192 lines
19 KiB
XML
192 lines
19 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5700103" ID-GBIF-Dataset="8d8c1ff6-4422-428a-9f44-5c0d16451c25" ID-GBIF-Taxon="190304787" ID-ISSN="1211-87 8 8" ID-Zenodo-Dep="5700103" approvedBy="valdenar" checkinTime="1636646813558" checkinUser="valdenar" docAuthor="Löbl, Ivan" docDate="2021" docId="03AE7920FF9CFFF45038FD8E02BF4DCD" docLanguage="en" docName="ActaMusMoraviae.106.2.227-248.pdf" docOrigin="Acta Musei Moraviae 106 (2)" docTitle="Bironium grande Löbl 2021, sp. nov." docType="treatment" docVersion="10" lastPageId="9" lastPageNumber="234" masterDocId="FF970158FF9BFFFD515DFFB103354C2C" masterDocTitle="A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas" masterLastPageNumber="248" masterPageNumber="227" pageId="7" pageNumber="234" updateTime="1637334954976" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Löbl, Ivan</mods:namePart>
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<mods:affiliation>MusØum d ̓ histoire naturelle, Case postale 6434, 1211 Genève, Switzerland</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Acta Musei Moraviae</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-11-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>106</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>227</mods:start>
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<mods:end>248</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.5706173</mods:identifier>
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<mods:identifier type="GBIF-Dataset">8d8c1ff6-4422-428a-9f44-5c0d16451c25</mods:identifier>
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<mods:identifier type="ISSN">1211-8788</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5706173</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5700103" ID-GBIF-Taxon="190304787" ID-Zenodo-Dep="5700103" LSID="urn:lsid:plazi:treatment:03AE7920FF9CFFF45038FD8E02BF4DCD" httpUri="http://treatment.plazi.org/id/03AE7920FF9CFFF45038FD8E02BF4DCD" lastPageId="9" lastPageNumber="234" pageId="7" pageNumber="234">
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<subSubSection box="[357,643,575,600]" pageId="7" pageNumber="234" type="nomenclature">
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<paragraph blockId="7.[357,643,575,600]" box="[357,643,575,600]" pageId="7" pageNumber="234">
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<heading bold="true" box="[357,643,575,600]" fontSize="10" level="3" pageId="7" pageNumber="234" reason="2">
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<emphasis bold="true" box="[357,635,575,600]" pageId="7" pageNumber="234">
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<taxonomicName authority="Löbl, 2021" authorityName="Löbl" authorityYear="2021" box="[357,548,575,599]" class="Insecta" family="Staphylinidae" genus="Bironium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="234" phylum="Arthropoda" rank="species" species="grande" status="sp. nov.">
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<emphasis bold="true" box="[357,548,575,599]" italics="true" pageId="7" pageNumber="234">Bironium grande</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[555,643,576,600]" pageId="7" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection box="[1072,1210,574,600]" pageId="7" pageNumber="234" type="description">
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<paragraph blockId="7.[1072,1210,574,600]" box="[1072,1210,574,600]" pageId="7" pageNumber="234">
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(
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<figureCitation box="[1079,1159,574,600]" captionStart="Figs 11-19" captionStartId="8.[332,370,1698,1717]" captionTargetBox="[350,1258,391,1670]" captionTargetId="figure-104@8.[349,1262,390,1675]" captionTargetPageId="8" captionText="Figs 11-19. 1114, Bironium biroi (Pic): 11 Aedeagus in lateral view, scale = 0.2 mm; 12 Internal sac, scale = 0.05 mm; 13 Paramere in ventral view, scale = 0.1 mm; 14 Gonocoxite, scale = 0.05 mm. 1519, Bironium grande sp. nov.: 15, 16 Aedeagus in dorsal and lateral views, scale = 0.4 mm; 17 Paramere in ventral view, scale = 0.2 mm; 18 Internal sac, scale = 0.05 mm; 19 Gonocoxite, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5706179" httpUri="https://zenodo.org/record/5706179/files/figure.png" pageId="7" pageNumber="234">Figs 15</figureCitation>
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19)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="234" type="materials_examined">
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<paragraph blockId="7.[304,1252,620,693]" pageId="7" pageNumber="234">
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<emphasis bold="true" box="[304,440,622,642]" pageId="7" pageNumber="234">Type material.</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3400798335" collectingDate="1998-03-12" collectingDateMax="1998-03-12" collectingDateMin="1998-03-11" collectionCode="MHNG" collectorName="A. Riedel" country="PAPUA N. G." elevation="1600-1700" location="Morobe Prov." pageId="7" pageNumber="234" specimenCount="1" specimenCount-male="1" stateProvince="Morobe Prov." typeStatus="holotype">
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<typeStatus box="[451,530,623,642]" pageId="7" pageNumber="234">Holotype</typeStatus>
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<specimenCount box="[540,556,620,642]" pageId="7" pageNumber="234" type="male">♂</specimenCount>
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,
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<collectingCountry box="[572,703,623,642]" pageId="7" pageNumber="234">
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<taxonomicName authority="Löbl, 2021" authorityName="Löbl" authorityYear="2021" box="[572,640,623,642]" class="Insecta" family="Pyralidae" genus="Papua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="234" phylum="Arthropoda" rank="genus" status="N. G.">PAPUA</taxonomicName>
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<taxonomicNameLabel box="[650,703,623,642]" pageId="7" pageNumber="234" rank="genus">N. G.</taxonomicNameLabel>
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</collectingCountry>
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<collectingRegion box="[712,835,623,642]" country="Papua New Guinea" name="Morobe" pageId="7" pageNumber="234">Morobe Prov.</collectingRegion>
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leg.
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<collectorName box="[884,971,623,642]" pageId="7" pageNumber="234">A. Riedel</collectorName>
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/ Aseki, Oiwa,
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<elevation box="[1123,1248,621,642]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="234" unit="m" value="1700.0">
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1600-
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<quantity box="[1177,1248,621,642]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="234" unit="m" value="1700.0">1700 m</quantity>
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</elevation>
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,
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<date box="[304,431,647,668]" pageId="7" pageNumber="234" value="1998-03-12">
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11-
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<collectingDate box="[336,431,647,668]" pageId="7" pageNumber="234" value="1998-03-12">12.III.1998</collectingDate>
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</date>
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(
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<collectionCode box="[446,516,648,667]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="7" pageNumber="234" type="Museum">MHNG</collectionCode>
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).
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</materialsCitation>
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<materialsCitation ID-GBIF-Occurrence="3400798308" collectingDate="1957-11" collectionCode="MHNG" collectorName="J. Sedlacek" country="Papua New Guinea" location="Kassem Pass" pageId="7" pageNumber="234" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Highlands" typeStatus="paratype">
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<typeStatus box="[534,609,649,668]" pageId="7" pageNumber="234">Paratype</typeStatus>
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<specimenCount box="[618,634,646,668]" pageId="7" pageNumber="234" type="female">♀</specimenCount>
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,
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<collectingCountry box="[649,825,648,668]" name="Papua New Guinea" pageId="7" pageNumber="234">
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<taxonomicName authority="Löbl, 2021" authorityName="Löbl" authorityYear="2021" box="[649,701,649,668]" class="Insecta" family="Pyralidae" genus="Papua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="234" phylum="Arthropoda" rank="genus" status="N. G.">Papua</taxonomicName>
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New Guinea
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</collectingCountry>
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,
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<collectingRegion box="[837,1000,648,668]" country="Papua New Guinea" name="Eastern Highlands" pageId="7" pageNumber="234">Eastern Highlands</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:03AE7920FF9CFFF45038FD8E02BF4DCD:8ED89EEDFF9CFFFA52A8FD38075F4EB0" box="[1013,1130,649,668]" country="Papua New Guinea" name="Kassem Pass" pageId="7" pageNumber="234" stateProvince="Eastern Highlands">Kassem Pass</location>
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,
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<date box="[1143,1225,648,668]" pageId="7" pageNumber="234" value="1957-11">
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<collectingDate box="[1143,1225,648,668]" pageId="7" pageNumber="234" value="1957-11">XI. 1957</collectingDate>
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</date>
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,
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<collectorName pageId="7" pageNumber="234">J. Sedlacek</collectorName>
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(
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<collectionCode box="[393,463,674,693]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="7" pageNumber="234" type="Museum">MHNG</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="234" type="description">
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<paragraph blockId="7.[304,1251,720,1761]" pageId="7" pageNumber="234">
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<emphasis bold="true" box="[304,443,722,746]" pageId="7" pageNumber="234">Description.</emphasis>
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Length 3.0-
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<quantity box="[584,684,720,746]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="234" unit="mm" value="3.2">3.20 mm</quantity>
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, width 1.77 -
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<quantity box="[840,939,720,746]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="7" pageNumber="234" unit="mm" value="1.92">1.92 mm</quantity>
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. Head and body black, with apex of abdomen lighter. Femora and tibiae black, tarsi blackishbrown. Antennae blackish. Pronotum very finely punctate. Scutellum rounded. Elytra with distinct humeral humps; adsutural areas raised; sutural striae with scattered, fine punctures; basal striae shortened, broadly interrupted laterally of elytral midwidth, lateral striae finely punctate in anterior halves. Elytra lacking sulci and impressions, punctation near sutural and lateral striae, and at bases very fine, similar to pronotal punctation, on apical fourth somewhat less fine, on prevailing discal surface rather coarse and irregular. Mesoventrite hardly convex, with robust mesal ridge nearly reaching margin of mesocoxal process, and with oblique striae. Metaventrite impunctate, not microsculptured; impressed mesally; mesocoxal process below level of mesoventrite, longitudinally striate. Metacoxal process slightly oblique, with transverse margin. Metanepisterna narrow, near anterior margins about
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<quantity box="[377,478,1106,1130]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="234" unit="mm" value="0.04">0.04 mm</quantity>
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||
wide, weakly widened apically. Metanepisternal suture impunctate, straight and nearly reaching margin of mesepimeron. Abdomen very finely punctate, except for to distinct punctures along basal margin of ventrite I. Mesal area of ventrite I lacking basal bump, with strigulate microsculpture becoming punctulate near apical margin, lateral parts of ventrite Inot microsculptured. Ventrites II to IV and base of ventrite Vwith strigulate microsculpture, prevailing surface of ventrite Vwith punctulate microsculpture. Protarsomeres III lacking prominent ventral lobe.
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</paragraph>
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||
<paragraph blockId="7.[304,1251,720,1761]" pageId="7" pageNumber="234">
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||
<specimenCount box="[357,377,1327,1354]" pageId="7" pageNumber="234">♂</specimenCount>
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||
: Protarsomeres Ito III moderately widened, similar in width, each about as wide as two thirds of protibial apex. Protarsomere III longer than following three tarsomeres combined, with apicoventral pair of long setae not widened. Aedeagus as
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<figureCitation box="[1122,1204,1392,1418]" captionStart="Figs 11-19" captionStartId="8.[332,370,1698,1717]" captionTargetBox="[350,1258,391,1670]" captionTargetId="figure-104@8.[349,1262,390,1675]" captionTargetPageId="8" captionText="Figs 11-19. 1114, Bironium biroi (Pic): 11 Aedeagus in lateral view, scale = 0.2 mm; 12 Internal sac, scale = 0.05 mm; 13 Paramere in ventral view, scale = 0.1 mm; 14 Gonocoxite, scale = 0.05 mm. 1519, Bironium grande sp. nov.: 15, 16 Aedeagus in dorsal and lateral views, scale = 0.4 mm; 17 Paramere in ventral view, scale = 0.2 mm; 18 Internal sac, scale = 0.05 mm; 19 Gonocoxite, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5706179" httpUri="https://zenodo.org/record/5706179/files/figure.png" pageId="7" pageNumber="234">Figs 15</figureCitation>
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-18,
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||
<quantity box="[304,399,1426,1450]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="234" unit="mm" value="1.5">1.50 mm</quantity>
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long.
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||
</paragraph>
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||
<paragraph blockId="7.[304,1251,720,1761]" box="[357,808,1455,1482]" pageId="7" pageNumber="234">
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||
<specimenCount box="[357,377,1455,1482]" pageId="7" pageNumber="234">♀</specimenCount>
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: Gonocoxite with gonostyle, as
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<figureCitation box="[725,804,1458,1482]" captionStart="Figs 11-19" captionStartId="8.[332,370,1698,1717]" captionTargetBox="[350,1258,391,1670]" captionTargetId="figure-104@8.[349,1262,390,1675]" captionTargetPageId="8" captionText="Figs 11-19. 1114, Bironium biroi (Pic): 11 Aedeagus in lateral view, scale = 0.2 mm; 12 Internal sac, scale = 0.05 mm; 13 Paramere in ventral view, scale = 0.1 mm; 14 Gonocoxite, scale = 0.05 mm. 1519, Bironium grande sp. nov.: 15, 16 Aedeagus in dorsal and lateral views, scale = 0.4 mm; 17 Paramere in ventral view, scale = 0.2 mm; 18 Internal sac, scale = 0.05 mm; 19 Gonocoxite, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5706179" httpUri="https://zenodo.org/record/5706179/files/figure.png" pageId="7" pageNumber="234">Fig. 19</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection box="[304,672,1497,1522]" pageId="7" pageNumber="234" type="distribution">
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<paragraph blockId="7.[304,1251,720,1761]" box="[304,672,1497,1522]" pageId="7" pageNumber="234">
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<emphasis bold="true" box="[304,450,1497,1521]" pageId="7" pageNumber="234">Distribution.</emphasis>
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<collectingCountry box="[457,668,1498,1522]" name="Papua New Guinea" pageId="7" pageNumber="234">
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<taxonomicName authority="Löbl, 2021" authorityName="Löbl" authorityYear="2021" box="[457,522,1498,1521]" class="Insecta" family="Pyralidae" genus="Papua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="234" phylum="Arthropoda" rank="genus" status="N. G.">Papua</taxonomicName>
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New Guinea
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</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection box="[304,1023,1537,1561]" pageId="7" pageNumber="234" type="etymology">
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<paragraph blockId="7.[304,1251,720,1761]" box="[304,1023,1537,1561]" pageId="7" pageNumber="234">
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<emphasis bold="true" box="[304,432,1537,1561]" pageId="7" pageNumber="234">Etymology.</emphasis>
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The species epithet is a Latin adjective meaning large.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="9" lastPageNumber="236" pageId="7" pageNumber="234" type="discussion">
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<paragraph blockId="7.[304,1251,720,1761]" lastBlockId="9.[304,1251,393,481]" lastPageId="9" lastPageNumber="236" pageId="7" pageNumber="234">
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<emphasis bold="true" box="[304,434,1577,1601]" pageId="7" pageNumber="234">Comments.</emphasis>
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The species may be distinguished from its Papuan congeners by the bodysize. Only four Asian species,
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<taxonomicName authority="(Achard, 1914)" baseAuthorityName="Achard" baseAuthorityYear="1914" box="[638,887,1609,1633]" class="Insecta" family="Staphylinidae" genus="Bironium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="234" phylum="Arthropoda" rank="species" species="feai">
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<emphasis box="[638,710,1609,1633]" italics="true" pageId="7" pageNumber="234">B. feai</emphasis>
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(
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<bibRefCitation author="ACHARD J." box="[726,879,1609,1633]" journalOrPublisher="Bulletin de la SociOtO entomologique de France" pageId="7" pageNumber="234" pagination="394" part="1914" refId="ref9778" refString="ACHARD J. 1914: Un scaphidiide nouveau de Birmanie (Col.). Bulletin de la SociOtO entomologique de France 1914: 394 396." title="Un scaphidiide nouveau de Birmanie (Col.)" type="journal article" year="1914">Achard, 1914</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicName authority="(Achard, 1920)" baseAuthorityName="Achard" baseAuthorityYear="1920" box="[899,1215,1609,1633]" class="Insecta" family="Staphylinidae" genus="Bironium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="234" phylum="Arthropoda" rank="species" species="grouvellei">
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<emphasis box="[899,1038,1609,1633]" italics="true" pageId="7" pageNumber="234">B. grouvellei</emphasis>
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(Achard, 1920)
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</taxonomicName>
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,
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<taxonomicName authority="(Pic, 1922)" baseAuthorityName="Pic" baseAuthorityYear="1922" class="Insecta" family="Staphylinidae" genus="Bironium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="234" phylum="Arthropoda" rank="species" species="tonkineum">
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<emphasis italics="true" pageId="7" pageNumber="234">B. tonkineum</emphasis>
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(Pic, 1922)
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</taxonomicName>
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and
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<taxonomicName authority="Lobl, 2020" authorityName="Lobl" authorityYear="2020" box="[595,865,1641,1665]" class="Insecta" family="Staphylinidae" genus="Bironium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="234" phylum="Arthropoda" rank="species" species="yunnanum">
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<emphasis box="[595,737,1642,1665]" italics="true" pageId="7" pageNumber="234">B. yunnanum</emphasis>
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<bibRefCitation author="LOBL I. & LESCHEN R. A. B. & KODADA J." box="[744,865,1641,1665]" journalOrPublisher="Annales Zoologici (Warszawa)" pageId="7" pageNumber="234" pagination="711" part="70" refId="ref10053" refString="LOBL I., LESCHEN R. A. B. & KODADA J. 2020: Review of the Asian species and cladistic analysis of Bironium Csiki (Coleoptera: Staphylinidae: Scaphidiinae) with comments on biogeography. Annales Zoologici (Warszawa) 70 (4): 711 736." title="Review of the Asian species and cladistic analysis of Bironium Csiki (Coleoptera: Staphylinidae: Scaphidiinae) with comments on biogeography" type="journal article" year="2020">Löbl, 2020</bibRefCitation>
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</taxonomicName>
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, have the body
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<quantity box="[1035,1097,1641,1665]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="234" unit="mm" value="3.0">3 mm</quantity>
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long, or more than
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<quantity box="[358,421,1673,1697]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="234" unit="mm" value="3.0">3 mm</quantity>
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long, in combination with the entirely black pronotum and elytra. This new species differs also conspicuously from the Papuan congeners by the punctate ventrites and the ventrite Ihaving microsculpture to part consisting of striae. The narrow notch situated posterior of the expanded apical section of the parameres is a unique character, and the gonostyle separates this species from the remaining Papuan
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<taxonomicName authorityName="Csiki" authorityYear="1909" box="[1042,1142,426,449]" class="Insecta" family="Staphylinidae" genus="Bironium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="236" phylum="Arthropoda" rank="genus">
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<emphasis box="[1042,1142,426,449]" italics="true" pageId="9" pageNumber="236">Bironium</emphasis>
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</taxonomicName>
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known in females.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |