEF0C4688C9F" box="[283,378,367,392]" pageId="10" pageNumber="211" value="1919-01-03">1 3, 19</date>
sex?, Shakaga-dake (alt.
<quantity id="4C8851E8FFD4FFFD51FEFEF0C7ED8C9F" box="[685,767,367,391]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="211" unit="m" value="1200.0">1200m</quantity>
), Yame-shi, Fukuoka-ken,
<date id="FFCEDACDFFD4FFFD576FFEF0C1C58C9F" box="[1084,1239,367,391]" pageId="10" pageNumber="211" value="1994-09-25">25. IX. 1994</date>
, S. Nomura leg. (KUM); 2 sex?, Sobo-san, Ôita-ken,
<date id="FFCEDACDFFD4FFFD5160FE0BC7D98CB4" box="[563,715,404,428]" pageId="10" pageNumber="211" value="1985-07-19">19. VII. 1985</date>
, K. Ogata leg. (KUM); 1 3, 2 ƤƤ, 1 sex?, Sobo-san, Taketa-shi, Ôita-ken,
<date id="FFCEDACDFFD4FFFD5259FE27C4818CD7" box="[266,403,439,463]" pageId="10" pageNumber="211" value="2009-06-07">7. VI. 2009</date>
, S. Yamamoto leg. (daytime on walkway; cYam); 13 sex?, same data, but
<date id="FFCEDACDFFD4FFFD57ACFE28C08E8CD7" box="[1279,1436,439,463]" pageId="10" pageNumber="211" value="2011-07-14">14. VII. 2011</date>
(besides these records, dozens of individuals were observed (only field observation, no collecting permit) at the top of the mountain which is situated on a boundary of Ôita-ken and Miyazaki-ken:
<figureCitation id="134BE088FFD4FFFD574BFE60C1798F0F" box="[1048,1131,511,535]" captionStart="FIGURES 42 43" captionStartId="17.[151,264,1257,1280]" captionTargetBox="[157,1430,811,1230]" captionTargetId="figure@17.[151,1436,807,1235]" captionTargetPageId="17" captionText="FIGURES 42 43. Habitats of the subgenus Coprochara of the genus Aleochara in Japan. 93. the summit of Sobo-san (located in boundaries between Ôita and Miyazaki Prefectures; only observed; A. (C.) verna: arrow); 43. Watari (Fukutsu, Fukuoka Prefecture; A. (C.) squalithorax: arrow)." httpUri="https://zenodo.org/record/223172/files/figure.png" pageId="10" pageNumber="211">Fig. 42</figureCitation>
); 3 ƤƤ, 2 sex?, Kurokawa, Aso-shi, Kumamoto-ken,
<date id="FFCEDACDFFD4FFFD52EAFDBBC7288F24" box="[441,570,548,572]" pageId="10" pageNumber="211" value="2009-06-06">6. VI. 2009</date>
, S. Yamamoto leg. (from cow dung on pasture during daytime; cYam); 1 3, 1 Ƥ, Ichifusa-yama, Mizukami-mura, Kuma-gun, Kumamoto-ken,
<date id="FFCEDACDFFD4FFFD502EFDD8C1218F47" box="[893,1075,583,607]" pageId="10" pageNumber="211" value="1988-08-01" valueMax="1988-08-02" valueMin="1988-08-01">12. VIII. 1988</date>
, S.-I. Naomi leg. (KUM); 1 Ƥ, Tomioka (Amakusa-syotô), Reihoku-machi, Amakusa-gun, Kumamoto-ken,
<date id="FFCEDACDFFD4FFFD50ABFDF3C16A8F9C" box="[1016,1144,620,644]" pageId="10" pageNumber="211" value="1977-04-03">3. IV. 1977</date>
, S.-I. Naomi leg. (KUM); 1 3, Nichinan-shi, Miyazaki-ken,
<date id="FFCEDACDFFD4FFFD517EFD10C7AE8FBF" box="[557,700,655,679]" pageId="10" pageNumber="211" value="1969-05-29">29. V. 1969</date>
, Watanabe leg. (without collectors given name; KUM); 1 Ƥ, Wanitsuka-yama, Kou, Tano-chô, Miyazaki-shi, Miyazaki-ken,
<date id="FFCEDACDFFD4FFFD502DFD2BC11E8FD4" box="[894,1036,692,716]" pageId="10" pageNumber="211" value="1993-09-06">6. IX. 1993</date>
, S. Nomura leg. (KUM); 1 sex?, Kirishima-yama, Kobayashi-shi, Miyazaki-ken,
<date id="FFCEDACDFFD4FFFD519CFD48C67E8FF7" box="[719,876,727,751]" pageId="10" pageNumber="211" value="1982-07-04">4. VII. 1982</date>
, M. Takanashi leg. (KUM); 1 3, Takachiho, Miyazaki-ken,
<date id="FFCEDACDFFD4FFFD526DFD63C4DF8E0C" box="[318,461,764,788]" pageId="10" pageNumber="211" value="1969-06-24">24. VI. 1969</date>
,
<collectingCountry id="F367BC9DFFD4FFFD528BFD62C4FE8E0C" box="[472,492,765,788]" name="Italy" pageId="10" pageNumber="211">Y</collectingCountry>
. Goto leg. (cShi); 28 sex?, Takachihonomine, Miike-chô, Miyakonojô-shi, Miyazakiken,
<date id="FFCEDACDFFD4FFFD539CFC80C4448E2F" box="[207,342,799,823]" pageId="10" pageNumber="211" value="1982-05-21">21. V. 1982</date>
, H. Takemoto leg. (KUM); 1 sex?, same locality,
<date id="FFCEDACDFFD4FFFD50C7FC80C1238E2F" box="[916,1073,799,823]" pageId="10" pageNumber="211" value="1983-07-19">19. VII. 1983</date>
, M. Ôhara leg. (KUM); 8 sex?, Takachihonomine,
<date id="FFCEDACDFFD4FFFD523FFCDBC4E28E44" box="[364,496,836,860]" pageId="10" pageNumber="211" value="1981-05-31">31. V. 1981</date>
,
<collectingCountry id="F367BC9DFFD4FFFD52AEFCDAC7038E44" box="[509,529,837,860]" name="Italy" pageId="10" pageNumber="211">Y</collectingCountry>
. Takai leg. (KUM); 5 sex?, same locality,?
<date id="FFCEDACDFFD4FFFD50ACFCDAC1658E44" box="[1023,1143,836,860]" pageId="10" pageNumber="211" value="1979-08">VIII. 1979</date>
, Koda leg. (without detail collecting date and collectors given name; KUM); 1 Ƥ, 19 sex?, Takachihonomine, kirishimataguchi, Kirishima-shi, Kagoshima-ken, N. Koda leg. (without collecting date; KUM); 1 3, 1 Ƥ, 3 sex?, Kirishima-yama (near Miike pond), Kagoshima-ken,
<date id="FFCEDACDFFD4FFFD5200FC30C4C68EDF" box="[339,468,943,967]" pageId="10" pageNumber="211" value="1983-05-18">18. V. 1983</date>
,
<collectingCountry id="F367BC9DFFD4FFFD528DFC2FC4E08EDF" box="[478,498,944,967]" name="Italy" pageId="10" pageNumber="211">Y</collectingCountry>
. Sawada leg. (KUM); 1 3, Unzen-dake, Unzen-shi, Nagasaki-ken,
<date id="FFCEDACDFFD4FFFD5785FC30C0448EDF" box="[1238,1366,943,967]" pageId="10" pageNumber="211" value="1986-05-17">17. V. 1986</date>
, T. Itô leg. (cIto); 3 sex?, same locality,
<date id="FFCEDACDFFD4FFFD5150FC4BC79E8EF4" box="[515,652,980,1004]" pageId="10" pageNumber="211" value="1977-07-03">3. VII. 1977</date>
, S. Imasaka leg. (KUM); 1 Ƥ, same data, but
<date id="FFCEDACDFFD4FFFD57D0FC4BC01F8EF4" box="[1155,1293,980,1004]" pageId="10" pageNumber="211" value="1978-07-04">4. VII. 1978</date>
; 1 3, 1 sex?, same data, but
<date id="FFCEDACDFFD4FFFD526FFC68C4DE8917" box="[316,460,1015,1039]" pageId="10" pageNumber="211" value="1979-06-24">24. VI. 1979</date>
; 1 Ƥ, same data, but
<date id="FFCEDACDFFD4FFFD51E7FC68C6558917" box="[692,839,1015,1039]" pageId="10" pageNumber="211" value="1980-08-05">5. VIII. 1980</date>
; 1 Ƥ, same data, but
<date id="FFCEDACDFFD4FFFD5763FC68C1D28917" box="[1072,1216,1015,1039]" pageId="10" pageNumber="211" value="1981-06-24">24. VI. 1981</date>
.
</paragraph>
<paragraph id="8BCFFC0DFFD4FFFD53EFFB84C16B894F" blockId="10.[151,1437,151,2006]" pageId="10" pageNumber="211">
<emphasis id="B904201FFFD4FFFD53EFFB84C46A892C" bold="true" box="[188,376,1051,1076]" pageId="10" pageNumber="211">Other material.</emphasis>
<collectingCountry id="F367BC9DFFD4FFFD522CFB83C4F5892C" box="[383,487,1052,1076]" name="Russia" pageId="10" pageNumber="211">RUSSIA</collectingCountry>
: [
<emphasis id="B904201FFFD4FFFD52A8FB83C611892C" bold="true" box="[507,771,1051,1076]" pageId="10" pageNumber="211">Kamchatka Peninsula</emphasis>
]: 1 Ƥ, Nagornyi (
<geoCoordinate id="EE449ACAFFD4FFFD509CFB83C13E892C" box="[975,1068,1052,1076]" direction="north" orientation="latitude" pageId="10" pageNumber="211" precision="925" value="53.116665">53°07'N</geoCoordinate>
<geoCoordinate id="EE449ACAFFD4FFFD5767FB83C18C892C" box="[1076,1182,1052,1076]" direction="east" orientation="longitude" pageId="10" pageNumber="211" precision="925" value="158.51666">158°31'E</geoCoordinate>
), near Petropavlovsk- Kamachatskiy,
<date id="FFCEDACDFFD4FFFD5210FBA0C4CF894F" box="[323,477,1087,1111]" pageId="10" pageNumber="211" value="1997-07-10">10. VII. 1997</date>
, A. Saitô leg. (Naomi
<emphasis id="B904201FFFD4FFFD518DFBDEC6028940" box="[734,784,1088,1112]" italics="true" pageId="10" pageNumber="211">et al</emphasis>
., 2000; CBM: CBM-ZI/81531).
</paragraph>
<paragraph id="8BCFFC0DFFD4FFFC5394FBFCC0868C7A" blockId="10.[151,1437,151,2006]" lastBlockId="11.[151,1437,151,2034]" lastPageId="11" lastPageNumber="212" pageId="10" pageNumber="211">
<emphasis id="B904201FFFD4FFFD5394FBFCC4A88964" bold="true" box="[199,442,1123,1148]" pageId="10" pageNumber="211">Redescription. Body</emphasis>
(
<figureCitation id="134BE088FFD4FFFD529AFBFBC7198964" box="[457,523,1124,1148]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[278,1306,191,1972]" captionTargetId="figure@2.[278,1308,161,1975]" captionTargetPageId="2" captionText="FIGURES 1 4. Japanese species of Aleochara (Coprochara) in habitus. 1. Aleochara (Coprochara) verna; 2. A. (C.) binotata; 3 4. A. (C.) squalithorax: gray type (Fig. 3) and yellow type (Fig. 4)." httpUri="https://zenodo.org/record/223166/files/figure.png" pageId="10" pageNumber="211">Fig. 1</figureCitation>
): slender and subparallel, normally moderate in size, ca.
<quantity id="4C8851E8FFD4FFFD57DCFBFBC1E18964" box="[1167,1267,1124,1148]" metricMagnitude="-3" metricUnit="m" metricValue="3.63" pageId="10" pageNumber="211" unit="mm" value="3.63">3.63 mm</quantity>
in body length (
<quantity id="4C8851E8FFD4FFFD53CDFB18C45A8987" box="[158,328,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="3.72" metricValueMax="4.32" metricValueMin="3.12" pageId="10" pageNumber="211" unit="mm" value="3.72" valueMax="4.32" valueMin="3.12">3.124.32 mm</quantity>
, N = 15), and
<quantity id="4C8851E8FFD4FFFD52A7FB18C7498987" box="[500,603,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" pageId="10" pageNumber="211" unit="mm" value="1.72">1.72 mm</quantity>
in fore body length (
<quantity id="4C8851E8FFD4FFFD5001FB18C6E88987" box="[850,1018,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" metricValueMax="1.83" metricValueMin="1.41" pageId="10" pageNumber="211" unit="mm" value="1.62" valueMax="1.83" valueMin="1.41">1.411.83 mm</quantity>
, N = 15), robust, and dorsal surface clearly glossy.
<emphasis id="B904201FFFD4FFFD5210FB34C49A89DC" bold="true" box="[323,392,1195,1220]" pageId="10" pageNumber="211">Color</emphasis>
(
<figureCitation id="134BE088FFD4FFFD52CAFB34C4FB89DB" box="[409,489,1195,1219]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[278,1306,191,1972]" captionTargetId="figure@2.[278,1308,161,1975]" captionTargetPageId="2" captionText="FIGURES 1 4. Japanese species of Aleochara (Coprochara) in habitus. 1. Aleochara (Coprochara) verna; 2. A. (C.) binotata; 3 4. A. (C.) squalithorax: gray type (Fig. 3) and yellow type (Fig. 4)." httpUri="https://zenodo.org/record/223166/files/figure.png" pageId="10" pageNumber="211">Figs. 1</figureCitation>
,
<figureCitation id="134BE088FFD4FFFD52AAFB34C73589DB" box="[505,551,1195,1219]" captionStart="FIGURES 5 13" captionStartId="3.[151,264,1073,1096]" captionTargetBox="[163,1424,194,1036]" captionTargetId="figure@3.[151,1436,171,1052]" captionTargetPageId="3" captionText="FIGURES 5 13. Japanese species of Aleochara (Coprochara). 5 7. Aleochara (Coprochara) verna; 8 10. A. (C.) binotata; 11 13. A. (C.) squalithorax. (5, 8, 11: elytra; 6, 9, 12: intercoxal process of metaventrite; 7, 10, 13: mesoventrite and metaventrite)." httpUri="https://zenodo.org/record/223167/files/figure.png" pageId="10" pageNumber="211">57</figureCitation>
): ground color completely black; legs, especially tarsal segments, and mouth parts brown to reddish brown; elytra with a pair of yellowish spots; antennae blackish brown to reddish brown, surface moderately to densely pubescent with minute whitish setae.
<emphasis id="B904201FFFD4FFFD50F2FB6CC6F38813" bold="true" box="[929,993,1267,1291]" pageId="10" pageNumber="211">Head</emphasis>
(
<figureCitation id="134BE088FFD4FFFD50A1FB6CC1248813" box="[1010,1078,1267,1291]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[278,1306,191,1972]" captionTargetId="figure@2.[278,1308,161,1975]" captionTargetPageId="2" captionText="FIGURES 1 4. Japanese species of Aleochara (Coprochara) in habitus. 1. Aleochara (Coprochara) verna; 2. A. (C.) binotata; 3 4. A. (C.) squalithorax: gray type (Fig. 3) and yellow type (Fig. 4)." httpUri="https://zenodo.org/record/223166/files/figure.png" pageId="10" pageNumber="211">Fig. 1</figureCitation>
): circular (head length =
<quantity id="4C8851E8FFD4FFFD560EFB6CC5EF8836" metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.699999999999999" metricValueMin="4.699999999999999" pageId="10" pageNumber="211" unit="mm" value="0.52" valueMax="0.57" valueMin="0.47">0.47 0.57 mm</quantity>
(mean =
<quantity id="4C8851E8FFD4FFFD523FFA89C4C48836" box="[364,470,1302,1326]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="10" pageNumber="211" unit="mm" value="0.53">0.53 mm</quantity>
), head width =
<quantity id="4C8851E8FFD4FFFD51DAFA89C63C8836" box="[649,814,1302,1326]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" metricValueMax="6.3" metricValueMin="4.9" pageId="10" pageNumber="211" unit="mm" value="0.56" valueMax="0.63" valueMin="0.49">0.490.63 mm</quantity>
(mean =
<quantity id="4C8851E8FFD4FFFD50CDFA89C1158836" box="[926,1031,1302,1326]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="10" pageNumber="211" unit="mm" value="0.57">0.57 mm</quantity>
), N = 15), slightly transverse (head width / head length = 1.08 (mean), N = 15), widest at eyes; surface covered with long and thin setae sparsely, punctation shallow and inconspicuous.
<emphasis id="B904201FFFD4FFFD510EFAC1C7C2886F" bold="true" box="[605,720,1374,1399]" pageId="10" pageNumber="211">Antennae</emphasis>
(
<figureCitation id="134BE088FFD4FFFD51B0FAC1C63A886E" box="[739,808,1374,1398]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[278,1306,191,1972]" captionTargetId="figure@2.[278,1308,161,1975]" captionTargetPageId="2" captionText="FIGURES 1 4. Japanese species of Aleochara (Coprochara) in habitus. 1. Aleochara (Coprochara) verna; 2. A. (C.) binotata; 3 4. A. (C.) squalithorax: gray type (Fig. 3) and yellow type (Fig. 4)." httpUri="https://zenodo.org/record/223166/files/figure.png" pageId="10" pageNumber="211">Fig. 1</figureCitation>
): moniliform, thick and robust, shorter than head and pronotum combined (antenna length =
<quantity id="4C8851E8FFD4FFFD5100FA1EC7E58881" box="[595,759,1409,1433]" metricMagnitude="-4" metricUnit="m" metricValue="8.45" metricValueMax="9.5" metricValueMin="7.4" pageId="10" pageNumber="211" unit="mm" value="0.845" valueMax="0.95" valueMin="0.74">0.740.95 mm</quantity>
(mean =
<quantity id="4C8851E8FFD4FFFD5035FA1EC6DD8881" box="[870,975,1409,1433]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="10" pageNumber="211" unit="mm" value="0.87">0.87 mm</quantity>
), N = 15); segment I, about 2.3 times as long as broad, segment II clearly shorter than I, segment III slightly shorter than II, segment IV spherical, as long as width except for stem of segment, segments V to X clearly transverse, segment XI thick and subconical, nearly 1.8 times as long as width, longer than segment I, approximate relative length of segments from basal to apex: 12.0: 10.5: 7.5: 4.0: 4.5: 5.0: 5.0: 5.0: 5.0: 5.0: 15.0.
<emphasis id="B904201FFFD4FFFD51F7F98EC7EF8B31" bold="true" box="[676,765,1553,1577]" pageId="10" pageNumber="211">Thorax</emphasis>
: pronotum (
<figureCitation id="134BE088FFD4FFFD50DBF98FC6DB8B30" box="[904,969,1552,1576]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[278,1306,191,1972]" captionTargetId="figure@2.[278,1308,161,1975]" captionTargetPageId="2" captionText="FIGURES 1 4. Japanese species of Aleochara (Coprochara) in habitus. 1. Aleochara (Coprochara) verna; 2. A. (C.) binotata; 3 4. A. (C.) squalithorax: gray type (Fig. 3) and yellow type (Fig. 4)." httpUri="https://zenodo.org/record/223166/files/figure.png" pageId="10" pageNumber="211">Fig. 1</figureCitation>
) moderately transverse (pronotum length =
<quantity id="4C8851E8FFD4FFFD53E2F9ABC44B8B54" box="[177,345,1588,1612]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="7.9" metricValueMin="5.4" pageId="10" pageNumber="211" unit="mm" value="0.665" valueMax="0.79" valueMin="0.54">0.540.79 mm</quantity>
(mean =
<quantity id="4C8851E8FFD4FFFD529CF9ABC7298B54" box="[463,571,1588,1612]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="211" unit="mm" value="0.65">0.65 mm</quantity>
), pronotum width =
<quantity id="4C8851E8FFD4FFFD5063F9ABC6C58B54" box="[816,983,1588,1612]" metricMagnitude="-4" metricUnit="m" metricValue="7.95" metricValueMax="9.1" metricValueMin="6.8" pageId="10" pageNumber="211" unit="mm" value="0.795" valueMax="0.91" valueMin="0.68">0.680.91 mm</quantity>
(mean =
<quantity id="4C8851E8FFD4FFFD571DF9ABC1A88B54" box="[1102,1210,1588,1612]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="10" pageNumber="211" unit="mm" value="0.84">0.84 mm</quantity>
), pronotum width / pronotum length = 1.29 (mean), N = 15), wider than head (pronotum width / head width =1.49 (mean), N = 15), widest just after middle; surface coarsely and shallowly punctured, with weak longitudinal rows of punctures along midline. Intercoxal process of mesoventrite (
<figureCitation id="134BE088FFD4FFFD51FFF93FC7E48BA0" box="[684,758,1696,1720]" captionStart="FIGURES 5 13" captionStartId="3.[151,264,1073,1096]" captionTargetBox="[163,1424,194,1036]" captionTargetId="figure@3.[151,1436,171,1052]" captionTargetPageId="3" captionText="FIGURES 5 13. Japanese species of Aleochara (Coprochara). 5 7. Aleochara (Coprochara) verna; 8 10. A. (C.) binotata; 11 13. A. (C.) squalithorax. (5, 8, 11: elytra; 6, 9, 12: intercoxal process of metaventrite; 7, 10, 13: mesoventrite and metaventrite)." httpUri="https://zenodo.org/record/223167/files/figure.png" pageId="10" pageNumber="211">Fig. 7</figureCitation>
) with strong carina along midline. Intercoxal process of metaventrite (
<figureCitation id="134BE088FFD4FFFD5267F95CC4B08BC3" box="[308,418,1731,1755]" captionStart="FIGURES 5 13" captionStartId="3.[151,264,1073,1096]" captionTargetBox="[163,1424,194,1036]" captionTargetId="figure@3.[151,1436,171,1052]" captionTargetPageId="3" captionText="FIGURES 5 13. Japanese species of Aleochara (Coprochara). 5 7. Aleochara (Coprochara) verna; 8 10. A. (C.) binotata; 11 13. A. (C.) squalithorax. (5, 8, 11: elytra; 6, 9, 12: intercoxal process of metaventrite; 7, 10, 13: mesoventrite and metaventrite)." httpUri="https://zenodo.org/record/223167/files/figure.png" pageId="10" pageNumber="211">Figs. 67</figureCitation>
) broad and short, rounded apically. Metaventrite (
<figureCitation id="134BE088FFD4FFFD50B7F95CC1388BC3" box="[996,1066,1731,1755]" captionStart="FIGURES 5 13" captionStartId="3.[151,264,1073,1096]" captionTargetBox="[163,1424,194,1036]" captionTargetId="figure@3.[151,1436,171,1052]" captionTargetPageId="3" captionText="FIGURES 5 13. Japanese species of Aleochara (Coprochara). 5 7. Aleochara (Coprochara) verna; 8 10. A. (C.) binotata; 11 13. A. (C.) squalithorax. (5, 8, 11: elytra; 6, 9, 12: intercoxal process of metaventrite; 7, 10, 13: mesoventrite and metaventrite)." httpUri="https://zenodo.org/record/223167/files/figure.png" pageId="10" pageNumber="211">Fig. 7</figureCitation>
) smooth, pubescent like elytra. Elytra (
<figureCitation id="134BE088FFD4FFFD53BFF978C4238BE7" box="[236,305,1767,1791]" captionStart="FIGURES 5 13" captionStartId="3.[151,264,1073,1096]" captionTargetBox="[163,1424,194,1036]" captionTargetId="figure@3.[151,1436,171,1052]" captionTargetPageId="3" captionText="FIGURES 5 13. Japanese species of Aleochara (Coprochara). 5 7. Aleochara (Coprochara) verna; 8 10. A. (C.) binotata; 11 13. A. (C.) squalithorax. (5, 8, 11: elytra; 6, 9, 12: intercoxal process of metaventrite; 7, 10, 13: mesoventrite and metaventrite)." httpUri="https://zenodo.org/record/223167/files/figure.png" pageId="10" pageNumber="211">Fig. 5</figureCitation>
) wider than long (elytra length =
<quantity id="4C8851E8FFD4FFFD51EAF978C64C8BE7" box="[697,862,1767,1791]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" metricValueMax="7.0" metricValueMin="5.4" pageId="10" pageNumber="211" unit="mm" value="0.62" valueMax="0.7" valueMin="0.54">0.540.70 mm</quantity>
(mean =
<quantity id="4C8851E8FFD4FFFD509CF978C12B8BE7" box="[975,1081,1767,1791]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="211" unit="mm" value="0.62">0.62 mm</quantity>
), elytra width =
<quantity id="4C8851E8FFD4FFFD57ABF978C08E8BE7" box="[1272,1436,1767,1791]" metricMagnitude="-4" metricUnit="m" metricValue="9.4" metricValueMax="11.1" metricValueMin="7.7" pageId="10" pageNumber="211" unit="mm" value="0.94" valueMax="1.11" valueMin="0.77">0.771.11 mm</quantity>
(mean =
<quantity id="4C8851E8FFD4FFFD53AEF894C4758A3B" box="[253,359,1803,1827]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="10" pageNumber="211" unit="mm" value="0.98">0.98 mm</quantity>
), N = 15) with a pair of yellowish small spots extending from posterior margin close to elytral suture; surface shallowly punctured and with inconspicuous setae; posterior margin weakly rounded.
<emphasis id="B904201FFFD4FFFD5651F8B0C0288A5F" bold="true" box="[1282,1338,1839,1863]" pageId="10" pageNumber="211">Legs</emphasis>
(
<figureCitation id="134BE088FFD4FFFD561AF8B1C0998A5E" box="[1353,1419,1838,1862]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[278,1306,191,1972]" captionTargetId="figure@2.[278,1308,161,1975]" captionTargetPageId="2" captionText="FIGURES 1 4. Japanese species of Aleochara (Coprochara) in habitus. 1. Aleochara (Coprochara) verna; 2. A. (C.) binotata; 3 4. A. (C.) squalithorax: gray type (Fig. 3) and yellow type (Fig. 4)." httpUri="https://zenodo.org/record/223166/files/figure.png" pageId="10" pageNumber="211">Fig. 1</figureCitation>
): short (hind tibia length =
<quantity id="4C8851E8FFD4FFFD52EFF8CDC7728A72" box="[444,608,1874,1898]" metricMagnitude="-4" metricUnit="m" metricValue="5.45" metricValueMax="6.5" metricValueMin="4.4" pageId="10" pageNumber="211" unit="mm" value="0.545" valueMax="0.65" valueMin="0.44">0.440.65 mm</quantity>
(mean =
<quantity id="4C8851E8FFD4FFFD519CF8CDC62A8A72" box="[719,824,1874,1898]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="211" unit="mm" value="0.55">0.55 mm</quantity>
), N =15), dozens of long spines on fore and midtibia, respectively; relative lengths of tarsal segments from basal to apical: 4.5: 3.0: 3.0: 3.0:
<quantity id="4C8851E8FFD4FFFD5724F8E9C1AA8A96" box="[1143,1208,1910,1934]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" pageId="10" pageNumber="211" unit="in" value="8.5">8.5 in</quantity>
foretarsus, 9.0: 5.0: 5.0: 5.0: 11.0 in midtarsus, 13.0: 8.0: 8.0: 8.0:
<quantity id="4C8851E8FFD4FFFD51FCF806C7ED8AA9" box="[687,767,1945,1969]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" pageId="10" pageNumber="211" unit="in" value="14.5">14.5 in</quantity>
hindtarsus.
<emphasis id="B904201FFFD4FFFD50D8F806C1128AAA" bold="true" box="[907,1024,1945,1970]" pageId="10" pageNumber="211">Abdomen</emphasis>
(
<figureCitation id="134BE088FFD4FFFD5742F806C1478AA9" box="[1041,1109,1945,1969]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[278,1306,191,1972]" captionTargetId="figure@2.[278,1308,161,1975]" captionTargetPageId="2" captionText="FIGURES 1 4. Japanese species of Aleochara (Coprochara) in habitus. 1. Aleochara (Coprochara) verna; 2. A. (C.) binotata; 3 4. A. (C.) squalithorax: gray type (Fig. 3) and yellow type (Fig. 4)." httpUri="https://zenodo.org/record/223166/files/figure.png" pageId="10" pageNumber="211">Fig. 1</figureCitation>
): weakly glossy, covered by short and thin setae.
<emphasis id="B904201FFFD4FFFD52C7F821C4F18ACE" bold="true" box="[404,483,1982,2006]" pageId="10" pageNumber="211">
[
<collectingCountry id="F367BC9DFFD4FFFD52C8F821C4CE8ACE" box="[411,476,1982,2006]" name="Malta" pageId="10" pageNumber="211">Male</collectingCountry>
]
</emphasis>
: posterior margin of tergite VIII (
<figureCitation id="134BE088FFD4FFFD502CF822C6C48ACD" box="[895,982,1981,2005]" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="10" pageNumber="211">Fig. 14</figureCitation>
) clearly serrate, with approximately 8 macrosetae on each side. Sternite VIII (
<figureCitation id="134BE088FFD5FFFC5126FF08C7DE8DB7" box="[629,716,151,175]" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="11" pageNumber="212">Fig. 16</figureCitation>
) with approximately 8 macrosetae; posterior margin pointed triangularly. Median lobe of aedeagus as in
<figureCitation id="134BE088FFD5FFFC51C0FF24C60C8DCB" box="[659,798,187,211]" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="11" pageNumber="212">Figs. 1819</figureCitation>
: elongated and narrowed toward apex; flagellum inside projecting, longer than median lobe of aedeagus; basal plate of flagellum small and short; sclerite Z with long attachment, waving apically; apical lobe in lateral view moderately bent, narrowing subapically; basal swelling of median lobe circularly rounded; median lobe in ventral view moderately tear-shaped; basal swelling of median lobe small; apical lobe looks short isosceles. Apical lobe of paramerite (
<figureCitation id="134BE088FFD5FFFC50D5FED5C6C58C7A" box="[902,983,330,354]" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="11" pageNumber="212">Fig. 20</figureCitation>
) long and slender, bearing 4 long setae.
</paragraph>
</subSubSection>
<subSubSection id="C36AAF86FFD5FFFC5394FEF1C6D58F44" pageId="11" pageNumber="212" type="description">
<paragraph id="8BCFFC0DFFD5FFFC5394FEF1C6D58F44" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="212">
<emphasis id="B904201FFFD5FFFC5394FEF1C4228C9E" bold="true" box="[199,304,366,390]" pageId="11" pageNumber="212">[Female]</emphasis>
: tergite VIII (
<figureCitation id="134BE088FFD5FFFC5280FEF1C7348C9E" box="[467,550,366,390]" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="11" pageNumber="212">Fig. 15</figureCitation>
) with weakly serrate posterior margin, with 7 macrosetae. Sternite VIII (
<figureCitation id="134BE088FFD5FFFC563DFEF1C5A68CB2" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="11" pageNumber="212">Fig. 17</figureCitation>
) rounded posteriorly, with 8 macrosetae. Spermatheca (
<figureCitation id="134BE088FFD5FFFC5062FE0DC69A8CB2" box="[817,904,402,426]" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="11" pageNumber="212">Fig. 21</figureCitation>
): apical invagination of spermatheca small but deep; spermathecal head spherical, as long as spermathecal neck; attachment of spermathecal duct inconspicuous; (sn) clearly narrowing toward basal portion of spermathecal stem, stem moderately curved to almost angled at its base; coiled portion of moderate length, composed of maximally 20 thin coils, not strongly extended laterally; coils directly connected with thin membranous portion of spermathecal duct; each part of spermatheca except for (sm) entirely and moderately sclerotized; inner wall of (sh) finely striate.
</paragraph>
</subSubSection>
<subSubSection id="C36AAF86FFD5FFFC5394FDF7C4808E62" pageId="11" pageNumber="212" type="diagnosis">
<paragraph id="8BCFFC0DFFD5FFFC5394FDF7C1238E2B" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="212">
<emphasis id="B904201FFFD5FFFC5394FDF7C4538F99" bold="true" box="[199,321,616,641]" pageId="11" pageNumber="212">Diagnosis.</emphasis>
This species is similar to the other Japanese species of the subgenus
<taxonomicName id="4C70878EFFD5FFFC5719FDF7C1C08F98" box="[1098,1234,616,640]" class="Insecta" family="Staphylinidae" genus="Coprochara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="B904201FFFD5FFFC5719FDF7C1C08F98" box="[1098,1234,616,640]" italics="true" pageId="11" pageNumber="212">Coprochara</emphasis>
</taxonomicName>
, but can be easily distinguished from them by the following character states: surface smooth and glossy (
<figureCitation id="134BE088FFD5FFFC57D5FD13C1D98FBC" box="[1158,1227,652,676]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[278,1306,191,1972]" captionTargetId="figure@2.[278,1308,161,1975]" captionTargetPageId="2" captionText="FIGURES 1 4. Japanese species of Aleochara (Coprochara) in habitus. 1. Aleochara (Coprochara) verna; 2. A. (C.) binotata; 3 4. A. (C.) squalithorax: gray type (Fig. 3) and yellow type (Fig. 4)." httpUri="https://zenodo.org/record/223166/files/figure.png" pageId="11" pageNumber="212">Fig. 1</figureCitation>
); elytral posterior margin rounded with a pair of yellowish spots on dorsal surface of elytra (
<figureCitation id="134BE088FFD5FFFC5744FD30C1788FDF" box="[1047,1130,687,711]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[278,1306,191,1972]" captionTargetId="figure@2.[278,1308,161,1975]" captionTargetPageId="2" captionText="FIGURES 1 4. Japanese species of Aleochara (Coprochara) in habitus. 1. Aleochara (Coprochara) verna; 2. A. (C.) binotata; 3 4. A. (C.) squalithorax: gray type (Fig. 3) and yellow type (Fig. 4)." httpUri="https://zenodo.org/record/223166/files/figure.png" pageId="11" pageNumber="212">Figs. 1</figureCitation>
,
<figureCitation id="134BE088FFD5FFFC572EFD30C19F8FDF" box="[1149,1165,687,711]" captionStart="FIGURES 5 13" captionStartId="3.[151,264,1073,1096]" captionTargetBox="[163,1424,194,1036]" captionTargetId="figure@3.[151,1436,171,1052]" captionTargetPageId="3" captionText="FIGURES 5 13. Japanese species of Aleochara (Coprochara). 5 7. Aleochara (Coprochara) verna; 8 10. A. (C.) binotata; 11 13. A. (C.) squalithorax. (5, 8, 11: elytra; 6, 9, 12: intercoxal process of metaventrite; 7, 10, 13: mesoventrite and metaventrite)." httpUri="https://zenodo.org/record/223167/files/figure.png" pageId="11" pageNumber="212">5</figureCitation>
); intercoxal process of metaventrite moderately rounded apically (
<figureCitation id="134BE088FFD5FFFC5128FD4CC7FB8FF3" box="[635,745,723,747]" captionStart="FIGURES 5 13" captionStartId="3.[151,264,1073,1096]" captionTargetBox="[163,1424,194,1036]" captionTargetId="figure@3.[151,1436,171,1052]" captionTargetPageId="3" captionText="FIGURES 5 13. Japanese species of Aleochara (Coprochara). 5 7. Aleochara (Coprochara) verna; 8 10. A. (C.) binotata; 11 13. A. (C.) squalithorax. (5, 8, 11: elytra; 6, 9, 12: intercoxal process of metaventrite; 7, 10, 13: mesoventrite and metaventrite)." httpUri="https://zenodo.org/record/223167/files/figure.png" pageId="11" pageNumber="212">Figs. 67</figureCitation>
); tergite VIII without thick spines in posterior half (
<figureCitation id="134BE088FFD5FFFC5661FD4CC5A48E17" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="11" pageNumber="212">Figs. 14 15</figureCitation>
). [
<emphasis id="B904201FFFD5FFFC538BFD67C4078E08" bold="true" box="[216,277,760,784]" pageId="11" pageNumber="212">
<collectingCountry id="F367BC9DFFD5FFFC538BFD67C4078E08" box="[216,277,760,784]" name="Malta" pageId="11" pageNumber="212">Male</collectingCountry>
</emphasis>
]: sclerite Z inside the median lobe with long waving attachment at apex (
<figureCitation id="134BE088FFD5FFFC57D6FD68C0018E17" box="[1157,1299,759,783]" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="11" pageNumber="212">Figs. 1819</figureCitation>
). [
<emphasis id="B904201FFFD5FFFC5667FD67C0998E08" bold="true" box="[1332,1419,760,784]" pageId="11" pageNumber="212">Female</emphasis>
]: spermatheca with maximally 20 thin coils, not greatly extended laterally (
<figureCitation id="134BE088FFD5FFFC5082FC84C1368E2B" box="[977,1060,795,819]" captionStart="FIGURES 14 21" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[213,1379,182,1923]" captionTargetId="figure@5.[191,1401,159,1944]" captionTargetPageId="5" captionText="FIGURES 14 21. Terminalia of Aleochara (Coprochara) verna. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of paramerite, lateral view; 21. female genitalia: spermatheca." httpUri="https://zenodo.org/record/223168/files/figure.png" pageId="11" pageNumber="212">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BCFFC0DFFD5FFFC5392FCA1C4808E62" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="212">
<emphasis id="B904201FFFD5FFFC5392FCA1C7CC8E4F" bold="true" box="[193,734,830,855]" pageId="11" pageNumber="212">Confirmed distribution by the present study.</emphasis>
[
<emphasis id="B904201FFFD5FFFC51A3FCA1C65B8E4F" bold="true" box="[752,841,830,855]" pageId="11" pageNumber="212">
<collectingCountry id="F367BC9DFFD5FFFC51A3FCA1C65B8E4F" box="[752,841,830,855]" name="Japan" pageId="11" pageNumber="212">JAPAN</collectingCountry>
</emphasis>
]: Hokkaidô, Honshû, Shikoku, Kyûshû, [
<emphasis id="B904201FFFD5FFFC5677FCA1C0988E4F" bold="true" box="[1316,1418,830,855]" pageId="11" pageNumber="212">
<collectingCountry id="F367BC9DFFD5FFFC5677FCA1C0988E4F" box="[1316,1418,830,855]" name="Russia" pageId="11" pageNumber="212">RUSSIA</collectingCountry>
</emphasis>
]: Kamchatka Peninsula.
</paragraph>
</subSubSection>
<subSubSection id="C36AAF86FFD5FFFC5396FC1AC1CC89A4" pageId="11" pageNumber="212" type="description">
<paragraph id="8BCFFC0DFFD5FFFC5396FC1AC1CC89A4" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="212">
<emphasis id="B904201FFFD5FFFC5396FC1AC75D8E86" bold="true" box="[197,591,901,926]" pageId="11" pageNumber="212">Other localities in the literature.</emphasis>
[
<emphasis id="B904201FFFD5FFFC5131FC1AC7C48E86" bold="true" box="[610,726,901,926]" pageId="11" pageNumber="212">EUROPE</emphasis>
]:
<collectingCountry id="F367BC9DFFD5FFFC51A2FC19C6548E86" box="[753,838,902,926]" name="Austria" pageId="11" pageNumber="212">Austria</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5000FC19C6C38E86" box="[851,977,902,926]" name="Azerbaijan" pageId="11" pageNumber="212">Azerbaijan</collectingCountry>
, Azores,
<collectingCountry id="F367BC9DFFD5FFFC576DFC19C1B38E86" box="[1086,1185,902,926]" name="Belgium" pageId="11" pageNumber="212">Belgium</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC57FCFC19C08A8E86" box="[1199,1432,902,926]" name="Bosnia and Herzegovina" pageId="11" pageNumber="212">Bosnia Herzegovina</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC53C4FC35C5E88EDA" box="[151,250,938,962]" name="Bulgaria" pageId="11" pageNumber="212">Bulgaria</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC525BFC35C44F8EDA" box="[264,349,938,962]" name="Croatia" pageId="11" pageNumber="212">Croatia</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC523FFC35C7368EDA" box="[364,548,938,962]" name="Czech Republic" pageId="11" pageNumber="212">Czech Republic</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5161FC35C78F8EDA" box="[562,669,938,962]" name="Denmark" pageId="11" pageNumber="212">Denmark</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC51F8FC34C7E88EDA" box="[683,762,939,962]" name="France" pageId="11" pageNumber="212">France</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC505BFC35C6768EDA" box="[776,868,938,962]" name="Georgia" pageId="11" pageNumber="212">Georgia</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5021FC35C6C98EDA" box="[882,987,938,962]" name="Germany" pageId="11" pageNumber="212">Germany</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC50B9FC35C12E8EDA" box="[1002,1084,938,962]" name="Greece" pageId="11" pageNumber="212">Greece</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5719FC34C1BF8EDB" box="[1098,1197,939,963]" name="Hungary" pageId="11" pageNumber="212">Hungary</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC57E8FC35C1E28EDA" box="[1211,1264,938,962]" name="Italy" pageId="11" pageNumber="212">Italy</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC57ADFC35C07E8EDA" box="[1278,1388,938,962]" name="Lithuania" pageId="11" pageNumber="212">Lithuania</collectingCountry>
, the Nertherlands,
<collectingCountry id="F367BC9DFFD5FFFC5210FC51C4868EFE" box="[323,404,974,998]" name="Poland" pageId="11" pageNumber="212">Poland</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC52FEFC51C71D8EFE" box="[429,527,974,998]" name="Portugal" pageId="11" pageNumber="212">Portugal</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC517BFC51C7828EFE" box="[552,656,974,998]" name="Romania" pageId="11" pageNumber="212">Romania</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC51FBFC51C7E58EFE" box="[680,759,974,998]" name="Russia" pageId="11" pageNumber="212">Russia</collectingCountry>
: South European Territory,
<collectingCountry id="F367BC9DFFD5FFFC5730FC51C1DA8EFE" box="[1123,1224,974,998]" name="Slovakia" pageId="11" pageNumber="212">Slovakia</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC57B2FC51C0318EFE" box="[1249,1315,974,998]" name="Spain" pageId="11" pageNumber="212">Spain</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC566FFC51C0858EFE" box="[1340,1431,974,998]" name="Sweden" pageId="11" pageNumber="212">Sweden</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC53C4FC6EC4328911" box="[151,288,1009,1033]" name="Switzerland" pageId="11" pageNumber="212">Switzerland</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5278FC6DC4448911" box="[299,342,1010,1033]" name="United Kingdom" pageId="11" pageNumber="212">UK</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5230FC6EC4AF8911" box="[355,445,1009,1033]" name="Ukraine" pageId="11" pageNumber="212">Ukraine</collectingCountry>
(Maus, 1998; Smetana, 2004; Assing, 2006; Prado e Castro
<emphasis id="B904201FFFD5FFFC5738FC6DC18F8911" box="[1131,1181,1009,1033]" italics="true" pageId="11" pageNumber="212">et al</emphasis>
., 2010; Tamutis
<emphasis id="B904201FFFD5FFFC560FFC6DC09C8911" box="[1372,1422,1009,1033]" italics="true" pageId="11" pageNumber="212">et al</emphasis>
., 2011), [
<emphasis id="B904201FFFD5FFFC53A1FB8AC4C28936" bold="true" box="[242,464,1045,1070]" pageId="11" pageNumber="212">NORTH AFRICA</emphasis>
]:
<collectingCountry id="F367BC9DFFD5FFFC52BFFB8AC7518935" box="[492,579,1045,1069]" name="Algeria" pageId="11" pageNumber="212">Algeria</collectingCountry>
, Canary Islands,
<collectingCountry id="F367BC9DFFD5FFFC505CFB89C6478936" box="[783,853,1046,1070]" name="Egypt" pageId="11" pageNumber="212">Egypt</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5030FB8AC6B58935" box="[867,935,1045,1069]" name="Libya" pageId="11" pageNumber="212">Libya</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC50E4FB89C10D8935" box="[951,1055,1046,1069]" name="Morocco" pageId="11" pageNumber="212">Morocco</collectingCountry>
, Madeira Archipelago (Smetana, 2004), [
<emphasis id="B904201FFFD5FFFC53A1FBA7C4278949" bold="true" box="[242,309,1080,1105]" pageId="11" pageNumber="212">ASIA</emphasis>
]:
<collectingCountry id="F367BC9DFFD5FFFC5203FBA6C4CE8949" box="[336,476,1081,1105]" name="Afghanistan" pageId="11" pageNumber="212">Afghanistan</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC52B9FBA6C7228949" box="[490,560,1081,1105]" name="China" pageId="11" pageNumber="212">China</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC516DFBA6C7688949" box="[574,634,1081,1105]" name="India" pageId="11" pageNumber="212">India</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC51DBFBA5C7AA8949" box="[648,696,1082,1105]" name="Iran" pageId="11" pageNumber="212">Iran</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5195FBA6C6158949" box="[710,775,1081,1105]" name="Israel" pageId="11" pageNumber="212">Israel</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5046FBA6C6898949" box="[789,923,1081,1105]" name="Kazakhstan" pageId="11" pageNumber="212">Kazakhstan</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC50FAFBA6C10A8949" box="[937,1048,1081,1105]" name="Mongolia" pageId="11" pageNumber="212">Mongolia</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5775FBA6C17E8949" box="[1062,1132,1081,1105]" name="Nepal" pageId="11" pageNumber="212">Nepal</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5729FBA6C01E8949" box="[1146,1292,1081,1105]" name="North Korea" pageId="11" pageNumber="212">North Korea</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5649FBA6C07B8949" box="[1306,1385,1081,1105]" name="Russia" pageId="11" pageNumber="212">Russia</collectingCountry>
: Far East,
<collectingCountry id="F367BC9DFFD5FFFC5385FBC3C47A896C" box="[214,360,1116,1140]" name="South Korea" pageId="11" pageNumber="212">South Korea</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5227FBC3C4A0896C" box="[372,434,1116,1140]" name="Syria" pageId="11" pageNumber="212">Syria</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC52EDFBC3C73D896C" box="[446,559,1116,1140]" name="Tajikistan" pageId="11" pageNumber="212">Tajikistan</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5168FBC3C7CB896C" box="[571,729,1116,1140]" name="Turkmenistan" pageId="11" pageNumber="212">Turkmenistan</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC51B5FBC3C625896C" box="[742,823,1116,1140]" name="Turkey" pageId="11" pageNumber="212">Turkey</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC5010FBC3C6D3896C" box="[835,961,1116,1140]" name="Uzbekistan" pageId="11" pageNumber="212">Uzbekistan</collectingCountry>
(Maus, 1998; Smetana, 2004; Park
<emphasis id="B904201FFFD5FFFC560FFBC1C09C896D" box="[1372,1422,1117,1141]" italics="true" pageId="11" pageNumber="212">et al</emphasis>
., 2011), [
<emphasis id="B904201FFFD5FFFC53A1FB1FC4FE8981" bold="true" box="[242,492,1152,1177]" pageId="11" pageNumber="212">
NORTH
<collectingCountry id="F367BC9DFFD5FFFC5230FB1FC4FE8981" box="[355,492,1152,1177]" name="United States of America" pageId="11" pageNumber="212">AMERICA</collectingCountry>
</emphasis>
]: Alaska,
<collectingCountry id="F367BC9DFFD5FFFC513BFB1FC7AD8980" box="[616,703,1152,1176]" name="Canada" pageId="11" pageNumber="212">Canada</collectingCountry>
, Hawaii,
<collectingCountry id="F367BC9DFFD5FFFC5062FB1FC69B8980" box="[817,905,1152,1176]" name="Mexico" pageId="11" pageNumber="212">Mexico</collectingCountry>
,
<collectingCountry id="F367BC9DFFD5FFFC50CBFB1FC6C28980" box="[920,976,1152,1176]" name="United States of America" pageId="11" pageNumber="212">USA</collectingCountry>
(Klimaszewski, 1984; Navarrete
<emphasis id="B904201FFFD5FFFC560AFB1EC09C8980" box="[1369,1422,1152,1176]" italics="true" pageId="11" pageNumber="212">et al</emphasis>
., 2002; Gouix &amp; Klimaszewski, 2007), [
<emphasis id="B904201FFFD5FFFC511FFB3BC62D89A5" bold="true" box="[588,831,1188,1213]" pageId="11" pageNumber="212">
SOUTH
<collectingCountry id="F367BC9DFFD5FFFC51E5FB3BC62D89A5" box="[694,831,1188,1213]" name="United States of America" pageId="11" pageNumber="212">AMERICA</collectingCountry>
</emphasis>
]:
<collectingCountry id="F367BC9DFFD5FFFC5005FB3BC6DB89A4" box="[854,969,1188,1212]" name="Venezuela" pageId="11" pageNumber="212">Venezuela</collectingCountry>
(Navarrete
<emphasis id="B904201FFFD5FFFC571CFB3AC19389A4" box="[1103,1153,1188,1212]" italics="true" pageId="11" pageNumber="212">et al</emphasis>
., 2002).
</paragraph>
</subSubSection>
<subSubSection id="C36AAF86FFD5FFFB5394FB57C08F8F47" lastPageId="12" lastPageNumber="213" pageId="11" pageNumber="212" type="discussion">
<paragraph id="8BCFFC0DFFD5FFFC5394FB57C7B18877" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="212">
<emphasis id="B904201FFFD5FFFC5394FB57C42989F8" bold="true" box="[199,315,1224,1248]" pageId="11" pageNumber="212">Remarks.</emphasis>
<taxonomicName id="4C70878EFFD5FFFC5217FB57C71389F8" box="[324,513,1224,1248]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD5FFFC5217FB57C71389F8" box="[324,513,1224,1248]" italics="true" pageId="11" pageNumber="212">Aleochara verna</emphasis>
</taxonomicName>
is very common and widely distributed in the Holarctic region. The distribution ranges through all of Europe and North Africa, across northern
<collectingCountry id="F367BC9DFFD5FFFC503DFB74C6B5881B" box="[878,935,1259,1283]" name="India" pageId="11" pageNumber="212">India</collectingCountry>
and the Russian Far East to North
<collectingCountry id="F367BC9DFFD5FFFC5668FB74C089881B" box="[1339,1435,1259,1283]" name="United States of America" pageId="11" pageNumber="212">America</collectingCountry>
(Maus, 1998; Smetana, 2004). In
<collectingCountry id="F367BC9DFFD5FFFC514AFA8FC74E883F" box="[537,604,1296,1319]" name="Japan" pageId="11" pageNumber="212">Japan</collectingCountry>
, reliable records had been reported from only two localities, Gunma and Kanagawa Prefectures (Maus, 1998; Park
<emphasis id="B904201FFFD5FFFC51DBFAABC7AC8853" box="[648,702,1331,1355]" italics="true" pageId="11" pageNumber="212">et al</emphasis>
., 2011). Numerous records are added from all of
<collectingCountry id="F367BC9DFFD5FFFC5640FAABC0448853" box="[1299,1366,1332,1355]" name="Japan" pageId="11" pageNumber="212">Japan</collectingCountry>
, from northern Hokkaidô to southern Kyûshû,
<collectingCountry id="F367BC9DFFD5FFFC5108FAC7C78D8877" box="[603,671,1368,1391]" name="Japan" pageId="11" pageNumber="212">Japan</collectingCountry>
.
</paragraph>
<paragraph id="8BCFFC0DFFD5FFFC5394FAE5C15F88C2" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="212">
This species was originally described from Missouri,
<collectingCountry id="F367BC9DFFD5FFFC5074FAE5C671888A" box="[807,867,1402,1426]" name="United States of America" pageId="11" pageNumber="212">USA</collectingCountry>
.
<collectingCountry id="F367BC9DFFD5FFFC503DFAE4C69F888B" box="[878,909,1403,1427]" name="American Samoa" pageId="11" pageNumber="212">As</collectingCountry>
the
<typeStatus id="54CB42AFFFD5FFFC50ECFAE4C6FD888B" box="[959,1007,1403,1427]" pageId="11" pageNumber="212">type</typeStatus>
specimen was destroyed, the
<typeStatus id="54CB42AFFFD5FFFC5610FAE4C08E888B" box="[1347,1436,1403,1427]" pageId="11" pageNumber="212" type="neotype">neotype</typeStatus>
was designated by Klimaszewski (1984). The distribution of this species in the Palearctic region is unclear; it has been suggested that
<taxonomicName id="4C70878EFFD5FFFC522AFA5DC4CA88C2" box="[377,472,1474,1498]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD5FFFC522AFA5DC4CA88C2" box="[377,472,1474,1498]" italics="true" pageId="11" pageNumber="212">A. verna</emphasis>
</taxonomicName>
is not a Nearctic, but a Palearctic species (Maus, 1998).
</paragraph>
<paragraph id="8BCFFC0DFFD5FFFC5394FA79C4FD8B94" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="212">
The taxonomic status of
<taxonomicName id="4C70878EFFD5FFFC5289FA79C72B88E6" box="[474,569,1510,1534]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD5FFFC5289FA79C72B88E6" box="[474,569,1510,1534]" italics="true" pageId="11" pageNumber="212">A. verna</emphasis>
</taxonomicName>
was unstable until Lohse (1986) and thus far several cases of confusion have occurred, e.g., misidentifications of species in Europe and the proposed use of many synonyms for some species and subspecies under
<taxonomicName id="4C70878EFFD5FFFC52C5F9B2C4EB8B5D" box="[406,505,1581,1605]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD5FFFC52C5F9B2C4EB8B5D" box="[406,505,1581,1605]" italics="true" pageId="11" pageNumber="212">A. verna</emphasis>
</taxonomicName>
(Klimaszewski, 1984; Welch 1997; Maus, 1998). Recent studies such as Welch (1990, 1997) and Maus (1998) have allowed the precise identification of the
<taxonomicName id="4C70878EFFD5FFFC575CF9CEC1858B71" box="[1039,1175,1617,1641]" class="Insecta" family="Staphylinidae" genus="Coprochara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="B904201FFFD5FFFC575CF9CEC1858B71" box="[1039,1175,1617,1641]" italics="true" pageId="11" pageNumber="212">Coprochara</emphasis>
</taxonomicName>
-complex, including
<taxonomicName id="4C70878EFFD5FFFC56D7F9CEC5C58B95" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD5FFFC56D7F9CEC5C58B95" italics="true" pageId="11" pageNumber="212">A. verna</emphasis>
</taxonomicName>
, in the Palearctic region.
</paragraph>
<paragraph id="8BCFFC0DFFD5FFFC5394F907C4458A27" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="212">
The Japanese specimens examined have a blackish body with a pair of small yellow markings on the elytra; no geographical variations were detected. This species is commonly found in
<collectingCountry id="F367BC9DFFD5FFFC5089F922C10F8BCC" box="[986,1053,1725,1748]" name="Japan" pageId="11" pageNumber="212">Japan</collectingCountry>
; many records have been reported as “
<taxonomicName id="4C70878EFFD5FFFC5394F97FC4D48BE0" box="[199,454,1760,1784]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bipustulata">
<emphasis id="B904201FFFD5FFFC5394F97FC4D48BE0" box="[199,454,1760,1784]" italics="true" pageId="11" pageNumber="212">Aleochara bipustulata</emphasis>
</taxonomicName>
”. However, true specimens of
<taxonomicName id="4C70878EFFD5FFFC506FF97FC6CC8BE0" box="[828,990,1760,1784]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bipustulata">
<emphasis id="B904201FFFD5FFFC506FF97FC6CC8BE0" box="[828,990,1760,1784]" italics="true" pageId="11" pageNumber="212">A. bipustulata</emphasis>
</taxonomicName>
were not found in the present study. Therefore, the distribution of
<taxonomicName id="4C70878EFFD5FFFC52B1F89BC76D8A04" box="[482,639,1796,1820]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bipustulata">
<emphasis id="B904201FFFD5FFFC52B1F89BC76D8A04" box="[482,639,1796,1820]" italics="true" pageId="11" pageNumber="212">A. bipustulata</emphasis>
</taxonomicName>
in
<collectingCountry id="F367BC9DFFD5FFFC51F0F89BC7F18A03" box="[675,739,1796,1819]" name="Japan" pageId="11" pageNumber="212">Japan</collectingCountry>
is doubtful and it should be excluded from the Japanese fauna (see Discussion).
</paragraph>
<paragraph id="8BCFFC0DFFD5FFFC5394F8D4C04A8A9F" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="212">
Park
<emphasis id="B904201FFFD5FFFC5250F8D3C4258A7B" box="[259,311,1867,1891]" italics="true" pageId="11" pageNumber="212">et al</emphasis>
. (2011) redescribed and recorded
<taxonomicName id="4C70878EFFD5FFFC5193F8D4C6338A7B" box="[704,801,1867,1891]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD5FFFC5193F8D4C6338A7B" box="[704,801,1867,1891]" italics="true" pageId="11" pageNumber="212">A. verna</emphasis>
</taxonomicName>
from
<collectingCountry id="F367BC9DFFD5FFFC5039F8D4C6EE8A7B" box="[874,1020,1867,1891]" name="South Korea" pageId="11" pageNumber="212">South Korea</collectingCountry>
, with diagnoses of
<taxonomicName id="4C70878EFFD5FFFC578FF8D4C02F8A7B" box="[1244,1341,1867,1891]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD5FFFC578FF8D4C02F8A7B" box="[1244,1341,1867,1891]" italics="true" pageId="11" pageNumber="212">A. verna</emphasis>
</taxonomicName>
and “
<taxonomicName id="4C70878EFFD5FFFC56D7F8D4C4078A9F" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bipustulata">
<emphasis id="B904201FFFD5FFFC56D7F8D4C4078A9F" italics="true" pageId="11" pageNumber="212">A. bipustulata</emphasis>
</taxonomicName>
”. Following the Japanese example, Korean specimens of “
<taxonomicName id="4C70878EFFD5FFFC50F9F8F0C1558A9F" box="[938,1095,1903,1927]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bipustulata">
<emphasis id="B904201FFFD5FFFC50F9F8F0C1558A9F" box="[938,1095,1903,1927]" italics="true" pageId="11" pageNumber="212">A. bipustulata</emphasis>
</taxonomicName>
” should be reexamined.
</paragraph>
<paragraph id="8BCFFC0DFFD5FFFC5394F80CC4828AEA" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="212">
In
<collectingCountry id="F367BC9DFFD5FFFC53B9F80CC43F8AB2" box="[234,301,1939,1962]" name="Japan" pageId="11" pageNumber="212">Japan</collectingCountry>
,
<taxonomicName id="4C70878EFFD5FFFC526EF80CC4EC8AB3" box="[317,510,1939,1963]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD5FFFC526EF80CC4EC8AB3" box="[317,510,1939,1963]" italics="true" pageId="11" pageNumber="212">Aleochara verna</emphasis>
</taxonomicName>
might be the most efficient candidate as a biological control agent due to its abundance, and wide distribution range, and the availability of information that has been accumulated in previous studies on the species.
</paragraph>
<paragraph id="8BCFFC0DFFD2FFFB5394FF08C6E88CB4" blockId="12.[151,1437,151,607]" pageId="12" pageNumber="213">
<emphasis id="B904201FFFD2FFFB5394FF08C4588DA8" bold="true" box="[199,330,151,176]" pageId="12" pageNumber="213">Bionomics.</emphasis>
Species of the subgenus
<taxonomicName id="4C70878EFFD2FFFB5139FF07C7E08DA8" box="[618,754,152,176]" class="Insecta" family="Staphylinidae" genus="Coprochara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B904201FFFD2FFFB5139FF07C7E08DA8" box="[618,754,152,176]" italics="true" pageId="12" pageNumber="213">Coprochara</emphasis>
</taxonomicName>
occur predominantly in the dung of herbivorous vertebrates and in a wide range of organic materials (Klimaszewski &amp; Jansen, 1994).
<taxonomicName id="4C70878EFFD2FFFB5740FF23C1C68DCC" box="[1043,1236,188,212]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD2FFFB5740FF23C1C68DCC" box="[1043,1236,188,212]" italics="true" pageId="12" pageNumber="213">Aleochara verna</emphasis>
</taxonomicName>
is considered as inhabiting in the same environment. Adults were collected from human feces and other animal droppings such as cow, horse, dog, and woodchuck, as well as carrion (Klimaszewski, 1984). The first author collected the species from cow dung in a pasture at Aso, Kumamoto Prefecture,
<collectingCountry id="F367BC9DFFD2FFFB5063FEB7C6618C27" box="[816,883,296,319]" name="Japan" pageId="12" pageNumber="213">Japan</collectingCountry>
. He also confirmed that
<taxonomicName id="4C70878EFFD2FFFB57D6FEB7C1F78C58" box="[1157,1253,296,320]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD2FFFB57D6FEB7C1F78C58" box="[1157,1253,296,320]" italics="true" pageId="12" pageNumber="213">A. verna</emphasis>
</taxonomicName>
was attracted to small prawns (leavings from a packed lunch) on a walkway in 2009 at Mt. Sobo,
<collectingCountry id="F367BC9DFFD2FFFB5729FED2C1AF8C7C" box="[1146,1213,333,356]" name="Japan" pageId="12" pageNumber="213">Japan</collectingCountry>
. SY also collected specimens of this species on a walkway in summer 2011 on the same mountain, and observed dozens of individuals walking actively on rocks and pebbly surface on the summit (
<figureCitation id="134BE088FFD2FFFB501BFE0BC68E8CB4" box="[840,924,404,428]" captionStart="FIGURES 42 43" captionStartId="17.[151,264,1257,1280]" captionTargetBox="[157,1430,811,1230]" captionTargetId="figure@17.[151,1436,807,1235]" captionTargetPageId="17" captionText="FIGURES 42 43. Habitats of the subgenus Coprochara of the genus Aleochara in Japan. 93. the summit of Sobo-san (located in boundaries between Ôita and Miyazaki Prefectures; only observed; A. (C.) verna: arrow); 43. Watari (Fukutsu, Fukuoka Prefecture; A. (C.) squalithorax: arrow)." httpUri="https://zenodo.org/record/223172/files/figure.png" pageId="12" pageNumber="213">Fig. 42</figureCitation>
: arrow).
</paragraph>
<paragraph id="8BCFFC0DFFD2FFFB5394FE28C45D8F0F" blockId="12.[151,1437,151,607]" pageId="12" pageNumber="213">
The altitudinal range of
<taxonomicName id="4C70878EFFD2FFFB5287FE27C7208CC8" box="[468,562,440,464]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD2FFFB5287FE27C7208CC8" box="[468,562,440,464]" italics="true" pageId="12" pageNumber="213">A. verna</emphasis>
</taxonomicName>
is quite wide. Klimaszewski (1984) noted that adult specimens were collected at altitudes from less than
<quantity id="4C8851E8FFD2FFFB5293FE43C71A8CEC" box="[448,520,476,500]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="213" unit="m" value="200.0">200 m</quantity>
to nearly
<quantity id="4C8851E8FFD2FFFB5129FE43C7C88CEC" box="[634,730,476,500]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="213" unit="m" value="4000.0">4,000 m</quantity>
. Japanese specimens were found at altitudes from 0 (seashore) to over
<quantity id="4C8851E8FFD2FFFB53BFFE60C4598F0F" box="[236,331,511,535]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="213" unit="m" value="2000.0">2,000 m</quantity>
.
</paragraph>
<paragraph id="8BCFFC0DFFD2FFFB5394FDBBC08F8F47" blockId="12.[151,1437,151,607]" pageId="12" pageNumber="213">
<emphasis id="B904201FFFD2FFFB5394FDBBC47A8F24" bold="true" box="[199,360,548,572]" pageId="12" pageNumber="213">Host records.</emphasis>
Comprehensive dipteran host records for
<taxonomicName id="4C70878EFFD2FFFB5018FDBBC6BE8F24" box="[843,940,548,572]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="verna">
<emphasis id="B904201FFFD2FFFB5018FDBBC6BE8F24" box="[843,940,548,572]" italics="true" pageId="12" pageNumber="213">A. verna</emphasis>
</taxonomicName>
were provided by Maus
<emphasis id="B904201FFFD2FFFB579FFDBAC1ED8F24" box="[1228,1279,548,572]" italics="true" pageId="12" pageNumber="213">et al</emphasis>
. (1998). This paper gives the following four families as its hosts:
<taxonomicName id="4C70878EFFD2FFFB518AFDD8C66E8F47" box="[729,892,583,607]" class="Insecta" family="Anthomyiidae" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="family">Anthomyiidae</taxonomicName>
,
<taxonomicName id="4C70878EFFD2FFFB50D4FDD8C1298F47" box="[903,1083,583,607]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
,
<taxonomicName id="4C70878EFFD2FFFB5715FDD8C1A48F47" box="[1094,1206,583,607]" class="Insecta" family="Muscidae" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="family">Muscidae</taxonomicName>
, and
<taxonomicName id="4C70878EFFD2FFFB57A3FDD8C08B8F47" box="[1264,1433,583,607]" class="Insecta" family="Sarcophagidae" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="family">Sarcophagidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>