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<document ID-DOI="10.11646/zootaxa.5277.1.3" ID-GBIF-Dataset="195c7011-1a00-4c5f-a339-dc529aa9e104" ID-ISSN="1175-5326" ID-Zenodo-Dep="7891898" ID-ZooBank="2EBD6E19-89E9-4918-8EE3-3324DF3BBA13" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1682930661848" checkinUser="plazi" docAuthor="Jałoszyński, Paweł, Hlaváč, Peter &amp; Anlaş, Sinan" docDate="2023" docId="03AB722FA10AFFE525C9F8C09535F8A6" docLanguage="en" docName="zootaxa.5277.1.3.pdf" docOrigin="Zootaxa 5277 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Percussiopalpus Jaloszynski &amp; Hlavac 2023, gen. n." docType="treatment" docVersion="3" lastPageNumber="78" masterDocId="FF920A57A10BFFE2255EFFB59768FFDC" masterDocTitle="Percussiopalpus gen. n. from Turkey, a free-living relative of cave-dwelling Thaumastocephalus, and revised placement of Thaumastocephalini (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="90" masterPageNumber="71" pageNumber="72" updateTime="1683121264666" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Percussiopalpus gen. n. from Turkey, a free-living relative of cave-dwelling Thaumastocephalus, and revised placement of Thaumastocephalini (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jałoszyński, Paweł</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-2973-1803</mods:nameIdentifier>
<mods:affiliation>Museum ofNaturalHistory, UniversityofWrocław, Sienkiewicza 21, 50 - 335 Wrocław, Poland. https: // orcid. org / 0000 - 0003 - 2973 - 1803 Department of Entomology, National Museum, Natural History Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, The Czech Republic. peterhlavac @ gmail. com; https: // orcid. org / 0000 - 0001 - 5060 - 0811</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hlaváč, Peter</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-5060-0811</mods:nameIdentifier>
<mods:nameIdentifier type="email">peterhlavac@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Anlaş, Sinan</mods:namePart>
<mods:affiliation>Manisa Celal Bayar University, Alaşehir Vocational School, TR- 45600, Alaşehir, Manisa, Turkey.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5277</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>71</mods:start>
<mods:end>90</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5277.1.3</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5277.1.3</mods:identifier>
<mods:identifier type="GBIF-Dataset">195c7011-1a00-4c5f-a339-dc529aa9e104</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7891898</mods:identifier>
<mods:identifier type="ZooBank">2EBD6E19-89E9-4918-8EE3-3324DF3BBA13</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7891948" ID-Zenodo-Dep="7891948" LSID="urn:lsid:plazi:treatment:03AB722FA10AFFE525C9F8C09535F8A6" httpUri="http://treatment.plazi.org/id/03AB722FA10AFFE525C9F8C09535F8A6" lastPageId="7" lastPageNumber="78" pageId="1" pageNumber="72">
<subSubSection box="[151,723,1908,1936]" pageId="1" pageNumber="72" type="nomenclature">
<paragraph blockId="1.[151,723,1908,1971]" box="[151,723,1908,1936]" pageId="1" pageNumber="72">
<heading bold="true" box="[151,723,1908,1936]" fontSize="11" level="1" pageId="1" pageNumber="72" reason="1">
<emphasis bold="true" box="[151,723,1908,1936]" pageId="1" pageNumber="72">
<taxonomicName authority="Jaloszynski &amp; Hlavac" authorityName="Jaloszynski &amp; Hlavac" authorityYear="2023" box="[151,628,1908,1936]" class="Insecta" family="Staphylinidae" genus="Percussiopalpus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis bold="true" box="[151,348,1909,1936]" italics="true" pageId="1" pageNumber="72">Percussiopalpus</emphasis>
Jałoszyński &amp; Hlaváč
</taxonomicName>
,
<taxonomicNameLabel box="[641,723,1909,1935]" pageId="1" pageNumber="72" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,281,1945,1971]" pageId="1" pageNumber="72" type="description">
<paragraph blockId="1.[151,723,1908,1971]" box="[151,281,1945,1971]" pageId="1" pageNumber="72">
(
<figureCitation box="[159,272,1945,1971]" captionStart-0="FIGURES" captionStart-1="FIGURES 3–7" captionStart-2="FIGURES 8–13" captionStart-3="FIGURES 14–18" captionStart-4="FIGURES 19–23" captionStart-5="FIGURES 24–30" captionStart-6="FIGURES31–38" captionStart-7="FIGURES 39–40" captionStartId-0="2.[151,179,1725,1751]" captionStartId-1="3.[151,264,1751,1775]" captionStartId-2="5.[151,264,1836,1860]" captionStartId-3="6.[151,264,1030,1054]" captionStartId-4="7.[151,264,1457,1481]" captionStartId-5="9.[151,264,1860,1884]" captionStartId-6="10.[151,264,1945,1969]" captionStartId-7="11.[151,264,1947,1971]" captionTargetBox-0="[173,1416,513,1671]" captionTargetBox-1="[151,1436,381,1713]" captionTargetBox-2="[151,1436,181,1812]" captionTargetBox-3="[151,1436,181,1006]" captionTargetBox-4="[151,1436,181,1433]" captionTargetBox-5="[151,1436,345,1837]" captionTargetBox-6="[151,1436,885,1908]" captionTargetBox-7="[151,1436,453,1924]" captionTargetId-0="figure-140@2.[151,1436,486,1700]" captionTargetId-1="figure-86@3.[151,1436,381,1728]" captionTargetId-2="figure-18@5.[151,1436,181,1812]" captionTargetId-3="figure-18@6.[151,1436,181,1006]" captionTargetId-4="figure-18@7.[151,1436,181,1433]" captionTargetId-5="figure-90@9.[151,1436,345,1837]" captionTargetId-6="figure-284@10.[151,1436,885,1921]" captionTargetId-7="figure-101@11.[151,1436,453,1924]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionText-0="FIGURES 1–2. Percussiopalpus inusitatus sp. n. Dorsal habitus of male (1) and female (2)." captionText-1="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." captionText-2="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." captionText-3="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." captionText-4="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." captionText-5="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." captionText-6="FIGURES31–38. Percussiopalpusinusitatussp. n. Protarsusin ventral(31) andlateral (32) views, metatarsus inlateral view (33), aedeagus in abparameral (34, 36), parameral (37) and lateral (35, 38) views. Abbreviations: pr, paramere; t1‒3, tarsomere 1‒3." captionText-7="FIGURES 39–40. Percussiopalpus inusitatus sp. n. Habitat in Balıkesir Province in Turkey. Collecting site in Erdemit, 202 m, 40°27.610”N, 27°54.605” (39); collecting site in Erdek, Kirazlı Manastırı road, 358 m, 40°26.742”N, 27°54.182” (40)." figureDoi-0="http://doi.org/10.5281/zenodo.7891902" figureDoi-1="http://doi.org/10.5281/zenodo.7891906" figureDoi-2="http://doi.org/10.5281/zenodo.7891910" figureDoi-3="http://doi.org/10.5281/zenodo.7891914" figureDoi-4="http://doi.org/10.5281/zenodo.7891916" figureDoi-5="http://doi.org/10.5281/zenodo.7891922" figureDoi-6="http://doi.org/10.5281/zenodo.7891926" figureDoi-7="http://doi.org/10.5281/zenodo.7891930" httpUri-0="https://zenodo.org/record/7891902/files/figure.png" httpUri-1="https://zenodo.org/record/7891906/files/figure.png" httpUri-2="https://zenodo.org/record/7891910/files/figure.png" httpUri-3="https://zenodo.org/record/7891914/files/figure.png" httpUri-4="https://zenodo.org/record/7891916/files/figure.png" httpUri-5="https://zenodo.org/record/7891922/files/figure.png" httpUri-6="https://zenodo.org/record/7891926/files/figure.png" httpUri-7="https://zenodo.org/record/7891930/files/figure.png" pageId="1" pageNumber="72">Figs 1‒40</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,626,2014,2038]" pageId="1" pageNumber="72" type="type_taxon">
<paragraph blockId="1.[151,626,2014,2038]" box="[151,626,2014,2038]" pageId="1" pageNumber="72">
<typeStatus box="[151,201,2014,2038]" pageId="1" pageNumber="72">Type</typeStatus>
species.
<taxonomicName authority="Jałoszyński &amp; Hlaváč &amp; Anlaş, 2023" authorityName="Jałoszyński &amp; Hlaváč &amp; Anlaş" authorityYear="2023" box="[294,565,2014,2038]" class="Insecta" family="Staphylinidae" genus="Percussiopalpus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="inusitatus" status="sp. n.">
<emphasis box="[294,565,2014,2038]" italics="true" pageId="1" pageNumber="72">Percussiopalpus inusitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[571,626,2014,2037]" pageId="1" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="73" type="diagnosis">
<paragraph blockId="2.[151,1437,150,465]" pageId="2" pageNumber="73">
<emphasis bold="true" box="[151,274,150,176]" pageId="2" pageNumber="73">Diagnosis.</emphasis>
<taxonomicName authorityName="Jałoszyński &amp; Hlaváč &amp; Anlaş" authorityYear="2023" box="[289,516,151,177]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="71" phylum="Arthropoda" rank="tribe" tribe="Thaumastocephalini">Thaumastocephalini</taxonomicName>
with sharply delimited frontal sulcus; frontal rostrum gradually broadening posteriorly; tempora abruptly delimited from neck region; posterior tentorial pits situated closer to neck region than to submental bulge; maxillary palpomere 4 with convex distal margin; pronotum approximately cordiform, with lateral foveae, lateral longitudinal sulci and median longitudinal sulcus; mesoscutellar shield about as long as broad; metaventrite with only one pair of foveae (laterad mesocoxal insertion); each elytron with two basal foveae, complete sutural stria and incomplete discal stria; tergite IV with discal carinae and one pair of lateral foveae; sternite IV with pair of submedian foveae directed mesally.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="78" pageId="2" pageNumber="73" type="description">
<paragraph blockId="2.[151,1437,150,465]" pageId="2" pageNumber="73">
<emphasis bold="true" box="[199,345,402,428]" pageId="2" pageNumber="73">Description.</emphasis>
Body (
<figureCitation box="[434,541,403,429]" captionStart="FIGURES" captionStartId="2.[151,179,1725,1751]" captionTargetBox="[173,1416,513,1671]" captionTargetId="figure-140@2.[151,1436,486,1700]" captionTargetPageId="2" captionText="FIGURES 1–2. Percussiopalpus inusitatus sp. n. Dorsal habitus of male (1) and female (2)." figureDoi="http://doi.org/10.5281/zenodo.7891902" httpUri="https://zenodo.org/record/7891902/files/figure.png" pageId="2" pageNumber="73">Figs 1, 2</figureCitation>
) elongate, moderately slender, moderately convex, with distinct constrictions between head and prothorax and between prothorax and elytra.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7891902" ID-Zenodo-Dep="7891902" box="[151,1090,1725,1751]" httpUri="https://zenodo.org/record/7891902/files/figure.png" pageId="2" pageNumber="73" startId="2.[151,179,1725,1751]" targetBox="[173,1416,513,1671]" targetPageId="2" targetType="figure">
<paragraph blockId="2.[151,1090,1725,1751]" box="[151,1090,1725,1751]" pageId="2" pageNumber="73">
<emphasis bold="true" box="[151,317,1725,1751]" pageId="2" pageNumber="73">FIGURES 1–2.</emphasis>
<taxonomicName authority="Jałoszyński &amp; Hlaváč &amp; Anlaş, 2023" authorityName="Jałoszyński &amp; Hlaváč &amp; Anlaş" authorityYear="2023" box="[323,595,1726,1750]" class="Insecta" family="Staphylinidae" genus="Percussiopalpus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="inusitatus" status="sp. n.">
<emphasis box="[323,595,1726,1750]" italics="true" pageId="2" pageNumber="73">Percussiopalpus inusitatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[601,658,1727,1751]" pageId="2" pageNumber="73">
<taxonomicNameLabel box="[601,658,1727,1751]" pageId="2" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Dorsal habitus of male (
<emphasis bold="true" box="[909,922,1726,1750]" pageId="2" pageNumber="73">1</emphasis>
) and female (
<emphasis bold="true" box="[1063,1076,1726,1750]" pageId="2" pageNumber="73">2</emphasis>
).
</paragraph>
</caption>
<paragraph blockId="2.[151,1437,1797,2039]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="74" pageId="2" pageNumber="73">
Head (
<figureCitation box="[272,371,1797,1823]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="2" pageNumber="73">Figs 3‒7</figureCitation>
) approximately cordiform, weakly elongate, broadest posteriorly, strongly flattened in posterior half. Temples round, distinctly longer than eyes; vertex and frons confluent, lacking frontal fovea and dorsal tentorial pits (but dorsal tentorial arms are fused with vertex on inner surface of head capsule (
<figureCitation box="[1128,1197,1869,1895]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="2" pageNumber="73">Fig. 6</figureCitation>
;
<emphasis box="[1211,1219,1870,1895]" italics="true" pageId="2" pageNumber="73">t</emphasis>
)), head with broad and shallow V-shaped vertexal and transverse rectangular frontal sulcus (
<figureCitation box="[1000,1072,1905,1931]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="2" pageNumber="73">Fig. 3</figureCitation>
;
<emphasis box="[1088,1107,1905,1931]" italics="true" pageId="2" pageNumber="73">fs</emphasis>
), both confluent; compound eyes (
<figureCitation box="[217,321,1941,1967]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="2" pageNumber="73">Figs 3–7</figureCitation>
;
<emphasis box="[335,360,1942,1967]" italics="true" pageId="2" pageNumber="73">ce</emphasis>
) minute and situated in front of middle, slightly below median coronal plane of head capsule. Gular sutures lacking; posterior tentorial pits (
<figureCitation box="[676,780,1977,2004]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="2" pageNumber="73">Figs 5, 7</figureCitation>
;
<emphasis box="[794,830,1978,2003]" italics="true" pageId="2" pageNumber="73">ptp</emphasis>
) large, circular, widely separated, situated in front of transverse impression demarcating neck region ventrally; submental region strongly transverse, with prominent oval submental bulge (
<figureCitation box="[411,514,151,177]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="3" pageNumber="74">Figs 4‒7</figureCitation>
,
<figureCitation box="[527,558,151,177]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="3" pageNumber="74">10</figureCitation>
;
<emphasis box="[571,616,151,177]" italics="true" pageId="3" pageNumber="74">smb</emphasis>
) strongly projecting ventrally; hypostomal ridge (
<figureCitation box="[1187,1272,151,177]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="3" pageNumber="74">Fig. 10</figureCitation>
;
<emphasis box="[1286,1311,151,177]" italics="true" pageId="3" pageNumber="74">hr</emphasis>
) short, not reaching margin of submental bulge.
</paragraph>
<paragraph blockId="3.[151,1437,151,357]" pageId="3" pageNumber="74">
Antennae (
<figureCitation box="[322,423,223,249]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="3" pageNumber="74">Figs 8–9</figureCitation>
) composed of eleven antennomeres, slender, antennal insertions broadly separated. Scape cylindrical and elongate, longer than pedicel, antennal club trimerous and indistinctly demarcated, antennomere 11 elongate, symmetrical, rapidly narrowing in distal region to pointed apex, lacking cavities and clusters of basiconic sensilla.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7891906" ID-Zenodo-Dep="7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="3" pageNumber="74" startId="3.[151,264,1751,1775]" targetBox="[151,1436,381,1713]" targetPageId="3" targetType="figure">
<paragraph blockId="3.[151,1437,1751,1848]" pageId="3" pageNumber="74">
<emphasis bold="true" box="[151,317,1751,1775]" pageId="3" pageNumber="74">FIGURES 3–7.</emphasis>
<taxonomicName authority="Jałoszyński &amp; Hlaváč &amp; Anlaş, 2023" authorityName="Jałoszyński &amp; Hlaváč &amp; Anlaş" authorityYear="2023" box="[325,599,1752,1776]" class="Insecta" family="Staphylinidae" genus="Percussiopalpus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="inusitatus" status="sp. n.">
<emphasis box="[325,599,1752,1776]" italics="true" pageId="3" pageNumber="74">Percussiopalpus inusitatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[607,666,1752,1776]" pageId="3" pageNumber="74">
<taxonomicNameLabel box="[607,666,1752,1776]" pageId="3" pageNumber="74" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Head in dorsal (
<emphasis bold="true" box="[844,857,1751,1775]" pageId="3" pageNumber="74">3</emphasis>
), lateral (
<emphasis bold="true" box="[960,973,1751,1775]" pageId="3" pageNumber="74">4</emphasis>
,
<emphasis bold="true" box="[988,1001,1751,1775]" pageId="3" pageNumber="74">6</emphasis>
), and ventral (
<emphasis bold="true" box="[1156,1169,1752,1776]" pageId="3" pageNumber="74">5</emphasis>
,
<emphasis bold="true" box="[1183,1196,1752,1776]" pageId="3" pageNumber="74">7</emphasis>
) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium.
</paragraph>
</caption>
<paragraph blockId="3.[151,1436,1894,2028]" pageId="3" pageNumber="74">
Labium (
<figureCitation box="[301,381,1894,1920]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="3" pageNumber="74">Fig. 10</figureCitation>
) with elongate mentum (
<figureCitation box="[659,741,1894,1920]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="3" pageNumber="74">Fig. 10</figureCitation>
;
<emphasis box="[752,786,1895,1920]" italics="true" pageId="3" pageNumber="74">mn</emphasis>
), which is inversely subtrapezoidal, broadest near anterior third, with sinuate lateral margins; prementum short, transverse, lacking ligula; labial palpi (
<figureCitation box="[1219,1304,1930,1956]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="3" pageNumber="74">Fig. 10</figureCitation>
) trimerous, palpomere 1 (
<figureCitation box="[308,392,1966,1992]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="3" pageNumber="74">Fig. 10</figureCitation>
;
<emphasis box="[404,440,1966,1992]" italics="true" pageId="3" pageNumber="74">lp1</emphasis>
) short, nearly annular, palpomere 2 (
<figureCitation box="[860,945,1966,1992]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="3" pageNumber="74">Fig. 10</figureCitation>
;
<emphasis box="[957,993,1966,1992]" italics="true" pageId="3" pageNumber="74">lp2</emphasis>
) strongly elongate and weakly clavate, palpomere 3 setiform, barely discernible, obscured by long outer apical seta of palpomere 2.
</paragraph>
<paragraph blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="75">
Maxilla with large, nearly semicircular cardo (
<figureCitation box="[753,842,151,177]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="4" pageNumber="75">Fig. 10</figureCitation>
;
<emphasis box="[860,886,151,177]" italics="true" pageId="4" pageNumber="75">cd</emphasis>
); basistipes (
<figureCitation box="[1046,1135,151,177]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="4" pageNumber="75">Fig. 10</figureCitation>
;
<emphasis box="[1152,1185,151,177]" italics="true" pageId="4" pageNumber="75">bst</emphasis>
) triangular, elongate, mediostipes (
<figureCitation box="[300,382,187,213]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="4" pageNumber="75">Fig. 10</figureCitation>
;
<emphasis box="[394,433,187,212]" italics="true" pageId="4" pageNumber="75">mst</emphasis>
) subtriangular, about as long as basistipes; galea (
<figureCitation box="[990,1072,187,213]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="4" pageNumber="75">Fig. 10</figureCitation>
;
<emphasis box="[1084,1120,187,213]" italics="true" pageId="4" pageNumber="75">gal</emphasis>
) elongate and with fringe of dense, thick setae along mesal margin; lacinia (not shown) similar to galea but shorter; palpifer elongate. Maxillary palpi (
<figureCitation box="[222,324,259,285]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="4" pageNumber="75">Figs 4–7</figureCitation>
;
<emphasis box="[336,383,259,284]" italics="true" pageId="4" pageNumber="75">mxp</emphasis>
, 12, 13) shorter than head, palpomere 1 (
<figureCitation box="[845,929,259,285]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="4" pageNumber="75">Fig. 12</figureCitation>
;
<emphasis box="[941,1001,259,285]" italics="true" pageId="4" pageNumber="75">mxp1</emphasis>
) minute, elongate, about twice as long as wide; palpomere 2 (
<figureCitation box="[418,504,295,321]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="4" pageNumber="75">Fig. 12</figureCitation>
;
<emphasis box="[519,580,295,321]" italics="true" pageId="4" pageNumber="75">mxp2</emphasis>
) pedunculate, rapidly and strongly broadening in distal half, distal region strongly transverse and projecting outwards, with lateral sensory process inserted on conspicuously long and slender petiole broadened distally to form short collar bearing large spherical structure sparsely covered with flattened setae; palpomere 3 (
<figureCitation box="[310,395,403,429]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="4" pageNumber="75">Fig. 12</figureCitation>
;
<emphasis box="[409,470,403,429]" italics="true" pageId="4" pageNumber="75">mxp3</emphasis>
) non-pedunculate, strongly transverse and projecting outwards, with similar sensory process as palpomere 2; palpomere 4 (
<figureCitation box="[588,672,439,465]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="4" pageNumber="75">Fig. 12</figureCitation>
;
<emphasis box="[685,746,439,465]" italics="true" pageId="4" pageNumber="75">mxp4</emphasis>
) strongly transverse and projecting inwards, elliptical, wider than each of palpomeres 2 and 3, with apical sensory appendage (
<figureCitation box="[883,967,475,501]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="4" pageNumber="75">Fig. 12</figureCitation>
;
<emphasis box="[978,1003,475,500]" italics="true" pageId="4" pageNumber="75">sa</emphasis>
) rod-like, short and slender, pointed at apex, inserted in socket surrounded by distinctly broadened and raised margin.
</paragraph>
<paragraph blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="75">
Mandibles (
<figureCitation box="[330,395,547,573]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="4" pageNumber="75">Fig. 3</figureCitation>
;
<emphasis box="[404,438,547,573]" italics="true" pageId="4" pageNumber="75">md</emphasis>
) subtriangular, slightly convex dorsally and concave ventrally, large, slightly asymmetrical, differing in shapes and numbers of mesal row of several teeth gradually reducing in length towards mandibular base, outer mandibular margin with conspicuously long seta; prostheca and mola lacking.
</paragraph>
<paragraph blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="75">
Labrum (
<figureCitation box="[305,388,655,681]" captionStart="FIGURES 8–13" captionStartId="5.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure-18@5.[151,1436,181,1812]" captionTargetPageId="5" captionText="FIGURES 8–13. Percussiopalpus inusitatus sp. n. Right antenna in dorsal view (8, 9), anteromedian region of head in ventral view (10), mandibles and labrum in ventral view (11), left maxillary palp in ventral view (12), distal region of sensory process of maxillary palpomere 3 in ventral view (13). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process." figureDoi="http://doi.org/10.5281/zenodo.7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="4" pageNumber="75">Fig. 11</figureCitation>
;
<emphasis box="[401,420,655,681]" italics="true" pageId="4" pageNumber="75">lr</emphasis>
) transverse, subtrapezoidal, broadest anteriorly, with nearly straight anterior and rounded lateral margins; anteroventral margin with one pair of short peg-like sensilla (
<figureCitation box="[1067,1140,691,717]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="4" pageNumber="75">Fig. 4</figureCitation>
;
<emphasis box="[1157,1190,691,717]" italics="true" pageId="4" pageNumber="75">pls</emphasis>
) inserted on anterior epipharyngeal region.
</paragraph>
<paragraph blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="75">
Pronotum (
<figureCitation box="[327,409,763,789]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Fig. 14</figureCitation>
) widest distinctly in front of middle; lateral margins strongly rounded in anterior half and indistinctly sinuate in posterior half, anterior pronotal margin only slightly shorter than posterior one, anterior corners poorly marked, strongly obtuse-angled, posterior corners distinct, obtuse-angled; pronotal base arcuate. Pronotal disc with long median (
<figureCitation box="[536,673,871,897]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Figs 14, 17</figureCitation>
;
<emphasis box="[689,728,871,897]" italics="true" pageId="4" pageNumber="75">mls</emphasis>
) and long lateral (
<figureCitation box="[949,1087,871,897]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Figs 14, 17</figureCitation>
;
<emphasis box="[1102,1128,871,897]" italics="true" pageId="4" pageNumber="75">lls</emphasis>
) longitudinal sulci, lateral sulcus posteriorly originating from asetose lateral antebasal fovea (
<figureCitation box="[902,1032,907,933]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Figs 14, 17</figureCitation>
;
<emphasis box="[1044,1074,907,933]" italics="true" pageId="4" pageNumber="75">laf</emphasis>
) with short subcuticular pocket, median antebasal fovea (
<figureCitation box="[427,510,943,969]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Fig. 14</figureCitation>
;
<emphasis box="[521,563,943,969]" italics="true" pageId="4" pageNumber="75">maf</emphasis>
) vestigial, developed as broadening of median longitudinal sulcus and lacking subcuticular pocket; pronotal base with basal and sub-basal pairs of shallow but distinct suboval impressions, and with median impression adjacent to posterior margin.
</paragraph>
<paragraph blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="75">
Prosternum (
<figureCitation box="[342,468,1050,1077]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Figs 15–18</figureCitation>
) fused with hypomera, but with short grooves on anterior margin representing probable rudiments of notosternal sutures; hypomera (
<figureCitation box="[670,806,1086,1113]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Figs 15, 17</figureCitation>
;
<emphasis box="[822,848,1087,1113]" italics="true" pageId="4" pageNumber="75">hy</emphasis>
) broad and posteriorly weakly concave, each with shallow hypomeral groove (
<figureCitation box="[461,588,1123,1149]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Figs 16, 17</figureCitation>
;
<emphasis box="[599,627,1123,1149]" italics="true" pageId="4" pageNumber="75">hg</emphasis>
) clearly visible in lateral view; basisternal region (
<figureCitation box="[1187,1316,1122,1149]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Figs 15‒16</figureCitation>
;
<emphasis box="[1326,1359,1123,1149]" italics="true" pageId="4" pageNumber="75">bst</emphasis>
) much longer than very short coxal region (= furcasternum); prosternum with pair of submedian posterior foveae near middle of posterior basisternal margin, foveal pockets directed dorsally and slightly laterally from body midline (
<figureCitation box="[159,288,1231,1257]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Figs 16, 18</figureCitation>
; arrowheads); anterior prosternal margin weakly crenulate; prosternal process indistinct, not separating procoxae. Hypomera with hypomeral ridges (
<figureCitation box="[670,804,1266,1293]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="4" pageNumber="75">Figs 15, 17</figureCitation>
;
<emphasis box="[818,855,1267,1293]" italics="true" pageId="4" pageNumber="75">hyr</emphasis>
) demarcating conspicuously narrow, parallel-sided inner (adcoxal) regions.
</paragraph>
<paragraph blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="75">
Mesonotum (
<figureCitation box="[351,435,1338,1365]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="4" pageNumber="75">Fig. 25</figureCitation>
) with mesoscutum and mesoscutellum fused (
<figureCitation box="[968,1054,1338,1365]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="4" pageNumber="75">Fig. 25</figureCitation>
;
<emphasis box="[1068,1170,1339,1365]" italics="true" pageId="4" pageNumber="75">sc2+scl2</emphasis>
) forming subtriangular scutellar shield, only its minute tip projects posteriorly beyond transverse ridge demarcating basal articulating lobe and discal region of elytra, so that scutellar shield is not visible between elytral bases in intact specimens.
</paragraph>
<paragraph blockId="4.[151,1437,151,2013]" box="[199,1187,1447,1473]" pageId="4" pageNumber="75">Metanotum (not shown) strongly shortened and weakly sclerotized due to loss of wings.</paragraph>
<paragraph blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="75">
Mesoventrite (
<figureCitation box="[363,493,1483,1509]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Figs 19–23</figureCitation>
;
<emphasis box="[506,532,1483,1509]" italics="true" pageId="4" pageNumber="75">
v
<figureCitation box="[518,532,1483,1509]" captionStart="FIGURES" captionStartId="2.[151,179,1725,1751]" captionTargetBox="[173,1416,513,1671]" captionTargetId="figure-140@2.[151,1436,486,1700]" captionTargetPageId="2" captionText="FIGURES 1–2. Percussiopalpus inusitatus sp. n. Dorsal habitus of male (1) and female (2)." figureDoi="http://doi.org/10.5281/zenodo.7891902" httpUri="https://zenodo.org/record/7891902/files/figure.png" pageId="4" pageNumber="75">2</figureCitation>
</emphasis>
) broadening posteriorly, with massive prepectus (
<figureCitation box="[1096,1226,1483,1509]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Figs 20–23</figureCitation>
;
<emphasis box="[1239,1267,1483,1508]" italics="true" pageId="4" pageNumber="75">pp</emphasis>
); mesoventrite with three pairs of foveae in its anterior half (
<figureCitation box="[662,791,1519,1545]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Figs 20, 21</figureCitation>
, arrowheads): one sublateral fovea directed laterally with ventral opening, one lateral fovea directed anteromesally, and one lateral fovea directed mesally, lateral foveae share one large laterally visible setose opening (
<figureCitation box="[629,714,1591,1617]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Fig. 22</figureCitation>
, upper arrowhead). Median subtrapezoidal area of mesoventrite demarcated laterally by complete ridges. Mesocoxae narrowly separated by elongate, subtriangular mesoventral process (
<figureCitation box="[253,433,1663,1689]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Figs 19, 20, 23</figureCitation>
;
<emphasis box="[448,506,1663,1688]" italics="true" pageId="4" pageNumber="75">msvp</emphasis>
) with rounded apex reaching middle of mesocoxae, meeting, but not fused with anterior tip of anterior metaventral process (
<figureCitation box="[657,835,1699,1725]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Figs 19, 20, 23</figureCitation>
;
<emphasis box="[849,905,1699,1724]" italics="true" pageId="4" pageNumber="75">amtp</emphasis>
), both processes flat. Division between mesoand metaventrite marked with narrow sutures lateral to mesocoxal cavities (visible in
<figureCitation box="[1112,1283,1735,1761]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Figs 20, 22, 23</figureCitation>
), but division obliterated within mesocoxal rests.
</paragraph>
<paragraph blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="75">
Metaventrite (
<figureCitation box="[361,588,1807,1833]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Figs 19, 20, 22, 23</figureCitation>
;
<emphasis box="[603,630,1807,1833]" italics="true" pageId="4" pageNumber="75">
v
<figureCitation box="[616,630,1807,1833]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="4" pageNumber="75">3</figureCitation>
</emphasis>
) strongly transverse and broadening posteriorly; posterior margins of mesocoxal rests carinate; metaventrite with pair of lateral setose foveae directed anteromesally (
<figureCitation box="[1257,1431,1843,1869]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Fig. 20, 22, 23</figureCitation>
; lower arrowhead); metaventral intermetacoxal process (
<figureCitation box="[775,905,1879,1905]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Figs 19, 20</figureCitation>
,
<emphasis box="[916,970,1879,1904]" italics="true" pageId="4" pageNumber="75">mtvp</emphasis>
) broad and clearly separating metacoxae, with posteromedian emargination.
</paragraph>
<paragraph blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="75">
Metendosternite with broad and subquadrate lamellate stem and short lateral furcal arms (
<figureCitation box="[1192,1274,1951,1977]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="4" pageNumber="75">Fig. 20</figureCitation>
;
<emphasis box="[1285,1315,1951,1977]" italics="true" pageId="4" pageNumber="75">lfa</emphasis>
) diverging anteriorly.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7891910" ID-Zenodo-Dep="7891910" httpUri="https://zenodo.org/record/7891910/files/figure.png" pageId="5" pageNumber="76" startId="5.[151,264,1836,1860]" targetBox="[151,1436,181,1812]" targetPageId="5" targetType="figure">
<paragraph blockId="5.[151,1437,1836,2005]" pageId="5" pageNumber="76">
<emphasis bold="true" box="[151,328,1836,1860]" pageId="5" pageNumber="76">FIGURES 8–13.</emphasis>
<taxonomicName authority="Jałoszyński &amp; Hlaváč &amp; Anlaş, 2023" authorityName="Jałoszyński &amp; Hlaváč &amp; Anlaş" authorityYear="2023" box="[334,606,1836,1860]" class="Insecta" family="Staphylinidae" genus="Percussiopalpus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="76" phylum="Arthropoda" rank="species" species="inusitatus" status="sp. n.">
<emphasis box="[334,606,1836,1860]" italics="true" pageId="5" pageNumber="76">Percussiopalpus inusitatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[612,669,1836,1860]" pageId="5" pageNumber="76">
<taxonomicNameLabel box="[612,669,1836,1860]" pageId="5" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Right antenna in dorsal view (
<emphasis bold="true" box="[983,996,1836,1860]" pageId="5" pageNumber="76">8</emphasis>
,
<emphasis bold="true" box="[1008,1021,1836,1860]" pageId="5" pageNumber="76">9</emphasis>
), anteromedian region of head in ventral view (
<emphasis bold="true" box="[214,239,1872,1896]" pageId="5" pageNumber="76">10</emphasis>
), mandibles and labrum in ventral view (
<emphasis bold="true" box="[651,675,1872,1896]" pageId="5" pageNumber="76">11</emphasis>
), left maxillary palp in ventral view (
<emphasis bold="true" box="[1049,1074,1872,1896]" pageId="5" pageNumber="76">12</emphasis>
), distal region of sensory process of maxillary palpomere 3 in ventral view (
<emphasis bold="true" box="[547,572,1908,1932]" pageId="5" pageNumber="76">13</emphasis>
). Abbreviations: bst, basistipes; cd, cardo; gal, galea; hr, hypostomal ridge; lr, labrum; lp1‒2, labial palpomere 1‒2; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pd, pedicel; pls, peg-like sensillum; ptp, posterior tentorial pit; sa, sensory appendage; sc, scape; smb, submental bulge; sp, sensory process.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7891914" ID-Zenodo-Dep="7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="6" pageNumber="77" startId="6.[151,264,1030,1054]" targetBox="[151,1436,181,1006]" targetPageId="6" targetType="figure">
<paragraph blockId="6.[151,1437,1030,1162]" pageId="6" pageNumber="77">
<emphasis bold="true" box="[151,341,1030,1054]" pageId="6" pageNumber="77">FIGURES 14–18.</emphasis>
<taxonomicName authority="Jałoszyński &amp; Hlaváč &amp; Anlaş, 2023" authorityName="Jałoszyński &amp; Hlaváč &amp; Anlaş" authorityYear="2023" box="[348,621,1030,1054]" class="Insecta" family="Staphylinidae" genus="Percussiopalpus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="inusitatus" status="sp. n.">
<emphasis box="[348,621,1030,1054]" italics="true" pageId="6" pageNumber="77">Percussiopalpus inusitatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[628,686,1030,1054]" pageId="6" pageNumber="77">
<taxonomicNameLabel box="[628,686,1030,1054]" pageId="6" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Prothorax in dorsal (
<emphasis bold="true" box="[906,931,1030,1054]" pageId="6" pageNumber="77">14</emphasis>
), ventral (
<emphasis bold="true" box="[1040,1065,1030,1054]" pageId="6" pageNumber="77">15</emphasis>
,
<emphasis bold="true" box="[1079,1104,1030,1054]" pageId="6" pageNumber="77">16</emphasis>
), and left lateral (
<emphasis bold="true" box="[1291,1316,1030,1054]" pageId="6" pageNumber="77">17</emphasis>
,
<emphasis bold="true" box="[1330,1355,1030,1054]" pageId="6" pageNumber="77">18</emphasis>
) views. Abbreviations: bst, basisternal region of prosternum; fs, furcasternal region of prosternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae.
</paragraph>
</caption>
<paragraph blockId="6.[151,1437,1208,1991]" pageId="6" pageNumber="77">
Elytra (
<figureCitation box="[283,371,1208,1234]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="6" pageNumber="77">Figs 22</figureCitation>
,
<figureCitation box="[382,454,1208,1235]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">24, 25</figureCitation>
) together subtrapezoidal, dorsally flattened, each elytron with two asetose basal foveae (
<figureCitation box="[159,249,1244,1270]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Figs 24</figureCitation>
;
<emphasis box="[264,298,1244,1270]" italics="true" pageId="6" pageNumber="77">bef</emphasis>
, 25; arrowheads), minute humeral denticle (
<figureCitation box="[811,896,1244,1271]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Fig. 25</figureCitation>
,
<emphasis box="[911,939,1244,1270]" italics="true" pageId="6" pageNumber="77">hd</emphasis>
), short discal stria (
<figureCitation box="[1172,1257,1244,1270]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Fig. 24</figureCitation>
,
<emphasis box="[1271,1296,1244,1270]" italics="true" pageId="6" pageNumber="77">ds</emphasis>
), and entire sutural stria (
<figureCitation box="[298,380,1280,1306]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Fig. 24</figureCitation>
,
<emphasis box="[392,414,1281,1306]" italics="true" pageId="6" pageNumber="77">ss</emphasis>
). Posterior elytral margin slightly concave and lacking fringe of modified setae.
</paragraph>
<paragraph blockId="6.[151,1437,1208,1991]" pageId="6" pageNumber="77">
Abdomen (
<figureCitation box="[326,415,1316,1342]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="6" pageNumber="77">Figs 19</figureCitation>
,
<figureCitation box="[428,544,1316,1342]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">24, 26–30</figureCitation>
) broadest near middle, weakly convex dorsally and strongly convex ventrally, with well-developed broad paratergites on segments IV–VI. Tergite IV distinctly longer than remaining dorsally visible tergites (i.e., V–VII) combined, with transverse mediobasal impression laterally demarcated by short discal carinae, with one pair of basolateral foveae each adjacent to mesal margin of discal carina and with subcuticular pocket directed laterally (
<figureCitation box="[435,518,1460,1486]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Fig. 28</figureCitation>
; arrowhead). Sternite III (
<figureCitation box="[806,892,1460,1486]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="6" pageNumber="77">Figs 19</figureCitation>
,
<figureCitation box="[903,934,1460,1486]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">26</figureCitation>
;
<emphasis box="[945,992,1461,1486]" italics="true" pageId="6" pageNumber="77">stIII</emphasis>
) short and partly exposed only between metacoxae, with dense fringe of long setae along posterior margin; sternite IV (
<figureCitation box="[1080,1167,1496,1522]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Fig. 26</figureCitation>
;
<emphasis box="[1182,1227,1497,1522]" italics="true" pageId="6" pageNumber="77">stIV</emphasis>
) about as long as remaining visible sternites (i.e., V–VIII in female and V–IX in male) combined, with two pairs of foveae: submedian pair with subcuticular pockets directed mesally and touching at midline (
<figureCitation box="[948,1074,1568,1594]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Figs 27, 29</figureCitation>
, right arrowhead), and sublateral pair with pockets directed slightly mesally (
<figureCitation box="[640,723,1604,1630]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Fig. 27</figureCitation>
; left arrowhead) and strongly dorsally (
<figureCitation box="[1163,1246,1604,1630]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Fig. 29</figureCitation>
; left arrowhead). Male with sternite VIII (
<figureCitation box="[431,516,1640,1666]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Fig. 30</figureCitation>
;
<emphasis box="[529,593,1641,1666]" italics="true" pageId="6" pageNumber="77">stVIII</emphasis>
) slightly modified, with trisinuate posterior margin; male sternite IX with large and elongate median penial plate (
<figureCitation box="[605,690,1676,1702]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">Fig. 30</figureCitation>
;
<emphasis box="[703,748,1677,1702]" italics="true" pageId="6" pageNumber="77">stIX</emphasis>
), lateral portions subtriangular and shorter than penial plate, weakly visible in intact specimens.
</paragraph>
<paragraph blockId="6.[151,1437,1208,1991]" pageId="6" pageNumber="77">
Legs (
<figureCitation box="[271,376,1748,1774]" captionStart="FIGURES" captionStartId="2.[151,179,1725,1751]" captionTargetBox="[173,1416,513,1671]" captionTargetId="figure-140@2.[151,1436,486,1700]" captionTargetPageId="2" captionText="FIGURES 1–2. Percussiopalpus inusitatus sp. n. Dorsal habitus of male (1) and female (2)." figureDoi="http://doi.org/10.5281/zenodo.7891902" httpUri="https://zenodo.org/record/7891902/files/figure.png" pageId="6" pageNumber="77">Figs 1, 2</figureCitation>
,
<figureCitation box="[391,421,1748,1775]" captionStart="FIGURES 14–18" captionStartId="6.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1006]" captionTargetId="figure-18@6.[151,1436,181,1006]" captionTargetPageId="6" captionText="FIGURES 14–18. Percussiopalpus inusitatus sp. n. Prothorax in dorsal (14), ventral (15, 16), and left lateral (17, 18) views. Abbreviations:bst, basisternal region of prosternum; fs, furcasternalregion of prosternum;hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; maf, median antebasal fovea; mls, median longitudinal sulcus. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891914" httpUri="https://zenodo.org/record/7891914/files/figure.png" pageId="6" pageNumber="77">15</figureCitation>
,
<figureCitation box="[435,598,1748,1774]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1457,1481]" captionTargetBox="[151,1436,181,1433]" captionTargetId="figure-18@7.[151,1436,181,1433]" captionTargetPageId="7" captionText="FIGURES 19–23. Percussiopalpus inusitatus sp. n. Pterothorax in ventral (19, 20) and left lateral (22, 23) views, anterior region of mesoventrite in ventral view (21). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae." figureDoi="http://doi.org/10.5281/zenodo.7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="6" pageNumber="77">19, 20, 22, 23</figureCitation>
,
<figureCitation box="[613,643,1748,1774]" captionStart="FIGURES 24–30" captionStartId="9.[151,264,1860,1884]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-90@9.[151,1436,345,1837]" captionTargetPageId="9" captionText="FIGURES 24–30. Percussiopalpus inusitatus sp. n. Pterothorax and abdomen in dorsal view (24), elytral base in dorsal view (25), abdomen in ventral view (26), base of abdomen in ventral (27) and dorsal (28) views, abdomen in anterior view (29), abdominal apex of male in ventral view (30). Abbreviations: bef, basal elytral fovea; ds, discal stria; hd, humeral denticle; sc2+scl2, mesoscutum + mesoscutellum; ss, sutural stria; stIII‒IX, sternite III‒IX; tIII‒VII, tergite IV‒VII.Arrowheads indicate foveae." figureDoi="http://doi.org/10.5281/zenodo.7891922" httpUri="https://zenodo.org/record/7891922/files/figure.png" pageId="6" pageNumber="77">26</figureCitation>
,
<figureCitation box="[657,728,1748,1774]" captionStart="FIGURES31–38" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[151,1436,885,1908]" captionTargetId="figure-284@10.[151,1436,885,1921]" captionTargetPageId="10" captionText="FIGURES31–38. Percussiopalpusinusitatussp. n. Protarsusin ventral(31) andlateral (32) views, metatarsus inlateral view (33), aedeagus in abparameral (34, 36), parameral (37) and lateral (35, 38) views. Abbreviations: pr, paramere; t1‒3, tarsomere 1‒3." figureDoi="http://doi.org/10.5281/zenodo.7891926" httpUri="https://zenodo.org/record/7891926/files/figure.png" pageId="6" pageNumber="77">31–33</figureCitation>
) long and slender, all femora slightly clavate, tibiae and tarsi slender; tarsomere 1 (
<figureCitation box="[392,522,1784,1810]" captionStart="FIGURES31–38" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[151,1436,885,1908]" captionTargetId="figure-284@10.[151,1436,885,1921]" captionTargetPageId="10" captionText="FIGURES31–38. Percussiopalpusinusitatussp. n. Protarsusin ventral(31) andlateral (32) views, metatarsus inlateral view (33), aedeagus in abparameral (34, 36), parameral (37) and lateral (35, 38) views. Abbreviations: pr, paramere; t1‒3, tarsomere 1‒3." figureDoi="http://doi.org/10.5281/zenodo.7891926" httpUri="https://zenodo.org/record/7891926/files/figure.png" pageId="6" pageNumber="77">Figs 31–33</figureCitation>
;
<emphasis box="[533,555,1784,1810]" italics="true" pageId="6" pageNumber="77">
t
<figureCitation box="[542,555,1784,1810]" captionStart="FIGURES" captionStartId="2.[151,179,1725,1751]" captionTargetBox="[173,1416,513,1671]" captionTargetId="figure-140@2.[151,1436,486,1700]" captionTargetPageId="2" captionText="FIGURES 1–2. Percussiopalpus inusitatus sp. n. Dorsal habitus of male (1) and female (2)." figureDoi="http://doi.org/10.5281/zenodo.7891902" httpUri="https://zenodo.org/record/7891902/files/figure.png" pageId="6" pageNumber="77">1</figureCitation>
</emphasis>
) minuscule, tarsomere 2 (
<figureCitation box="[845,974,1784,1810]" captionStart="FIGURES31–38" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[151,1436,885,1908]" captionTargetId="figure-284@10.[151,1436,885,1921]" captionTargetPageId="10" captionText="FIGURES31–38. Percussiopalpusinusitatussp. n. Protarsusin ventral(31) andlateral (32) views, metatarsus inlateral view (33), aedeagus in abparameral (34, 36), parameral (37) and lateral (35, 38) views. Abbreviations: pr, paramere; t1‒3, tarsomere 1‒3." figureDoi="http://doi.org/10.5281/zenodo.7891926" httpUri="https://zenodo.org/record/7891926/files/figure.png" pageId="6" pageNumber="77">Figs 31–33</figureCitation>
;
<emphasis box="[986,1008,1784,1810]" italics="true" pageId="6" pageNumber="77">
t
<figureCitation box="[994,1008,1784,1810]" captionStart="FIGURES" captionStartId="2.[151,179,1725,1751]" captionTargetBox="[173,1416,513,1671]" captionTargetId="figure-140@2.[151,1436,486,1700]" captionTargetPageId="2" captionText="FIGURES 1–2. Percussiopalpus inusitatus sp. n. Dorsal habitus of male (1) and female (2)." figureDoi="http://doi.org/10.5281/zenodo.7891902" httpUri="https://zenodo.org/record/7891902/files/figure.png" pageId="6" pageNumber="77">2</figureCitation>
</emphasis>
) longer than tarsomere 3 (
<figureCitation box="[1302,1432,1784,1810]" captionStart="FIGURES31–38" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[151,1436,885,1908]" captionTargetId="figure-284@10.[151,1436,885,1921]" captionTargetPageId="10" captionText="FIGURES31–38. Percussiopalpusinusitatussp. n. Protarsusin ventral(31) andlateral (32) views, metatarsus inlateral view (33), aedeagus in abparameral (34, 36), parameral (37) and lateral (35, 38) views. Abbreviations: pr, paramere; t1‒3, tarsomere 1‒3." figureDoi="http://doi.org/10.5281/zenodo.7891926" httpUri="https://zenodo.org/record/7891926/files/figure.png" pageId="6" pageNumber="77">Figs 31–33</figureCitation>
;
<emphasis box="[151,173,1820,1846]" italics="true" pageId="6" pageNumber="77">
t
<figureCitation box="[159,173,1820,1846]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[151,1436,381,1713]" captionTargetId="figure-86@3.[151,1436,381,1728]" captionTargetPageId="3" captionText="FIGURES 3–7. Percussiopalpus inusitatus sp. n. Head in dorsal (3), lateral (4, 6), and ventral (5, 7) views. Abbreviations: ce, compound eye; fs, frontal sulcus; gal, galea; gp, gular plate; lr, labrum; md, mandible; mxp, maxillary palp; ptp, posterior tentorial pit; smb, submental bulge; t, tentorium." figureDoi="http://doi.org/10.5281/zenodo.7891906" httpUri="https://zenodo.org/record/7891906/files/figure.png" pageId="6" pageNumber="77">3</figureCitation>
</emphasis>
); tarsal claw only slightly shorter than tarsomere 3; protarsi shorter than half length of tibiae, meso- and metatarsi longer than half length of tibiae.
</paragraph>
<paragraph blockId="6.[151,1437,1208,1991]" pageId="6" pageNumber="77">
Aedeagus (
<figureCitation box="[328,461,1892,1918]" captionStart="FIGURES31–38" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[151,1436,885,1908]" captionTargetId="figure-284@10.[151,1436,885,1921]" captionTargetPageId="10" captionText="FIGURES31–38. Percussiopalpusinusitatussp. n. Protarsusin ventral(31) andlateral (32) views, metatarsus inlateral view (33), aedeagus in abparameral (34, 36), parameral (37) and lateral (35, 38) views. Abbreviations: pr, paramere; t1‒3, tarsomere 1‒3." figureDoi="http://doi.org/10.5281/zenodo.7891926" httpUri="https://zenodo.org/record/7891926/files/figure.png" pageId="6" pageNumber="77">Figs 34–38</figureCitation>
) elongate, with weakly sclerotized walls and asymmetric median lobe bearing oval dorsal diaphragm; endophallic sclerites elongate and asymmetric, parameres (
<figureCitation box="[1045,1130,1928,1954]" captionStart="FIGURES31–38" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[151,1436,885,1908]" captionTargetId="figure-284@10.[151,1436,885,1921]" captionTargetPageId="10" captionText="FIGURES31–38. Percussiopalpusinusitatussp. n. Protarsusin ventral(31) andlateral (32) views, metatarsus inlateral view (33), aedeagus in abparameral (34, 36), parameral (37) and lateral (35, 38) views. Abbreviations: pr, paramere; t1‒3, tarsomere 1‒3." figureDoi="http://doi.org/10.5281/zenodo.7891926" httpUri="https://zenodo.org/record/7891926/files/figure.png" pageId="6" pageNumber="77">Fig. 37</figureCitation>
,
<emphasis box="[1145,1170,1929,1954]" italics="true" pageId="6" pageNumber="77">pr</emphasis>
) asymmetric, long and broad, with subapical setae.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7891916" ID-Zenodo-Dep="7891916" httpUri="https://zenodo.org/record/7891916/files/figure.png" pageId="7" pageNumber="78" startId="7.[151,264,1457,1481]" targetBox="[151,1436,181,1433]" targetPageId="7" targetType="figure">
<paragraph blockId="7.[151,1436,1457,1589]" pageId="7" pageNumber="78">
<emphasis bold="true" box="[151,339,1457,1481]" pageId="7" pageNumber="78">FIGURES 19–23.</emphasis>
<taxonomicName authority="Jałoszyński &amp; Hlaváč &amp; Anlaş, 2023" authorityName="Jałoszyński &amp; Hlaváč &amp; Anlaş" authorityYear="2023" box="[343,614,1457,1481]" class="Insecta" family="Staphylinidae" genus="Percussiopalpus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="inusitatus" status="sp. n.">
<emphasis box="[343,614,1457,1481]" italics="true" pageId="7" pageNumber="78">Percussiopalpus inusitatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[618,674,1458,1482]" pageId="7" pageNumber="78">
<taxonomicNameLabel box="[618,674,1458,1482]" pageId="7" pageNumber="78" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Pterothorax in ventral (
<emphasis bold="true" box="[910,935,1457,1481]" pageId="7" pageNumber="78">19</emphasis>
,
<emphasis bold="true" box="[947,972,1457,1481]" pageId="7" pageNumber="78">20</emphasis>
) and left lateral (
<emphasis bold="true" box="[1142,1167,1457,1481]" pageId="7" pageNumber="78">22</emphasis>
,
<emphasis bold="true" box="[1179,1204,1457,1481]" pageId="7" pageNumber="78">23</emphasis>
) views, anterior region of mesoventrite in ventral view (
<emphasis bold="true" box="[481,506,1493,1517]" pageId="7" pageNumber="78">21</emphasis>
). Abbreviations: amtvp, anterior metaventral process; cx3, metacoxa; lfa, lateral furcal arm; mscxc, mesocoxal cavity; msff, mesofurcal fovea; msvp, mesoventral process; mtvp, metaventral process; pp, prepectus; v2‒3, meso- and metaventrite. Arrowheads indicate ventral foveae.
</paragraph>
</caption>
<paragraph blockId="7.[151,1436,1635,1914]" pageId="7" pageNumber="78">
<emphasis bold="true" box="[199,440,1635,1661]" pageId="7" pageNumber="78">Sexual dimorphism.</emphasis>
Males with thickened protarsomere 2, submedian ventral tooth on protibiae and sternite IX exposed and tripartite.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="78" type="etymology">
<paragraph blockId="7.[151,1436,1635,1914]" pageId="7" pageNumber="78">
<emphasis bold="true" box="[199,334,1708,1734]" pageId="7" pageNumber="78">Etymology.</emphasis>
The name
<taxonomicName authorityName="Jaloszynski &amp; Hlavac" authorityYear="2023" box="[465,649,1708,1734]" class="Insecta" family="Staphylinidae" genus="Percussiopalpus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis box="[465,649,1708,1734]" italics="true" pageId="7" pageNumber="78">Percussiopalpus</emphasis>
</taxonomicName>
refers to the peculiar maxillary palps with sensory processes shaped like percussion mallets, typical of
<taxonomicName authorityName="Jałoszyński &amp; Hlaváč &amp; Anlaş" authorityYear="2023" box="[535,766,1744,1770]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="71" phylum="Arthropoda" rank="tribe" tribe="Thaumastocephalini">Thaumastocephalini</taxonomicName>
. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection box="[199,948,1779,1806]" pageId="7" pageNumber="78" type="distribution">
<paragraph blockId="7.[151,1436,1635,1914]" box="[199,948,1779,1806]" pageId="7" pageNumber="78">
<emphasis bold="true" box="[199,353,1779,1805]" pageId="7" pageNumber="78">Distribution.</emphasis>
Northwestern
<collectingCountry box="[517,616,1780,1806]" name="Turkey" pageId="7" pageNumber="78">Anatolia</collectingCountry>
,
<collectingCountry box="[627,707,1780,1806]" name="Turkey" pageId="7" pageNumber="78">Turkey</collectingCountry>
(
<collectingRegion box="[723,936,1780,1806]" country="Turkey" name="Balikesir" pageId="7" pageNumber="78">Balıkesir Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,968,1815,1842]" pageId="7" pageNumber="78" type="taxon_list">
<paragraph blockId="7.[151,1436,1635,1914]" box="[199,968,1815,1842]" pageId="7" pageNumber="78">
<emphasis bold="true" box="[199,403,1815,1841]" pageId="7" pageNumber="78">Species included.</emphasis>
<taxonomicName authority="Jaloszynski &amp; Hlavac." authorityName="Jaloszynski &amp; Hlavac." box="[410,968,1816,1842]" class="Insecta" family="Staphylinidae" genus="Percussiopalpus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="inusitatus">
<emphasis box="[410,710,1816,1842]" italics="true" pageId="7" pageNumber="78">Percussiopalpus inusitatus</emphasis>
Jałoszyński &amp; Hlaváč.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="78" type="discussion">
<paragraph blockId="7.[151,1436,1635,1914]" pageId="7" pageNumber="78">
<emphasis bold="true" box="[199,315,1852,1878]" pageId="7" pageNumber="78">Remarks.</emphasis>
Characters given in the diagnosis all clearly differentiate this genus from
<taxonomicName box="[1133,1357,1852,1878]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis box="[1133,1357,1852,1878]" italics="true" pageId="7" pageNumber="78">Thaumastocephalus</emphasis>
</taxonomicName>
; major differences are also compiled in
<tableCitation box="[515,599,1888,1914]" captionStart="TABLE" captionStartId="8.[151,167,152,176]" captionTargetPageId="8" captionText="TABLE 1. Major morphological differences between genera of Thaumastocephalini." httpUri="http://table.plazi.org/id/DF7D93B1A103FFEA25C9FF2D934DFF73" pageId="7" pageNumber="78" tableUuid="DF7D93B1A103FFEA25C9FF2D934DFF73">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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