treatments-xml/data/03/CB/D4/03CBD43FFFD7FFD0FF5B2C167C28FE08.xml
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<mods:title id="F31B18917D184FA173C405965C41764E">Two new species of Utivarachna Kishida, 1940 from southern China, with an updated key to the Chinese species (Araneae: Trachelidae)</mods:title>
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<mods:namePart id="D125B32AE67F82E0E8B961F1FFA3744A">Liu, Keke</mods:namePart>
<mods:affiliation id="A8D0EA5BFBA7A508789116949B0B9FA9">College of Life Science, Hunan Normal University, Changsha 410081, Hunan, China &amp; liukeke _ 1986 @ 126. com; https: // orcid. org / 0000 - 0001 - 7822 - 3667 &amp; yinhaiqiang @ hunnu. edu. cn; https: // orcid. org / 0000 - 0003 - 0276 - 2407 &amp; xux @ hunnu. edu. cn; https: // orcid. org / 0000 - 0001 - 9485 - 5373 &amp; 2521973053 @ qq. com; https: // orcid. org / 0000 - 0002 - 4477 - 1187 &amp; College of Life Science, Jinggangshan University, Jian 343009, Jiangxi, China &amp; liukeke _ 1986 @ 126. com; https: // orcid. org / 0000 - 0001 - 7822 - 3667</mods:affiliation>
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<mods:namePart id="3F99F16DD5E15CCF1BAD310BF8482BBF">Yin, Haiqiang</mods:namePart>
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<mods:namePart id="426929AB1C16B85D1CD3DFC4614A6303">Xu, Xiang</mods:namePart>
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<taxonomicName id="4C621EAAFFD7FFD4FF5B2C167B24FCF8" ID-CoL="9927T" authority="Liu, Xu &amp; Haddad" authorityName="Liu, Xu &amp; Haddad" authorityYear="2020" box="[151,616,837,864]" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="97" phylum="Arthropoda" rank="species" species="tangi" status="sp. nov.">
<emphasis id="B916B93BFFD7FFD4FF5B2C167838FCF8" bold="true" box="[151,372,837,864]" italics="true" pageId="10" pageNumber="97">Utivarachna tangi</emphasis>
Liu, Xu &amp; Haddad
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<figureCitation id="135979ACFFD7FFD4FF5B2C3A7997FC24" box="[151,219,873,899]" captionStart="FIGURE 7" captionStartId="11.[151,250,1945,1970]" captionTargetBox="[210,1378,181,1920]" captionTargetId="figure@11.[210,1378,181,1920]" captionTargetPageId="11" captionText="FIGURE 7AE. Utivarachna tangi sp. nov., female holotype. A. Habitus, dorsal view. B. Same, dorsal view. C. Eyes, ventral view. D. Chelicerae, ventral view. E. Maxillae, labium and sternum, ventral view. Scale bars: A, B, E = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908039" httpUri="https://zenodo.org/record/3908039/files/figure.png" pageId="10" pageNumber="97">Figs 7</figureCitation>
AE, 8AK, 9A, B
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<emphasis id="B916B93BFFD7FFD4FF5B2C86780AFC48" bold="true" box="[151,326,980,1007]" pageId="10" pageNumber="97">Type material.</emphasis>
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<emphasis id="B916B93BFFD7FFD4FEE42B4D782CFB90" box="[296,352,1053,1079]" italics="true" pageId="10" pageNumber="97">et al.</emphasis>
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<emphasis id="B916B93BFFD7FFD4FF0B2B127802FBFC" bold="true" box="[199,334,1089,1115]" pageId="10" pageNumber="97">Etymology.</emphasis>
The specific name is a patronym in honor of Dr Guo Tang, who made significant contributions to spider taxonomy in
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<emphasis id="B916B93BFFD7FFD4FF0B2BDB780EFB04" bold="true" box="[199,322,1160,1187]" pageId="10" pageNumber="97">Diagnosis.</emphasis>
This species is similar to those of
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<emphasis id="B916B93BFFD7FFD4FD0F2BD97A13FB04" box="[707,863,1161,1187]" italics="true" pageId="10" pageNumber="97">U. subfabaria</emphasis>
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</emphasis>
in having similar atria and bursae, but differs by the copulatory ducts being close to each other (
<figureCitation id="135979ACFFD7FFD4FD382BFE7A26FB60" box="[756,874,1197,1223]" captionStart="FIGURE 8" captionStartId="12.[151,250,1224,1249]" captionTargetBox="[339,1247,181,1199]" captionTargetId="figure@12.[339,1247,181,1199]" captionTargetPageId="12" captionText="FIGURE 8AK. Utivarachna tangi sp. nov., female holotype, legs and genitalia. A. Tarsus, metatarsus and tibia I, retrolateral view. B. Patella and femur I, retrolateral view. C. Tarsus, metatarsus and tibia II, retrolateral view. D. Patella and femur II, retrolateral view. E. Tarsus III, retrolateral view. F. Metatarsus, tibia, patella and femur III, retrolateral view. G. Tarsus IV, retrolateral view. H. Metatarsus and tibia IV, retrolateral view. I. Patella and femur IV, retrolateral view. J. Epigyne, ventral view. K. Vulva, dorsal view. Scale bars: AI = 0.5 mm; J, K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908041" httpUri="https://zenodo.org/record/3908041/files/figure.png" pageId="10" pageNumber="97">Figs 8J, K</figureCitation>
,
<figureCitation id="135979ACFFD7FFD4FCB92BFE7AF4FB60" box="[885,952,1197,1223]" captionStart="FIGURE 9" captionStartId="12.[151,250,1932,1957]" captionTargetBox="[334,1251,1421,1904]" captionTargetId="figure@12.[330,1257,1415,1907]" captionTargetPageId="12" captionText="FIGURE 9AB. Utivarachna tangi sp. nov. holotype, female genitalia.A. Epigyne, ventral view. B. Vulva, dorsal view.Abbreviations: A, atrium; BU, bursae; CD, copulatory duct; CO, copulatory openings; FD, fertilization duct; GP, glandular particles; SP, spermathecae. Scale bars: A, B = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908043" httpUri="https://zenodo.org/record/3908043/files/figure.png" pageId="10" pageNumber="97">9A, B</figureCitation>
)
<emphasis id="B916B93BFFD7FFD4FC0B2BFD7AAAFB60" box="[967,998,1198,1223]" italics="true" pageId="10" pageNumber="97">vs.</emphasis>
separated and parallel in
<taxonomicName id="4C621EAAFFD7FFD4FACD2BFD7CD0FB60" authority="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma, 2020" authorityName="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma" authorityYear="2020" box="[1281,1436,1197,1223]" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="97" phylum="Arthropoda" rank="species" species="subfabaria" status="sp. nov.">
<emphasis id="B916B93BFFD7FFD4FACD2BFD7CD0FB60" box="[1281,1436,1197,1223]" italics="true" pageId="10" pageNumber="97">U. subfabaria</emphasis>
</taxonomicName>
<emphasis id="B916B93BFFD7FFD4FF5B2B8279BDFB4C" bold="true" box="[151,241,1233,1259]" pageId="10" pageNumber="97">
<taxonomicNameLabel id="A2250440FFD7FFD4FF5B2B8279BDFB4C" box="[151,241,1233,1259]" pageId="10" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="135979ACFFD7FFD4FECC2B827827FB4C" box="[256,363,1232,1259]" captionStart="FIGURE 5" captionStartId="8.[151,250,1484,1509]" captionTargetBox="[225,1362,181,1459]" captionTargetId="figure@8.[225,1362,181,1459]" captionTargetPageId="8" captionText="FIGURE 5AJ. Utivarachna subfabaria sp. nov., female paratype, legs and genitalia.A. Tarsus, metatarsus and tibia I, retrolateral view. B. Patella and femur I, retrolateral view. C. Tarsus, metatarsus and tibia II, retrolateral view. D. Patella and femur II, retrolateral view. E. Tarsus and metatarsus III, retrolateral view. F. Tibia, patella and femur III, retrolateral view. G. Tarsus and metatarsus IV, retrolateral view. H. Tibia, patella and femur IV, retrolateral view. I. Epigyne, ventral view. J. Vulva, dorsal view. Scale bars: AH = 0.5 mm; I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908035" httpUri="https://zenodo.org/record/3908035/files/figure.png" pageId="10" pageNumber="97">Figs 5I, J</figureCitation>
,
<figureCitation id="135979ACFFD7FFD4FEBB2B8278F9FB4C" box="[375,437,1233,1259]" captionStart="FIGURE 6" captionStartId="9.[151,250,1852,1877]" captionTargetBox="[270,1318,597,1816]" captionTargetId="figure@9.[264,1322,597,1827]" captionTargetPageId="9" captionText="FIGURE 6AF. Utivarachna subfabaria sp. nov. male palp and female genitalia. A. Male palp, prolateral view. B. Same, ventral view. C. Same, retrolateral view. D. Same, dorsolateral view. E. Epigyne, ventral view. F. Vulva, dorsal view.Abbreviations: A, atrium; B, Bulb; BU, bursae; CD, copulatory duct; CO, copulatory openings; E, embolus; FD, fertilization duct; GP, glandular particles; RTA, retrolateral tibial apophysis; SD, sperm duct; SP, spermathecae; ST, subtegulum; T, tegulum. Scale bars: AF = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908037" httpUri="https://zenodo.org/record/3908037/files/figure.png" pageId="10" pageNumber="97">6E, F</figureCitation>
) and the blade-shaped bursae (
<figureCitation id="135979ACFFD7FFD4FCDD2B827AC4FB4C" box="[785,904,1233,1259]" captionStart="FIGURE 8" captionStartId="12.[151,250,1224,1249]" captionTargetBox="[339,1247,181,1199]" captionTargetId="figure@12.[339,1247,181,1199]" captionTargetPageId="12" captionText="FIGURE 8AK. Utivarachna tangi sp. nov., female holotype, legs and genitalia. A. Tarsus, metatarsus and tibia I, retrolateral view. B. Patella and femur I, retrolateral view. C. Tarsus, metatarsus and tibia II, retrolateral view. D. Patella and femur II, retrolateral view. E. Tarsus III, retrolateral view. F. Metatarsus, tibia, patella and femur III, retrolateral view. G. Tarsus IV, retrolateral view. H. Metatarsus and tibia IV, retrolateral view. I. Patella and femur IV, retrolateral view. J. Epigyne, ventral view. K. Vulva, dorsal view. Scale bars: AI = 0.5 mm; J, K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908041" httpUri="https://zenodo.org/record/3908041/files/figure.png" pageId="10" pageNumber="97">Figs 8J, K</figureCitation>
,
<figureCitation id="135979ACFFD7FFD4FC5F2B827A9AFB4C" box="[915,982,1233,1259]" captionStart="FIGURE 9" captionStartId="12.[151,250,1932,1957]" captionTargetBox="[334,1251,1421,1904]" captionTargetId="figure@12.[330,1257,1415,1907]" captionTargetPageId="12" captionText="FIGURE 9AB. Utivarachna tangi sp. nov. holotype, female genitalia.A. Epigyne, ventral view. B. Vulva, dorsal view.Abbreviations: A, atrium; BU, bursae; CD, copulatory duct; CO, copulatory openings; FD, fertilization duct; GP, glandular particles; SP, spermathecae. Scale bars: A, B = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908043" httpUri="https://zenodo.org/record/3908043/files/figure.png" pageId="10" pageNumber="97">9A, B</figureCitation>
)
<emphasis id="B916B93BFFD7FFD4FC2B2B817D49FB4C" box="[999,1029,1234,1259]" italics="true" pageId="10" pageNumber="97">vs.</emphasis>
ampullate in
<taxonomicName id="4C621EAAFFD7FFD4FB532B817C77FB4C" authority="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma, 2020" authorityName="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma" authorityYear="2020" box="[1183,1339,1233,1259]" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="97" phylum="Arthropoda" rank="species" species="subfabaria" status="sp. nov.">
<emphasis id="B916B93BFFD7FFD4FB532B817C77FB4C" box="[1183,1339,1233,1259]" italics="true" pageId="10" pageNumber="97">U. subfabaria</emphasis>
</taxonomicName>
<emphasis id="B916B93BFFD7FFD4FA8E2B827CD0FB4C" bold="true" box="[1346,1436,1233,1259]" pageId="10" pageNumber="97">
<taxonomicNameLabel id="A2250440FFD7FFD4FA8E2B827CD0FB4C" box="[1346,1436,1233,1259]" pageId="10" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="135979ACFFD7FFD4FF532BA67846FAA8" box="[159,266,1268,1295]" captionStart="FIGURE 5" captionStartId="8.[151,250,1484,1509]" captionTargetBox="[225,1362,181,1459]" captionTargetId="figure@8.[225,1362,181,1459]" captionTargetPageId="8" captionText="FIGURE 5AJ. Utivarachna subfabaria sp. nov., female paratype, legs and genitalia.A. Tarsus, metatarsus and tibia I, retrolateral view. B. Patella and femur I, retrolateral view. C. Tarsus, metatarsus and tibia II, retrolateral view. D. Patella and femur II, retrolateral view. E. Tarsus and metatarsus III, retrolateral view. F. Tibia, patella and femur III, retrolateral view. G. Tarsus and metatarsus IV, retrolateral view. H. Tibia, patella and femur IV, retrolateral view. I. Epigyne, ventral view. J. Vulva, dorsal view. Scale bars: AH = 0.5 mm; I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908035" httpUri="https://zenodo.org/record/3908035/files/figure.png" pageId="10" pageNumber="97">Figs 5I, J</figureCitation>
,
<figureCitation id="135979ACFFD7FFD4FEDA2BA6781AFAA8" box="[278,342,1269,1295]" captionStart="FIGURE 6" captionStartId="9.[151,250,1852,1877]" captionTargetBox="[270,1318,597,1816]" captionTargetId="figure@9.[264,1322,597,1827]" captionTargetPageId="9" captionText="FIGURE 6AF. Utivarachna subfabaria sp. nov. male palp and female genitalia. A. Male palp, prolateral view. B. Same, ventral view. C. Same, retrolateral view. D. Same, dorsolateral view. E. Epigyne, ventral view. F. Vulva, dorsal view.Abbreviations: A, atrium; B, Bulb; BU, bursae; CD, copulatory duct; CO, copulatory openings; E, embolus; FD, fertilization duct; GP, glandular particles; RTA, retrolateral tibial apophysis; SD, sperm duct; SP, spermathecae; ST, subtegulum; T, tegulum. Scale bars: AF = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908037" httpUri="https://zenodo.org/record/3908037/files/figure.png" pageId="10" pageNumber="97">6E, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37836A2FFD7FFD0FF0B2A4B7CD1FF70" lastPageId="14" lastPageNumber="101" pageId="10" pageNumber="97" type="description">
<paragraph id="8BDD6529FFD7FFD4FF0B2A4B7AAEF81C" blockId="10.[151,1437,980,1980]" pageId="10" pageNumber="97">
<emphasis id="B916B93BFFD7FFD4FF0B2A4B7815FA94" bold="true" box="[199,345,1304,1331]" pageId="10" pageNumber="97">Description.</emphasis>
Female. Habitus as in
<figureCitation id="135979ACFFD7FFD4FD9B2A4A7B9CFA94" box="[599,720,1305,1331]" captionStart="FIGURE 7" captionStartId="11.[151,250,1945,1970]" captionTargetBox="[210,1378,181,1920]" captionTargetId="figure@11.[210,1378,181,1920]" captionTargetPageId="11" captionText="FIGURE 7AE. Utivarachna tangi sp. nov., female holotype. A. Habitus, dorsal view. B. Same, dorsal view. C. Eyes, ventral view. D. Chelicerae, ventral view. E. Maxillae, labium and sternum, ventral view. Scale bars: A, B, E = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908039" httpUri="https://zenodo.org/record/3908039/files/figure.png" pageId="10" pageNumber="97">Fig. 7A, B</figureCitation>
. Total length 5.43. Carapace (
<figureCitation id="135979ACFFD7FFD4FBD02A4A7D38FA94" box="[1052,1140,1305,1331]" captionStart="FIGURE 7" captionStartId="11.[151,250,1945,1970]" captionTargetBox="[210,1378,181,1920]" captionTargetId="figure@11.[210,1378,181,1920]" captionTargetPageId="11" captionText="FIGURE 7AE. Utivarachna tangi sp. nov., female holotype. A. Habitus, dorsal view. B. Same, dorsal view. C. Eyes, ventral view. D. Chelicerae, ventral view. E. Maxillae, labium and sternum, ventral view. Scale bars: A, B, E = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908039" httpUri="https://zenodo.org/record/3908039/files/figure.png" pageId="10" pageNumber="97">Fig. 7A</figureCitation>
) dark brown, broadly oval in dorsal view (CL = 1.17 CW), pars cephalica strongly domed, anteriorly narrowed to 0.6 times its maximum width, with abundant black setae, posteriorly with indistinct cervical groove and distinct fovea, length 2.89, width 2.47. Eyes (
<figureCitation id="135979ACFFD7FFD4FF102AD6787DFA38" box="[220,305,1413,1439]" captionStart="FIGURE 7" captionStartId="11.[151,250,1945,1970]" captionTargetBox="[210,1378,181,1920]" captionTargetId="figure@11.[210,1378,181,1920]" captionTargetPageId="11" captionText="FIGURE 7AE. Utivarachna tangi sp. nov., female holotype. A. Habitus, dorsal view. B. Same, dorsal view. C. Eyes, ventral view. D. Chelicerae, ventral view. E. Maxillae, labium and sternum, ventral view. Scale bars: A, B, E = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908039" httpUri="https://zenodo.org/record/3908039/files/figure.png" pageId="10" pageNumber="97">Fig. 7C</figureCitation>
) diameters: AME 0.13, ALE 0.14, PME 0.14, PLE 0.14; interdistances:AMEAME 0.06, ALEAME 0.13, ALEALE 0.54, PMEPME 0.22, PLEPME 0.25, PLEPLE 0.95, AMEPME 0.10, ALEPLE 0.25. MOA 0.32 long, front width 0.34, back width 0.46, occupying most of anterior cephalic region. Mouthparts (
<figureCitation id="135979ACFFD7FFD4FAD92A9E7CC3FA40" box="[1301,1423,1485,1511]" captionStart="FIGURE 7" captionStartId="11.[151,250,1945,1970]" captionTargetBox="[210,1378,181,1920]" captionTargetId="figure@11.[210,1378,181,1920]" captionTargetPageId="11" captionText="FIGURE 7AE. Utivarachna tangi sp. nov., female holotype. A. Habitus, dorsal view. B. Same, dorsal view. C. Eyes, ventral view. D. Chelicerae, ventral view. E. Maxillae, labium and sternum, ventral view. Scale bars: A, B, E = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908039" httpUri="https://zenodo.org/record/3908039/files/figure.png" pageId="10" pageNumber="97">Fig. 7D, E</figureCitation>
): chelicerae yellow-brown (red-brown in furrow), straight, robust, with abundant setae along retromarginal teeth; 5 promarginal (4th largest, 5th smallest) and 3 retromarginal teeth (second largest); maxillae yellowish-pink, slightly convergent anteriorly, twice longer than wide, with numerous setae on anterior inner margin and dense maxillary setae tuft on anterior margin; labium pentagonal, with row of setae on anterior margin, slightly longer than wide, with trapezoidal base located at notch in anterior sternum, marginal grooves distinct in trapezoidal base. Sternum (
<figureCitation id="135979ACFFD7FFD4FAA329D279F4F918" captionStart="FIGURE 7" captionStartId="11.[151,250,1945,1970]" captionTargetBox="[210,1378,181,1920]" captionTargetId="figure@11.[210,1378,181,1920]" captionTargetPageId="11" captionText="FIGURE 7AE. Utivarachna tangi sp. nov., female holotype. A. Habitus, dorsal view. B. Same, dorsal view. C. Eyes, ventral view. D. Chelicerae, ventral view. E. Maxillae, labium and sternum, ventral view. Scale bars: A, B, E = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908039" httpUri="https://zenodo.org/record/3908039/files/figure.png" pageId="10" pageNumber="97">Fig. 7E</figureCitation>
) yellowish-white, as long as wide, with finely chain-shaped, raised, hollowed lines around edge. Abdomen (
<figureCitation id="135979ACFFD7FFD4FAA329F67997F944" captionStart="FIGURE 7" captionStartId="11.[151,250,1945,1970]" captionTargetBox="[210,1378,181,1920]" captionTargetId="figure@11.[210,1378,181,1920]" captionTargetPageId="11" captionText="FIGURE 7AE. Utivarachna tangi sp. nov., female holotype. A. Habitus, dorsal view. B. Same, dorsal view. C. Eyes, ventral view. D. Chelicerae, ventral view. E. Maxillae, labium and sternum, ventral view. Scale bars: A, B, E = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908039" httpUri="https://zenodo.org/record/3908039/files/figure.png" pageId="10" pageNumber="97">Fig. 7A, B</figureCitation>
) dark brown, 2.56 long, 2.45 wide, ovoid, with abundant short setae and three pairs of oval marks dorsally, and abundant pallid dots ventrally. Legs yellowish pink (
<figureCitation id="135979ACFFD7FFD4FCD229BE7A2CF8A0" box="[798,864,1773,1799]" captionStart="FIGURE 8" captionStartId="12.[151,250,1224,1249]" captionTargetBox="[339,1247,181,1199]" captionTargetId="figure@12.[339,1247,181,1199]" captionTargetPageId="12" captionText="FIGURE 8AK. Utivarachna tangi sp. nov., female holotype, legs and genitalia. A. Tarsus, metatarsus and tibia I, retrolateral view. B. Patella and femur I, retrolateral view. C. Tarsus, metatarsus and tibia II, retrolateral view. D. Patella and femur II, retrolateral view. E. Tarsus III, retrolateral view. F. Metatarsus, tibia, patella and femur III, retrolateral view. G. Tarsus IV, retrolateral view. H. Metatarsus and tibia IV, retrolateral view. I. Patella and femur IV, retrolateral view. J. Epigyne, ventral view. K. Vulva, dorsal view. Scale bars: AI = 0.5 mm; J, K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908041" httpUri="https://zenodo.org/record/3908041/files/figure.png" pageId="10" pageNumber="97">Fig. 8</figureCitation>
AH), with abundant setae on each segment; base of each tarsus notched dorsally, sub-base of femur enlarged, without conical ventral cusps and spines; legs III and IV distinctly thinner; femur IV longer than femora IIII, tibia IV longer than tibiae IIII, metatarsus IV longer than metatarsi IIII, tarsus I longer than tarsi IIIV; femora IIV wider basally; two claws with about 5 small teeth on each; measurements: I 6.44 (1.74, 0.93, 1.62, 1.24, 0.91); II 6.43 (1.79, 0.89, 1.63, 1.27, 0.86); III 5.49 (1.53, 0.80, 1.24, 1.32, 0.64); IV 7.26 (2.00, 0.81, 1.69, 1.95, 0.81). Leg formula: 4123.
</paragraph>
<caption id="DF1D35A1FFD6FFD5FF5B28CA798AF85E" ID-DOI="http://doi.org/10.5281/zenodo.3908039" ID-Zenodo-Dep="3908039" httpUri="https://zenodo.org/record/3908039/files/figure.png" pageId="11" pageNumber="98" startId="11.[151,250,1945,1970]" targetBox="[210,1378,181,1920]" targetPageId="11">
<paragraph id="8BDD6529FFD6FFD5FF5B28CA798AF85E" blockId="11.[151,1436,1945,2041]" pageId="11" pageNumber="98">
<emphasis id="B916B93BFFD6FFD5FF5B28CA7B13F815" bold="true" box="[151,607,1945,1970]" pageId="11" pageNumber="98">
FIGURE 7AE.
<taxonomicName id="4C621EAAFFD6FFD5FE8528CA7B45F815" authority="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma, 2020" authorityName="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma" authorityYear="2020" box="[329,521,1945,1970]" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="98" phylum="Arthropoda" rank="species" species="tangi" status="sp. nov.">
<emphasis id="B916B93BFFD6FFD5FE8528CA7B45F815" bold="true" box="[329,521,1945,1970]" italics="true" pageId="11" pageNumber="98">Utivarachna tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2250440FFD6FFD5FDC328C97B13F815" box="[527,607,1946,1970]" pageId="11" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female holotype. A. Habitus, dorsal view. B. Same, dorsal view. C. Eyes, ventral view. D. Chelicerae, ventral view. E. Maxillae, labium and sternum, ventral view. Scale bars: A, B, E = 0.5 mm; C, D = 0.1 mm.
</paragraph>
</caption>
<caption id="DF1D35A1FFD1FFD2FF5B2B9B7B8DFAD7" ID-DOI="http://doi.org/10.5281/zenodo.3908041" ID-Zenodo-Dep="3908041" httpUri="https://zenodo.org/record/3908041/files/figure.png" pageId="12" pageNumber="99" startId="12.[151,250,1224,1249]" targetBox="[339,1247,181,1199]" targetPageId="12">
<paragraph id="8BDD6529FFD1FFD2FF5B2B9B7B8DFAD7" blockId="12.[151,1436,1224,1393]" pageId="12" pageNumber="99">
<emphasis id="B916B93BFFD1FFD2FF5B2B9B7B2DFB46" bold="true" box="[151,609,1224,1249]" pageId="12" pageNumber="99">
FIGURE 8AK.
<taxonomicName id="4C621EAAFFD1FFD2FE802B9B7B47FB46" authority="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma, 2020" authorityName="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma" authorityYear="2020" box="[332,523,1224,1249]" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="99" phylum="Arthropoda" rank="species" species="tangi" status="sp. nov.">
<emphasis id="B916B93BFFD1FFD2FE802B9B7B47FB46" bold="true" box="[332,523,1224,1249]" italics="true" pageId="12" pageNumber="99">Utivarachna tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2250440FFD1FFD2FDDD2B9A7B2DFB46" box="[529,609,1225,1249]" pageId="12" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female holotype, legs and genitalia. A. Tarsus, metatarsus and tibia I, retrolateral view. B. Patella and femur I, retrolateral view. C. Tarsus, metatarsus and tibia II, retrolateral view. D. Patella and femur II, retrolateral view. E. Tarsus III, retrolateral view. F. Metatarsus, tibia, patella and femur III, retrolateral view. G. Tarsus IV, retrolateral view. H. Metatarsus and tibia IV, retrolateral view. I. Patella and femur IV, retrolateral view. J. Epigyne, ventral view. K. Vulva, dorsal view. Scale bars: AI = 0.5 mm; J, K = 0.1 mm.
</paragraph>
</caption>
<caption id="DF1D35A1FFD1FFD2FF5B28DF7B2BF84B" ID-DOI="http://doi.org/10.5281/zenodo.3908043" ID-Zenodo-Dep="3908043" httpUri="https://zenodo.org/record/3908043/files/figure.png" pageId="12" pageNumber="99" startId="12.[151,250,1932,1957]" targetBox="[334,1251,1421,1904]" targetPageId="12">
<paragraph id="8BDD6529FFD1FFD2FF5B28DF7B2BF84B" blockId="12.[151,1436,1932,2029]" pageId="12" pageNumber="99">
<emphasis id="B916B93BFFD1FFD2FF5B28DF7B15F802" bold="true" box="[151,601,1932,1957]" pageId="12" pageNumber="99">
FIGURE 9AB.
<taxonomicName id="4C621EAAFFD1FFD2FE8B28DF7B49F803" authority="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma, 2020" authorityName="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma" authorityYear="2020" box="[327,517,1932,1957]" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="99" phylum="Arthropoda" rank="species" species="tangi" status="sp. nov.">
<emphasis id="B916B93BFFD1FFD2FE8B28DF7B49F803" bold="true" box="[327,517,1932,1957]" italics="true" pageId="12" pageNumber="99">Utivarachna tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2250440FFD1FFD2FDC628DE7B15F802" box="[522,601,1933,1957]" pageId="12" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype, female genitalia. A. Epigyne, ventral view. B. Vulva, dorsal view.Abbreviations: A, atrium; BU, bursae; CD, copulatory duct; CO, copulatory openings; FD, fertilization duct; GP, glandular particles; SP, spermathecae. Scale bars: A, B = 0.1 mm.
</paragraph>
</caption>
<caption id="DF1D35A1FFD0FFD3FF5B284D7DEBF890" ID-DOI="http://doi.org/10.5281/zenodo.3908045" ID-Zenodo-Dep="3908045" box="[151,1191,1822,1847]" httpUri="https://zenodo.org/record/3908045/files/figure.png" pageId="13" pageNumber="100" startId="13.[151,250,1822,1847]" targetBox="[185,1397,218,1766]" targetPageId="13">
<paragraph id="8BDD6529FFD0FFD3FF5B284D7DEBF890" blockId="13.[151,1191,1822,1847]" box="[151,1191,1822,1847]" pageId="13" pageNumber="100">
<emphasis id="B916B93BFFD0FFD3FF5B284D786CF890" bold="true" box="[151,288,1822,1847]" pageId="13" pageNumber="100">FIGURE 10.</emphasis>
Collection localities for
<taxonomicName id="4C621EAAFFD0FFD3FDD0284C7A5CF890" authority="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma, 2020" authorityName="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma" authorityYear="2020" box="[540,784,1823,1847]" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="100" phylum="Arthropoda" rank="species" species="subfabaria" status="sp. nov.">
<emphasis id="B916B93BFFD0FFD3FDD0284C7A5CF890" box="[540,784,1823,1847]" italics="true" pageId="13" pageNumber="100">Utivarachna subfabaria</emphasis>
</taxonomicName>
<emphasis id="B916B93BFFD0FFD3FCDA284C7A2BF890" bold="true" box="[790,871,1823,1847]" pageId="13" pageNumber="100">
<taxonomicNameLabel id="A2250440FFD0FFD3FCDA284C7A2BF890" box="[790,871,1823,1847]" pageId="13" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C621EAAFFD0FFD3FC5428737AA7F890" authority="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma, 2020" authorityName="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma" authorityYear="2020" box="[920,1003,1824,1847]" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="100" phylum="Arthropoda" rank="species" species="tangi" status="sp. nov.">
<emphasis id="B916B93BFFD0FFD3FC5428737AA7F890" box="[920,1003,1824,1847]" italics="true" pageId="13" pageNumber="100">U. tangi</emphasis>
</taxonomicName>
<emphasis id="B916B93BFFD0FFD3FC3E284C7D0EF890" bold="true" box="[1010,1090,1823,1847]" pageId="13" pageNumber="100">
<taxonomicNameLabel id="A2250440FFD0FFD3FC3E284C7D0EF890" box="[1010,1090,1823,1847]" pageId="13" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in China.
</paragraph>
</caption>
<paragraph id="8BDD6529FFD0FFD0FF0B28367CD1FF70" blockId="13.[151,1436,1893,2028]" lastBlockId="14.[151,1437,153,431]" lastPageId="14" lastPageNumber="101" pageId="13" pageNumber="100">
Epigyne (
<figureCitation id="135979ACFFD0FFD3FEFD283678E8F827" box="[305,420,1893,1920]" captionStart="FIGURE 8" captionStartId="12.[151,250,1224,1249]" captionTargetBox="[339,1247,181,1199]" captionTargetId="figure@12.[339,1247,181,1199]" captionTargetPageId="12" captionText="FIGURE 8AK. Utivarachna tangi sp. nov., female holotype, legs and genitalia. A. Tarsus, metatarsus and tibia I, retrolateral view. B. Patella and femur I, retrolateral view. C. Tarsus, metatarsus and tibia II, retrolateral view. D. Patella and femur II, retrolateral view. E. Tarsus III, retrolateral view. F. Metatarsus, tibia, patella and femur III, retrolateral view. G. Tarsus IV, retrolateral view. H. Metatarsus and tibia IV, retrolateral view. I. Patella and femur IV, retrolateral view. J. Epigyne, ventral view. K. Vulva, dorsal view. Scale bars: AI = 0.5 mm; J, K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908041" httpUri="https://zenodo.org/record/3908041/files/figure.png" pageId="13" pageNumber="100">Figs 8J, K</figureCitation>
,
<figureCitation id="135979ACFFD0FFD3FE61283578F1F827" box="[429,445,1894,1920]" captionStart="FIGURE 9" captionStartId="12.[151,250,1932,1957]" captionTargetBox="[334,1251,1421,1904]" captionTargetId="figure@12.[330,1257,1415,1907]" captionTargetPageId="12" captionText="FIGURE 9AB. Utivarachna tangi sp. nov. holotype, female genitalia.A. Epigyne, ventral view. B. Vulva, dorsal view.Abbreviations: A, atrium; BU, bursae; CD, copulatory duct; CO, copulatory openings; FD, fertilization duct; GP, glandular particles; SP, spermathecae. Scale bars: A, B = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3908043" httpUri="https://zenodo.org/record/3908043/files/figure.png" pageId="13" pageNumber="100">9</figureCitation>
). Atrium large, as long as wide, occupying almost whole posterior half of plate; copulatory openings semilunar-shaped, closely spaced, located in middle of epigyne, connecting bursae by long thin copulatory ducts; copulatory ducts long, with median part between bursae straight and parallel, posteriorly connected to anterodorsal spermathecae; bursae large, 2 times longer than wide, with posterior margins touching anterior margins of comma- shaped spermathecae;bursae with several small clusters of glandular particles on posterior surface, occupying about2/3 of epigyne length; fertilization ducts broad, widely separated from each other, distance almost equal to width of atrium.
</paragraph>
</subSubSection>
<subSubSection id="C37836A2FFD3FFD0FF0B2FB2787AFE2C" pageId="14" pageNumber="101" type="discussion">
<paragraph id="8BDD6529FFD3FFD0FF0B2FB27AA9FEE4" blockId="14.[151,1437,153,431]" pageId="14" pageNumber="101">
<emphasis id="B916B93BFFD3FFD0FF0B2FB27877FF5C" bold="true" box="[199,315,225,251]" pageId="14" pageNumber="101">Remarks.</emphasis>
This new species occurs exactly in the same locality as the
<typeStatus id="54D9DB8BFFD3FFD0FC362FB27D29FF5C" box="[1018,1125,225,251]" pageId="14" pageNumber="101" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C621EAAFFD3FFD0FB5D2FB17C63FF5C" authority="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma, 2020" authorityName="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma" authorityYear="2020" box="[1169,1327,225,251]" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="101" phylum="Arthropoda" rank="species" species="subfabaria" status="sp. nov.">
<emphasis id="B916B93BFFD3FFD0FB5D2FB17C63FF5C" box="[1169,1327,225,251]" italics="true" pageId="14" pageNumber="101">U. subfabaria</emphasis>
</taxonomicName>
<emphasis id="B916B93BFFD3FFD0FAF52FB27CD9FF5C" bold="true" box="[1337,1429,225,251]" pageId="14" pageNumber="101">
<taxonomicNameLabel id="A2250440FFD3FFD0FAF52FB27CD9FF5C" box="[1337,1429,225,251]" pageId="14" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Nonggang National Nature Reserve. These two species are sympatric in at least some sites explored by the authors. Differences in color and epigyne morphology can be used to separate them.
</paragraph>
<paragraph id="8BDD6529FFD3FFD0FF0B2E1F787AFE2C" blockId="14.[151,1437,153,431]" pageId="14" pageNumber="101">
<emphasis id="B916B93BFFD3FFD0FF0B2E1F7866FEC0" bold="true" box="[199,298,332,359]" pageId="14" pageNumber="101">Habitat.</emphasis>
As with
<taxonomicName id="4C621EAAFFD3FFD0FE5C2E1D7B67FEC0" authority="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma, 2020" authorityName="Liu &amp; Yin &amp; Haddad &amp; Xu &amp; Ma" authorityYear="2020" box="[400,555,333,359]" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="101" phylum="Arthropoda" rank="species" species="subfabaria" status="sp. nov.">
<emphasis id="B916B93BFFD3FFD0FE5C2E1D7B67FEC0" box="[400,555,333,359]" italics="true" pageId="14" pageNumber="101">U. subfabaria</emphasis>
</taxonomicName>
<emphasis id="B916B93BFFD3FFD0FDFE2E1E7BC0FEC0" bold="true" box="[562,652,333,359]" pageId="14" pageNumber="101">
<taxonomicNameLabel id="A2250440FFD3FFD0FDFE2E1E7BC0FEC0" box="[562,652,333,359]" pageId="14" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, this species can be found in leaf litter in parts of rainforest with karst physiognomy.
</paragraph>
</subSubSection>
<subSubSection id="C37836A2FFD3FFD0FF0B2EC77C28FE08" box="[199,1380,404,431]" pageId="14" pageNumber="101" type="distribution">
<paragraph id="8BDD6529FFD3FFD0FF0B2EC77C28FE08" blockId="14.[151,1437,153,431]" box="[199,1380,404,431]" pageId="14" pageNumber="101">
<emphasis id="B916B93BFFD3FFD0FF0B2EC7782DFE08" bold="true" box="[199,353,404,431]" pageId="14" pageNumber="101">Distribution.</emphasis>
Known only from the
<typeStatus id="54D9DB8BFFD3FFD0FDAC2EC67BDCFE08" box="[608,656,405,431]" pageId="14" pageNumber="101">type</typeStatus>
locality,
<collectingRegion id="49A6ABCBFFD3FFD0FD342EC67AF9FE08" box="[760,949,405,431]" country="China" name="Guangxi" pageId="14" pageNumber="101">Guangxi Zhuang</collectingRegion>
Autonomous Region,
<collectingCountry id="F37525B9FFD3FFD0FB7D2EC67DB8FE08" box="[1201,1268,405,431]" name="China" pageId="14" pageNumber="101">China</collectingCountry>
(
<figureCitation id="135979ACFFD3FFD0FACF2EC67C1AFE08" box="[1283,1366,405,431]" captionStart="FIGURE 10" captionStartId="13.[151,250,1822,1847]" captionTargetBox="[185,1397,218,1766]" captionTargetId="figure@13.[169,1418,181,1797]" captionTargetPageId="13" captionText="FIGURE 10. Collection localities for Utivarachna subfabaria sp. nov. and U. tangi sp. nov. in China." figureDoi="http://doi.org/10.5281/zenodo.3908045" httpUri="https://zenodo.org/record/3908045/files/figure.png" pageId="14" pageNumber="101">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>