286 lines
35 KiB
XML
286 lines
35 KiB
XML
<document id="7ED1971ED76CEAD749FA6176EB927AEC" ID-CLB-Dataset="20176" ID-DOI="10.11646/zootaxa.5116.2.5" ID-GBIF-Dataset="dc6e6f10-3b06-448e-bf5f-8bc181884366" ID-ISSN="1175-5326" ID-Zenodo-Dep="6367323" ID-ZooBank="2558C8B0-FABB-4104-B103-56C0830AF277" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647595653746" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V., Chigray, Ivan A., Ntatsopoulos, Konstantinos & Papadopoulou, Anna" docDate="2022" docId="03D0A51AFFD8FFB89C97ACB3CDAA9123" docLanguage="en" docName="zootaxa.5116.2.5.pdf" docOrigin="Zootaxa 5116 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blaps verrucosa Adams 1817" docType="treatment" docVersion="7" lastPageNumber="286" masterDocId="FFE9DD62FFCAFFAB9C00A841CF16931E" masterDocTitle="A key to Russian and Eastern European species of Blaps Fabricius, 1775 (Coleoptera: Tenebrionidae: Blaptinae) with the description of a new species from the North Caucasus supported by morphological and molecular data" masterLastPageNumber="291" masterPageNumber="267" pageNumber="285" updateTime="1699318281145" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="35762AB6765A3B640D3B663B505F41D0">A key to Russian and Eastern European species of Blaps Fabricius, 1775 (Coleoptera: Tenebrionidae: Blaptinae) with the description of a new species from the North Caucasus supported by morphological and molecular data</mods:title>
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<mods:namePart id="9BDFD344334CD3C58730A8ED784B614D">Nabozhenko, Maxim V.</mods:namePart>
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<mods:affiliation id="BB45864D650BA06C712B051F06526978">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000, Russia. & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000, Russia & A. N. Severtsov Institute of ecology and evolution of the Russian Academy of Sciences, Leninsky Prospect 33, 119071 Moscow, Russia Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb., 1, Saint Petersburg 199034 Russia. chigray 93 @ bk. ru; https: // orcid. org / 0000 - 0002 - 3830 - 6860</mods:affiliation>
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<mods:namePart id="5EDBF91DC5A29D094AAAE2D62B647104">Chigray, Ivan A.</mods:namePart>
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<mods:nameIdentifier id="DB14EE3AC0DB103AC9D16C73CC459855" type="email">chigray93@bk.ru</mods:nameIdentifier>
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<mods:namePart id="BAB3DFACA21BAD1D27D099141BE1A533">Ntatsopoulos, Konstantinos</mods:namePart>
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<mods:affiliation id="8C447DC539CD0E2BB5DAAAFCE118D015">Department of Biological Sciences, University of Cyprus, 1678 Nicosia P. O. Box 20537, Cyprus. & ntatsopoulos. konstantinos @ ucy. ac. cy; https: // orcid. org / 0000 - 0003 - 0673 - 2416</mods:affiliation>
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<mods:namePart id="7DB4D064D0ACA7720C7D47BE959CAB44">Papadopoulou, Anna</mods:namePart>
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<mods:affiliation id="E320F932990533EFB8F11D5F1B2CD14B">Department of Biological Sciences, University of Cyprus, 1678 Nicosia P. O. Box 20537, Cyprus. & papadopoulou. g. anna @ ucy. ac. cy; https: // orcid. org / 0000 - 0002 - 4656 - 4894</mods:affiliation>
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<mods:nameIdentifier id="84BB4BA0ECE037DA76902B6009F024D5" type="email">papadopoulou.g.anna@ucy.ac.cy</mods:nameIdentifier>
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<mods:title id="C90A5C3DE600B4636837812066AAD595">Zootaxa</mods:title>
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<mods:date id="F0826573C08F10232B81730F5F90E9CA">2022</mods:date>
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<mods:number id="8086CDBE8DB338F7FCF2650BE0207B4C">5116</mods:number>
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<treatment id="03D0A51AFFD8FFB89C97ACB3CDAA9123" ID-DOI="http://doi.org/10.5281/zenodo.6369145" ID-GBIF-Taxon="194098882" ID-Zenodo-Dep="6369145" LSID="urn:lsid:plazi:treatment:03D0A51AFFD8FFB89C97ACB3CDAA9123" httpUri="http://treatment.plazi.org/id/03D0A51AFFD8FFB89C97ACB3CDAA9123" lastPageId="19" lastPageNumber="286" pageId="18" pageNumber="285">
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<subSubSection id="C3634787FFD8FFB99C97ACB3CD159612" box="[151,515,1266,1293]" pageId="18" pageNumber="285" type="nomenclature">
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<paragraph id="8BC6140CFFD8FFB99C97ACB3CD159612" blockId="18.[151,515,1266,1329]" box="[151,515,1266,1293]" pageId="18" pageNumber="285">
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<heading id="D08EA360FFD8FFB99C97ACB3CD159612" bold="true" box="[151,515,1266,1293]" fontSize="11" level="1" pageId="18" pageNumber="285" reason="1">
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<taxonomicName id="4C796F8FFFD8FFB99C97ACB3CD159612" authority="Adams, 1817" authorityName="Adams" authorityYear="1817" box="[151,515,1266,1293]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="285" phylum="Arthropoda" rank="species" species="verrucosa">
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<emphasis id="B90DC81EFFD8FFB99C97ACB3CD159612" bold="true" box="[151,515,1266,1293]" pageId="18" pageNumber="285">
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<emphasis id="B90DC81EFFD8FFB99C97ACB3CE4C9613" bold="true" box="[151,346,1266,1293]" italics="true" pageId="18" pageNumber="285">Blaps verrucosa</emphasis>
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Adams, 1817
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C3634787FFD8FFB99C97AD57CE58962F" box="[151,334,1302,1329]" pageId="18" pageNumber="285" type="description">
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<paragraph id="8BC6140CFFD8FFB99C97AD57CE58962F" blockId="18.[151,515,1266,1329]" box="[151,334,1302,1329]" pageId="18" pageNumber="285">
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(
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<figureCitation id="13420889FFD8FFB99C9FAD57CFFE962E" box="[159,232,1302,1328]" captionStart="FIGURE 6" captionStartId="5.[151,250,2010,2034]" captionTargetBox="[153,1432,1238,1984]" captionTargetId="figure-309@5.[151,1436,1236,1986]" captionTargetPageId="5" captionText="FIGURE 6. B. verrucosa, habitus. A, B = ♂; C = ♀; A, C = dorsal view; B = ventral view." figureDoi="http://doi.org/10.5281/zenodo.6367339" httpUri="https://zenodo.org/record/6367339/files/figure.png" pageId="18" pageNumber="285">Figs 6</figureCitation>
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,
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<figureCitation id="13420889FFD8FFB99CF4AD57CE53962F" box="[244,325,1302,1329]" captionStart="FIGURE 23" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[162,1430,186,1912]" captionTargetId="figure-23@20.[154,1433,181,1920]" captionTargetPageId="20" captionText="FIGURE 23. Blaps spp., habitus. A = non-rugose male of B. verrucosa from Baku, Azerbaijan, (the type locality of B. scabriuscula) ZM MSU; B = rugose male of B. verrucosa from Georgia (Kazbegi), ZIN; C = lectotype labels of B. subalpina, D = lectotype of B. subalpina, male (ZIN); E = lectotype labels of B. pterotapha; F = lectotype of B. pterotapha, female (ZIN)." figureDoi="http://doi.org/10.5281/zenodo.6367379" httpUri="https://zenodo.org/record/6367379/files/figure.png" pageId="18" pageNumber="285">23A, B</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3634787FFD8FFB99C97AD1ACC2F94B3" pageId="18" pageNumber="285" type="reference_group">
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<paragraph id="8BC6140CFFD8FFB99C97AD1ACD72966D" blockId="18.[151,612,1371,1425]" box="[151,612,1371,1395]" pageId="18" pageNumber="285">
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<emphasis id="B90DC81EFFD8FFB99C97AD1ACFB3966D" bold="true" box="[151,165,1371,1395]" pageId="18" pageNumber="285">=</emphasis>
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||
<taxonomicName id="4C796F8FFFD8FFB99CACAD1ACD0F966D" authority="Menetries, 1832" authorityName="Menetries" authorityYear="1832" box="[172,537,1371,1395]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="285" phylum="Arthropoda" rank="species" species="scabriuscula">
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<emphasis id="B90DC81EFFD8FFB99CACAD1ACE7B966D" box="[172,365,1371,1395]" italics="true" pageId="18" pageNumber="285">Blaps scabriuscula</emphasis>
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<bibRefCitation id="EFE869FDFFD8FFB99D74AD1ACD0F966D" author="Menetries, E." box="[372,537,1371,1395]" pageId="18" pageNumber="285" refId="ref14478" refString="Menetries, E. (1832) Catalogue raissonne des objets de Zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse. St. Petersbourg, L'Imprimerie de l'Academie Imperiale des Sciences, 272 pp., 5 pls. https: // doi. org / 10.5962 / bhl. title. 51784" type="book" year="1832">Ménétriés, 1832</bibRefCitation>
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</taxonomicName>
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||
<emphasis id="B90DC81EFFD8FFB99E1FAD1ACD72966D" bold="true" box="[543,612,1371,1395]" pageId="18" pageNumber="285">
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<taxonomicNameLabel id="A23E7565FFD8FFB99E1FAD1ACD72966D" box="[543,612,1371,1395]" pageId="18" pageNumber="285">syn. n.</taxonomicNameLabel>
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</emphasis>
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||
</paragraph>
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||
<paragraph id="8BC6140CFFD8FFB99C97AD38CD4E968F" blockId="18.[151,612,1371,1425]" box="[151,600,1401,1425]" pageId="18" pageNumber="285">
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<emphasis id="B90DC81EFFD8FFB99C97AD38CFB3968F" bold="true" box="[151,165,1401,1425]" pageId="18" pageNumber="285">=</emphasis>
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<taxonomicName id="4C796F8FFFD8FFB99CACAD38CD1A968F" authority="Motschulsky, 1839" authorityName="Motschulsky" authorityYear="1839" box="[172,524,1401,1425]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="285" phylum="Arthropoda" rank="species" species="montana">
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<emphasis id="B90DC81EFFD8FFB99CACAD38CE53968F" box="[172,325,1401,1425]" italics="true" pageId="18" pageNumber="285">Blaps montana</emphasis>
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<bibRefCitation id="EFE869FDFFD8FFB99D4BAD38CD1A968F" author="Motschulsky, V. de" box="[331,524,1401,1425]" pageId="18" pageNumber="285" pagination="44 - 93" refId="ref14537" refString="Motschulsky, V. de (1839) Insectes du Caucase et des Provinces Transcaucausiennes. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 12, 44 - 93, pls. I, II, V, VI." type="journal article" year="1839">Motschulsky, 1839</bibRefCitation>
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</taxonomicName>
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<emphasis id="B90DC81EFFD8FFB99E12AD38CD4E968F" bold="true" box="[530,600,1401,1425]" pageId="18" pageNumber="285">
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<taxonomicNameLabel id="A23E7565FFD8FFB99E12AD38CD4E968F" box="[530,600,1401,1425]" pageId="18" pageNumber="285">syn. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8BC6140CFFD8FFB99C97ADFECD669478" blockId="18.[151,1437,1471,1966]" pageId="18" pageNumber="285">
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<taxonomicName id="4C796F8FFFD8FFB99C97ADFECE4496C7" authorityName="Adams" authorityYear="1817" box="[151,338,1471,1497]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="285" phylum="Arthropoda" rank="species" species="verrucosa">
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<emphasis id="B90DC81EFFD8FFB99C97ADFECE4496C7" box="[151,338,1471,1497]" italics="true" pageId="18" pageNumber="285">Blaps verrucosa</emphasis>
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</taxonomicName>
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was described from the surroundings of
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<collectingRegion id="49BDDAEEFFD8FFB99F3AADFECC6096C7" box="[826,886,1471,1497]" country="Georgia" name="Tbilisi" pageId="18" pageNumber="285">Tiflis</collectingRegion>
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(now
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<collectingRegion id="49BDDAEEFFD8FFB99FC5ADFECB0796C7" box="[965,1041,1471,1497]" country="Georgia" name="Tbilisi" pageId="18" pageNumber="285">Tbilisi</collectingRegion>
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,
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<collectingCountry id="F36E549CFFD8FFB99821ADFECB9696C4" box="[1057,1152,1471,1498]" name="Georgia" pageId="18" pageNumber="285">Georgia</collectingCountry>
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). Adams’s description is detailed and completely corresponds to
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<taxonomicName id="4C796F8FFFD8FFB99E68ADA2CB1096E0" authority="Menetries, 1832" authorityName="Menetries" authorityYear="1832" box="[616,1030,1507,1534]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="285" phylum="Arthropoda" rank="species" species="scabriuscula">
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||
<emphasis id="B90DC81EFFD8FFB99E68ADA2CC5496E3" box="[616,834,1507,1533]" italics="true" pageId="18" pageNumber="285">Blaps scabriuscula</emphasis>
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<bibRefCitation id="EFE869FDFFD8FFB99F4DADA2CB1096E0" author="Menetries, E." box="[845,1030,1507,1534]" pageId="18" pageNumber="285" refId="ref14478" refString="Menetries, E. (1832) Catalogue raissonne des objets de Zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse. St. Petersbourg, L'Imprimerie de l'Academie Imperiale des Sciences, 272 pp., 5 pls. https: // doi. org / 10.5962 / bhl. title. 51784" type="book" year="1832">Ménétriés, 1832</bibRefCitation>
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</taxonomicName>
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described from
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||
<collectingRegion id="49BDDAEEFFD8FFB998C9ADA2CA1296E3" box="[1225,1284,1507,1533]" country="Azerbaijan" name="Baku" pageId="18" pageNumber="285">Baku</collectingRegion>
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(
|
||
<collectingCountry id="F36E549CFFD8FFB99916ADA2CA8396E0" box="[1302,1429,1507,1534]" name="Azerbaijan" pageId="18" pageNumber="285">Azerbaijan</collectingCountry>
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) (
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||
<bibRefCitation id="EFE869FDFFD8FFB99C9FAE46CE4E953C" author="Menetries, E." box="[159,344,1543,1570]" pageId="18" pageNumber="285" refId="ref14478" refString="Menetries, E. (1832) Catalogue raissonne des objets de Zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse. St. Petersbourg, L'Imprimerie de l'Academie Imperiale des Sciences, 272 pp., 5 pls. https: // doi. org / 10.5962 / bhl. title. 51784" type="book" year="1832">Ménétriés 1832</bibRefCitation>
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) and
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<taxonomicName id="4C796F8FFFD8FFB99DA7AE49CC0A953C" authority="Motschulsky, 1839" authorityName="Motschulsky" authorityYear="1839" box="[423,796,1543,1570]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="285" phylum="Arthropoda" rank="species" species="montana">
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||
<emphasis id="B90DC81EFFD8FFB99DA7AE49CD26953F" box="[423,560,1544,1569]" italics="true" pageId="18" pageNumber="285">B. montana</emphasis>
|
||
<bibRefCitation id="EFE869FDFFD8FFB99E3FAE46CC0A953C" author="Motschulsky, V. de" box="[575,796,1543,1570]" pageId="18" pageNumber="285" pagination="44 - 93" refId="ref14537" refString="Motschulsky, V. de (1839) Insectes du Caucase et des Provinces Transcaucausiennes. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 12, 44 - 93, pls. I, II, V, VI." type="journal article" year="1839">Motschulsky, 1839</bibRefCitation>
|
||
</taxonomicName>
|
||
described from Kurush (Southern
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||
<collectingRegion id="49BDDAEEFFD8FFB998CAAE49CA23953C" box="[1226,1333,1544,1570]" country="Russia" name="Dagestan" pageId="18" pageNumber="285">Dagestan</collectingRegion>
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,
|
||
<collectingCountry id="F36E549CFFD8FFB99948AE46CA82953C" box="[1352,1428,1543,1570]" name="Russia" pageId="18" pageNumber="285">Russia</collectingCountry>
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) (
|
||
<bibRefCitation id="EFE869FDFFD8FFB99C9FAE6ACE629558" author="Motschulsky, V. de" box="[159,372,1579,1606]" pageId="18" pageNumber="285" pagination="44 - 93" refId="ref14537" refString="Motschulsky, V. de (1839) Insectes du Caucase et des Provinces Transcaucausiennes. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 12, 44 - 93, pls. I, II, V, VI." type="journal article" year="1839">Motschulsky 1839</bibRefCitation>
|
||
).
|
||
<typeStatus id="54C2AAAEFFD8FFB99D88AE6DCEDC9558" box="[392,458,1580,1606]" pageId="18" pageNumber="285">Types</typeStatus>
|
||
of all species are lost, but the first author examined specimens from Baku, Kurush and
|
||
<collectingCountry id="F36E549CFFD8FFB99C97AE0ECFE79577" box="[151,241,1615,1641]" name="Georgia" pageId="18" pageNumber="285">Georgia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFE869FDFFD8FFB99D00AE0ECDA69574" author="Abdurakhmanov, G. M. & Nabozhenko, M. V." box="[256,688,1615,1642]" pageId="18" pageNumber="285" refId="ref12648" refString="Abdurakhmanov, G. M. & Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov & Nabozhenko 2011</bibRefCitation>
|
||
) and confirmed that all three taxa are morphologically conspecific and differ only in the level of elytral rugosity: from weak, poorly visible wrinkles in some specimens from Kurush and Baku to strongly and coarsely rugose specimens from mountains of Inner Dagestan, Chechnya and southern slopes of the Greater Caucasus in
|
||
<collectingCountry id="F36E549CFFD8FFB99E16AEFACD6595CB" box="[534,627,1723,1749]" name="Georgia" pageId="18" pageNumber="285">Georgia</collectingCountry>
|
||
. Adams mentioned that this very rare species has rugose flattened elytra with mucro, subquadrate pronotum, and long legs. Only one species with such characters occurs in
|
||
<collectingCountry id="F36E549CFFD8FFB9990AAE9ECA7195E7" box="[1290,1383,1759,1785]" name="Georgia" pageId="18" pageNumber="285">Georgia</collectingCountry>
|
||
, and it was listed for the eastern part of the country (Nakalakebi, Kazbegi) by
|
||
<bibRefCitation id="EFE869FDFFD8FFB99FD4AF42CBFE9400" author="Dzhambazishvili, M. Ya." box="[980,1256,1795,1822]" pageId="18" pageNumber="285" pagination="185 - 193" refId="ref13445" refString="Dzhambazishvili, M. Ya. (2000) Catalogue to darkling beetles (Coleoptera, Tenebrionidae) of Georgia. Trudy instituta zoologii Akademii nauk Gruzii, 20, 185 - 193. [in Russian]" type="journal article" year="2000">Dzhambazishvili (2000)</bibRefCitation>
|
||
. As a result, the following synonymy is proposed:
|
||
<taxonomicName id="4C796F8FFFD8FFB99E1AAF66CC67945C" authority="Adams, 1817" authorityName="Adams" authorityYear="1817" box="[538,881,1831,1858]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="285" phylum="Arthropoda" rank="species" species="verrucosa">
|
||
<emphasis id="B90DC81EFFD8FFB99E1AAF66CDC4945F" box="[538,722,1831,1857]" italics="true" pageId="18" pageNumber="285">Blaps verrucosa</emphasis>
|
||
Adams, 1817
|
||
</taxonomicName>
|
||
<emphasis id="B90DC81EFFD8FFB99F79AF69CC9F945C" bold="true" box="[889,905,1832,1858]" pageId="18" pageNumber="285">=</emphasis>
|
||
<taxonomicName id="4C796F8FFFD8FFB99F91AF66CA31945C" authority="Menetries, 1832" authorityName="Menetries" authorityYear="1832" box="[913,1319,1831,1858]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="285" phylum="Arthropoda" rank="species" species="scabriuscula">
|
||
<emphasis id="B90DC81EFFD8FFB99F91AF66CB7E945F" box="[913,1128,1831,1857]" italics="true" pageId="18" pageNumber="285">Blaps scabriuscula</emphasis>
|
||
<bibRefCitation id="EFE869FDFFD8FFB99870AF66CA31945C" author="Menetries, E." box="[1136,1319,1831,1858]" pageId="18" pageNumber="285" refId="ref14478" refString="Menetries, E. (1832) Catalogue raissonne des objets de Zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse. St. Petersbourg, L'Imprimerie de l'Academie Imperiale des Sciences, 272 pp., 5 pls. https: // doi. org / 10.5962 / bhl. title. 51784" type="book" year="1832">Ménétriés, 1832</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B90DC81EFFD8FFB9992FAF69CA8A945C" bold="true" box="[1327,1436,1832,1858]" pageId="18" pageNumber="285">
|
||
<taxonomicNameLabel id="A23E7565FFD8FFB9992FAF69CA68945C" box="[1327,1406,1832,1858]" pageId="18" pageNumber="285">syn. n.</taxonomicNameLabel>
|
||
, =
|
||
</emphasis>
|
||
<taxonomicName id="4C796F8FFFD8FFB99C97AF0ACD0A9478" authority="Motschulsky, 1839" authorityName="Motschulsky" authorityYear="1839" box="[151,540,1867,1894]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="285" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="B90DC81EFFD8FFB99C97AF0ACE56947B" box="[151,320,1867,1893]" italics="true" pageId="18" pageNumber="285">Blaps montana</emphasis>
|
||
<bibRefCitation id="EFE869FDFFD8FFB99D47AF0ACD0A9478" author="Motschulsky, V. de" box="[327,540,1867,1894]" pageId="18" pageNumber="285" pagination="44 - 93" refId="ref14537" refString="Motschulsky, V. de (1839) Insectes du Caucase et des Provinces Transcaucausiennes. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 12, 44 - 93, pls. I, II, V, VI." type="journal article" year="1839">Motschulsky, 1839</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B90DC81EFFD8FFB99E23AF0DCD669478" bold="true" box="[547,624,1868,1894]" pageId="18" pageNumber="285">
|
||
<taxonomicNameLabel id="A23E7565FFD8FFB99E23AF0DCD669478" box="[547,624,1868,1894]" pageId="18" pageNumber="285">syn. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC6140CFFD8FFB99CC7AF31CC2F94B3" blockId="18.[151,1437,1471,1966]" pageId="18" pageNumber="285">
|
||
<emphasis id="B90DC81EFFD8FFB99CC7AF31CE079494" bold="true" box="[199,273,1904,1930]" pageId="18" pageNumber="285">Notes.</emphasis>
|
||
A.V. Bogatshev confused the status and subspecies of
|
||
<taxonomicName id="4C796F8FFFD8FFB99F78AF31CB119497" authorityName="Adams" authorityYear="1817" box="[888,1031,1904,1929]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="285" phylum="Arthropoda" rank="species" species="verrucosa">
|
||
<emphasis id="B90DC81EFFD8FFB99F78AF31CB119497" box="[888,1031,1904,1929]" italics="true" pageId="18" pageNumber="285">B. verrucosa</emphasis>
|
||
</taxonomicName>
|
||
and mixed them with other species. The following interpretations were published by Bogatshev:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3634787FFD9FFB89C97A8D6CE9192B3" pageId="19" pageNumber="286" type="description">
|
||
<paragraph id="8BC6140CFFD9FFB89C97A8D6CDC793E7" blockId="19.[151,1437,151,429]" pageId="19" pageNumber="286">
|
||
1.
|
||
<bibRefCitation id="EFE869FDFFD9FFB89CC6A8D6CE8393AF" author="Bogatshev, A. V." box="[198,405,151,177]" pageId="19" pageNumber="286" pagination="14 - 71" refId="ref12704" refString="Bogatshev, A. V. (1934) Materials to the knowledge of the fauna of beetles of Apsheron Penisula. Trudy Azerbaijanskogo otdelenia Zakavkazskogo filiala of the AN SSR. Sektor zoologii, 7, 14 - 71." type="journal article" year="1934">Bogatshev (1934)</bibRefCitation>
|
||
:
|
||
<taxonomicName id="4C796F8FFFD9FFB89DA4A8D6CD4693AE" authorityName="Motschulsky" authorityYear="1839" box="[420,592,151,177]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="B90DC81EFFD9FFB89DA4A8D6CD4693AE" box="[420,592,151,177]" italics="true" pageId="19" pageNumber="286">Blaps montana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B90DC81EFFD9FFB89E64A8D6CD6293AF" bold="true" box="[612,628,151,177]" pageId="19" pageNumber="286">=</emphasis>
|
||
<taxonomicName id="4C796F8FFFD9FFB89E7FA8D6CC4E93AF" authorityName="Menetries" authorityYear="1832" box="[639,856,151,177]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="scabriuscula">
|
||
<emphasis id="B90DC81EFFD9FFB89E7FA8D6CC4E93AF" box="[639,856,151,177]" italics="true" pageId="19" pageNumber="286">Blaps scabriuscula</emphasis>
|
||
</taxonomicName>
|
||
, as a junior synonym). In addition, he mentioned two of its subspecies
|
||
<emphasis id="B90DC81EFFD9FFB89DB8A8FACD9993CA" box="[440,655,187,212]" italics="true" pageId="19" pageNumber="286">
|
||
<taxonomicName id="4C796F8FFFD9FFB89DB8A8FACD2F93CA" authorityName="Motschulsky" authorityYear="1839" box="[440,569,187,212]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="montana">B. montana</taxonomicName>
|
||
<taxonomicName id="4C796F8FFFD9FFB89E41A8FACD9993CA" authorityName="Gebler" authorityYear="1825" box="[577,655,187,212]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="rugosa">rugosa</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B90DC81EFFD9FFB89EC6A8FACCCA93CB" box="[710,988,187,213]" italics="true" pageId="19" pageNumber="286">
|
||
<taxonomicName id="4C796F8FFFD9FFB89EC6A8FACC5E93CA" authorityName="Motschulsky" authorityYear="1839" box="[710,840,187,212]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="montana">B. montana</taxonomicName>
|
||
chevsuretica
|
||
</emphasis>
|
||
. Both names are
|
||
<emphasis id="B90DC81EFFD9FFB898A1A8FACA2393CB" box="[1185,1333,187,213]" italics="true" pageId="19" pageNumber="286">nomina nuda</emphasis>
|
||
, because descriptions are absent. The Greater Caucasus.
|
||
</paragraph>
|
||
<paragraph id="8BC6140CFFD9FFB89C97A942CD4F925F" blockId="19.[151,1437,151,429]" pageId="19" pageNumber="286">
|
||
2.
|
||
<bibRefCitation id="EFE869FDFFD9FFB89CC6A942CE859203" author="Bogatshev, A. V." box="[198,403,259,285]" pageId="19" pageNumber="286" pagination="118 - 127" refId="ref12745" refString="Bogatshev, A. V. (1938) Review of species of the fam. Tenebrionidae of the Caucasus and neighboring countries. Bulleten' Gosudarstvennogo Muzeya Gruzii, 9, 118 - 127 [in Russian]." type="journal article" year="1938">Bogatshev (1938)</bibRefCitation>
|
||
:
|
||
<taxonomicName id="4C796F8FFFD9FFB89D9FA942CD5E9202" authorityName="Motschulsky" authorityYear="1839" box="[415,584,259,285]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="B90DC81EFFD9FFB89D9FA942CD5E9202" box="[415,584,259,285]" italics="true" pageId="19" pageNumber="286">Blaps montana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<collectingCountry id="F36E549CFFD9FFB89E57A942CDB09203" box="[599,678,259,285]" name="Russia" pageId="19" pageNumber="286">Russia</collectingCountry>
|
||
:
|
||
<collectingRegion id="49BDDAEEFFD9FFB89EB2A942CC0C9203" box="[690,794,259,285]" country="Russia" name="Dagestan" pageId="19" pageNumber="286">Dagestan</collectingRegion>
|
||
(Kurush, Chadakolob);
|
||
<collectingCountry id="F36E549CFFD9FFB8982CA942CBBD9203" box="[1068,1195,259,285]" name="Azerbaijan" pageId="19" pageNumber="286">Azerbaijan</collectingCountry>
|
||
: Sary-Bash;
|
||
<collectingCountry id="F36E549CFFD9FFB8993BA942CA8F9203" box="[1339,1433,259,285]" name="Georgia" pageId="19" pageNumber="286">Georgia</collectingCountry>
|
||
: Kheledula, Latal, Lailchala, Mestia.
|
||
</paragraph>
|
||
<paragraph id="8BC6140CFFD9FFB89C97A90ACE9192B3" blockId="19.[151,1437,151,429]" pageId="19" pageNumber="286">
|
||
3.
|
||
<bibRefCitation id="EFE869FDFFD9FFB89CC6A90ACE8E927B" author="Bogatshev, A. V." box="[198,408,330,357]" pageId="19" pageNumber="286" refId="ref12784" refString="Bogatshev, A. V. (1958) Darkling beetles. In: Kryzhanovsky, O. L. & Ter-Minasyan, M. E. Beetles - Coleoptera. In: Animal world of the USSR. Vol. 5. Mountain regions of the European part of the USSR. Academy of Sciences of the USSR, Moscow - Leningrad, pp. 384 - 421." type="book" year="1958">Bogatshev (1958)</bibRefCitation>
|
||
:
|
||
<taxonomicName id="4C796F8FFFD9FFB89DA8A90ACC09927B" authority="scabriuscula" box="[424,799,331,357]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="subSpecies" species="scabriuscula" subSpecies="scabriuscula">
|
||
<emphasis id="B90DC81EFFD9FFB89DA8A90ACC09927B" box="[424,799,331,357]" italics="true" pageId="19" pageNumber="286">Blaps scabriuscula scabriuscula</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="F36E549CFFD9FFB89F32A90ACCA7927B" box="[818,945,331,357]" name="Azerbaijan" pageId="19" pageNumber="286">Azerbaijan</collectingCountry>
|
||
:
|
||
<collectingRegion id="49BDDAEEFFD9FFB89FC1A90ACCEA927B" box="[961,1020,331,357]" country="Azerbaijan" name="Baku" pageId="19" pageNumber="286">Baku</collectingRegion>
|
||
and surrounds;
|
||
<taxonomicName id="4C796F8FFFD9FFB898C1A90ACE3E9296" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="subSpecies" species="scabriuscula" subSpecies="montana">
|
||
<emphasis id="B90DC81EFFD9FFB898C1A90ACE3E9296" italics="true" pageId="19" pageNumber="286">Blaps scabriuscula montana</emphasis>
|
||
</taxonomicName>
|
||
: Greater Caucasus;
|
||
<taxonomicName id="4C796F8FFFD9FFB89E04A92ECC999297" authority="Allard, 1881" authorityName="Menetries" authorityYear="1832" box="[516,911,367,393]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="scabriuscula">
|
||
<emphasis id="B90DC81EFFD9FFB89E04A92ECDED9297" box="[516,763,367,393]" italics="true" pageId="19" pageNumber="286">B. scabriuscula puella</emphasis>
|
||
Allard, 1881
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="F36E549CFFD9FFB89F96A92ECCED9297" box="[918,1019,367,393]" name="Armenia" pageId="19" pageNumber="286">Armenia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36E549CFFD9FFB89803A92ECB689297" box="[1027,1150,367,393]" name="Azerbaijan" pageId="19" pageNumber="286">Azerbaijan</collectingCountry>
|
||
(Karabakh,
|
||
<collectingRegion id="49BDDAEEFFD9FFB89905A92ECA879294" box="[1285,1425,367,394]" country="Azerbaijan" name="Naxcivan" pageId="19" pageNumber="286">Nakhchivan</collectingRegion>
|
||
), Eastern
|
||
<collectingCountry id="F36E549CFFD9FFB89D1FA9D2CE9592B3" box="[287,387,403,429]" name="Turkey" pageId="19" pageNumber="286">Anatolia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3634787FFD9FFB89C97A99ACDAA9123" pageId="19" pageNumber="286" type="reference_group">
|
||
<paragraph id="8BC6140CFFD9FFB89C97A99ACDAA9123" blockId="19.[151,1437,474,573]" pageId="19" pageNumber="286">
|
||
<taxonomicName id="4C796F8FFFD9FFB89C97A99ACE3292EB" authorityName="Allard" authorityYear="1881" box="[151,292,475,501]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="B90DC81EFFD9FFB89C97A99ACE3292EB" box="[151,292,475,501]" italics="true" pageId="19" pageNumber="286">Blaps puella</emphasis>
|
||
</taxonomicName>
|
||
, which was interpreted as a subspecies of
|
||
<taxonomicName id="4C796F8FFFD9FFB89EFDA99ACB6A92EB" authority="(Bogatshev 1958)" baseAuthorityName="Bogatshev" baseAuthorityYear="1958" box="[765,1148,474,501]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="scabriuscula">
|
||
<emphasis id="B90DC81EFFD9FFB89EFDA99ACCBD92EB" box="[765,939,475,501]" italics="true" pageId="19" pageNumber="286">B. scabriuscula</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE869FDFFD9FFB89FB9A99ACB6592EB" author="Bogatshev, A. V." box="[953,1139,474,501]" pageId="19" pageNumber="286" refId="ref12784" refString="Bogatshev, A. V. (1958) Darkling beetles. In: Kryzhanovsky, O. L. & Ter-Minasyan, M. E. Beetles - Coleoptera. In: Animal world of the USSR. Vol. 5. Mountain regions of the European part of the USSR. Academy of Sciences of the USSR, Moscow - Leningrad, pp. 384 - 421." type="book" year="1958">Bogatshev 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, belongs to the
|
||
<emphasis id="B90DC81EFFD9FFB8992BA99ACA8B92EA" box="[1323,1437,475,500]" italics="true" pageId="19" pageNumber="286">mortisaga</emphasis>
|
||
species-group with acute, strongly elongate reservoirs of the spermatheca (
|
||
<bibRefCitation id="EFE869FDFFD9FFB89801A9BECA4B9107" author="Chigray, I. A. & Nabozhenko, M. V." box="[1025,1373,511,537]" pageId="19" pageNumber="286" pagination="267 - 275" refId="ref13055" refString="Chigray, I. A. & Nabozhenko, M. V. (2016) To the knowledge of the genus Blaps Fabricius, 1775 (Coleoptera: Tenebrionidae) from Iran and Transcaucasia. Annales Zoologici, 66 (2), 267 - 275. https: // doi. org / 10.3161 / 00034541 ANZ 2016.66.2.007" type="journal article" year="2016">Chigray & Nabozhenko 2016</bibRefCitation>
|
||
,
|
||
<figureCitation id="13420889FFD9FFB8996CA9BECFCA9123" captionStart="FIGURE 7" captionStartId="6.[151,250,1157,1181]" captionTargetBox="[159,1428,319,1127]" captionTargetId="figure-310@6.[151,1436,315,1133]" captionTargetPageId="6" captionText="FIGURE 7. B. rugosa, habitus. A, B = ♂; C = ♀; A, C = dorsal view; B = ventral view (on: Chigray (2020)." figureDoi="http://doi.org/10.5281/zenodo.6367341" httpUri="https://zenodo.org/record/6367341/files/figure.png" pageId="19" pageNumber="286">Figs 7A–C</figureCitation>
|
||
) only slightly differing from
|
||
<emphasis id="B90DC81EFFD9FFB89E24AA62CDA39122" box="[548,693,547,572]" italics="true" pageId="19" pageNumber="286">B. mortisaga</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |