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<document id="C2D2ACAED20F8C0BA0B70E26923A09A2" ID-CLB-Dataset="2489" ID-DOI="10.21248/contrib.entomol.66.2.201-255" ID-GBIF-Dataset="991c89c8-c376-46a5-9aa8-6dfbbf623c23" ID-ISSN="0005-805X" ID-Zenodo-Dep="5212606" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629211447587" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="0391026EFF9DFF9FFF0B756FFB78F867" docLanguage="en" docName="ContribEntomol.66.2.201-255.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (2)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Anaulacaspis inexpectata GANGLBAUER 1895" docType="treatment" docVersion="11" lastPageNumber="253" masterDocId="FFA87A16FFA9FFABFFE8713CFFB4FFA6" masterDocTitle="Revision of the Anaulacaspis species of the Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="255" masterPageNumber="201" pageNumber="253" updateTime="1699087475910" updateUser="ExternalLinkService" zenodo-license-document="Abstyles">
<mods:mods id="D14A68B3CC96341BED9C650268AF508B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DD514CB08BCF4D523F881BB39A750800">
<mods:title id="556DB3C09930DECFD2B0B2ECF882156C">Revision of the Anaulacaspis species of the Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="41430B4EB42100500ACB99BAEFBBBB89" type="personal">
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<mods:namePart id="023766D851D030F0ABCF6A47D82FA594">Assing, Volker</mods:namePart>
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<mods:title id="1416DD1BE465724770C33D006657B477">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="29B7E1F2854E32663EB934F7D602A5A8">
<mods:date id="C4909420239E980B4C6DD4A8E9990358">2016</mods:date>
<mods:detail id="1C7C79A63D5140F9E06764244A876087" type="pubDate">
<mods:number id="BDE211D305DB9B81A5FCB93CD46C42EA">2016-12-20</mods:number>
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<mods:number id="4165B338A1221FC99A404C3CB9C32ADC">66</mods:number>
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<mods:url id="0A237149425CF4DF99C8474C70544E50">https://www.contributions-to-entomology.org/article/view/1904</mods:url>
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<mods:classification id="21CB3026637ABDF945DC48FF9CC44E12">journal article</mods:classification>
<mods:identifier id="E8EB780CD948821385A8844E85383960" type="CLB-Dataset">2489</mods:identifier>
<mods:identifier id="4EBE17A765262E2B290DA8C1B5838B7A" type="DOI">10.21248/contrib.entomol.66.2.201-255</mods:identifier>
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<treatment id="0391026EFF9DFF9FFF0B756FFB78F867" ID-DOI="http://doi.org/10.5281/zenodo.5884853" ID-GBIF-Taxon="192454149" ID-Zenodo-Dep="5884853" LSID="urn:lsid:plazi:treatment:0391026EFF9DFF9FFF0B756FFB78F867" httpUri="http://treatment.plazi.org/id/0391026EFF9DFF9FFF0B756FFB78F867" lastPageNumber="253" pageId="52" pageNumber="253">
<subSubSection id="C322E0F3FF9DFF9FFF0B756FFD24FBC8" box="[227,656,1107,1134]" pageId="52" pageNumber="253" type="nomenclature">
<paragraph id="8B87B378FF9DFF9FFF0B756FFD24FBC8" blockId="52.[227,656,1107,1167]" box="[227,656,1107,1134]" pageId="52" pageNumber="253">
<heading id="D0CF0414FF9DFF9FFF0B756FFD24FBC8" box="[227,656,1107,1134]" centered="true" fontSize="12" level="2" pageId="52" pageNumber="253" reason="2">
“
<taxonomicName id="4C38C8FBFF9DFF9FFF06756FFD24FBC8" ID-CoL="8VTS6" authority="(FAGEL, 1969)" authorityName="GANGLBAUER" authorityYear="1895" baseAuthorityName="FAGEL" baseAuthorityYear="1969" box="[238,656,1107,1134]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B94C6F6AFF9DFF9FFF06756FFEC3FBC8" box="[238,375,1107,1134]" italics="true" pageId="52" pageNumber="253">Anaulacaspis</emphasis>
”
<emphasis id="B94C6F6AFF9DFF9FFE60756FFDB4FBC8" box="[392,512,1107,1134]" italics="true" pageId="52" pageNumber="253">inexpectata</emphasis>
(
<bibRefCitation id="EFA9CE89FF9DFF9FFDE3756FFD3CFBC8" author="FAGEL, G." box="[523,648,1107,1134]" pageId="52" pageNumber="253" pagination="1 - 43" refId="ref48838" refString="FAGEL, G. 1969: Contribution a la connaissance des Staphylinidae. XIX. Les Melagria CASEY de la region palearctique occidentale. - Bulletin Institut Royal des Sciences Naturelles de Belgique 45 (34): 1 - 43." type="journal article" year="1969">FAGEL, 1969</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FF9DFF9FFF0B7545FE16FB29" box="[227,418,1145,1167]" pageId="52" pageNumber="253" type="description">
<paragraph id="8B87B378FF9DFF9FFF0B7545FE16FB29" blockId="52.[227,656,1107,1167]" box="[227,418,1145,1167]" pageId="52" pageNumber="253">
(
<figureCitation id="1303AFFDFF9DFF9FFF0F7545FEE9FB29" box="[231,349,1145,1167]" captionStart-0="Figs 211–232" captionStart-1="Figs 233–236" captionStartId-0="20.[189,229,1941,1963]" captionStartId-1="21.[151,191,725,747]" captionTargetBox-0="[201,1414,238,1920]" captionTargetBox-1="[160,1374,246,704]" captionTargetId-0="figure-41@20.[200,1426,232,1940]" captionTargetId-1="figure-716@21.[151,1398,232,724]" captionTargetPageId-1="21" captionText-0="Figs 211–232: Anaulacaspsis formosa (211), A. elegans (212–220), A. beesoni (221–228; 224–225: lectotype), and A. inexpectata (229–232; 231–232: holotype): habitus (211; photo by Marc Tronquet); male tergite VIII (212, 221); male sternite VIII (213, 222); median lobe of aedeagus in lateral and in ventral view (214–217, 224–227, 229–232); female tergite VIII (218); posterior margin of female tergite VIII (219); female sternite VIII (220, 228); postero-lateral portion of sternite VIII (223). Scale bars: 211: 1.0 mm; 212–218, 220–222, 224–232: 0.1 mm; 219, 223: 0.05 mm." captionText-1="Figs 233–236: Anaulacaspsis inexpectata: male tergite VIII (233); posterior margin of male tergite VIII (234); male sternite VIII (235); female sternite VIII (236). Scale bars: 233, 235–236: 0.1 mm; 234: 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5884779" figureDoi-1="http://doi.org/10.5281/zenodo.5212626" httpUri-0="https://zenodo.org/record/5884779/files/figure.png" httpUri-1="https://zenodo.org/record/5212626/files/figure.png" pageId="52" pageNumber="253">Figs 229–236</figureCitation>
,
<figureCitation id="1303AFFDFF9DFF9FFE8D7545FE29FB29" box="[357,413,1145,1167]" captionStart="Map 5" captionStartId="38.[189,234,2025,2046]" captionTargetBox="[189,1436,1328,2016]" captionTargetId="figure-506@38.[189,1436,1328,2016]" captionTargetPageId="38" captionText="Map 5: Distributions of Anaulacaspis flavomarginata (white diamonds), A. pamphylica (black star), A. inexpectata (white circles), and A. persica (black circles) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212639" httpUri="https://zenodo.org/record/5212639/files/figure.png" pageId="52" pageNumber="253">Map 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FF9DFF9FFF557597FD8DFB67" box="[189,569,1195,1217]" pageId="52" pageNumber="253" type="reference_group">
<paragraph id="8B87B378FF9DFF9FFF557597FD8DFB67" blockId="52.[189,569,1195,1217]" box="[189,569,1195,1217]" pageId="52" pageNumber="253">
<treatmentCitationGroup id="AB289456FF9DFF9FFF557597FD8DFB67" box="[189,569,1195,1217]" pageId="52" pageNumber="253">
<treatmentCitation id="0A999569FF9DFF9FFF557597FDAAFB66" author="FAGEL, G." box="[189,542,1195,1217]" page="41" pageId="52" pageNumber="253" year="1969">
<taxonomicName id="4C38C8FBFF9DFF9FFF557597FDAAFB66" ID-CoL="3Z3M4" authority="FAGEL, 1969: 41" authorityName="FAGEL" authorityPageNumber="41" authorityYear="1969" box="[189,542,1195,1217]" class="Insecta" family="Staphylinidae" genus="Melagria" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="inexpectata">
<emphasis id="B94C6F6AFF9DFF9FFF557597FE36FB67" box="[189,386,1195,1217]" italics="true" pageId="52" pageNumber="253">Melagria inexpectata</emphasis>
<bibRefCitation id="EFA9CE89FF9DFF9FFE6F7597FDAAFB66" author="FAGEL, G." box="[391,542,1195,1217]" pageId="52" pageNumber="253" pagination="1 - 43" refId="ref48838" refString="FAGEL, G. 1969: Contribution a la connaissance des Staphylinidae. XIX. Les Melagria CASEY de la region palearctique occidentale. - Bulletin Institut Royal des Sciences Naturelles de Belgique 45 (34): 1 - 43." type="journal article" year="1969">FAGEL, 1969: 41</bibRefCitation>
</taxonomicName>
</treatmentCitation>
ff.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FF9DFF9FFF5575D5FDD0FA27" pageId="52" pageNumber="253" type="materials_examined">
<paragraph id="8B87B378FF9DFF9FFF5575D5FDD0FA27" blockId="52.[189,794,1257,1410]" pageId="52" pageNumber="253">
<emphasis id="B94C6F6AFF9DFF9FFF5575D5FE6AFAA7" bold="true" box="[189,478,1257,1281]" pageId="52" pageNumber="253">Type material examined</emphasis>
:
<materialsCitation id="3B50B925FF9DFF9FFE1F75D5FD61FAC7" ID-GBIF-Occurrence="3449483726" collectionCode="IRSNB" collectorName="R. I. Sc." determinerName="A. Fauvel &amp; G. Fagel &amp; V. Assing" location="Coll" municipality="Tokmak" pageId="52" pageNumber="253" specimenCount="1" typeStatus="holotype">
<typeStatus id="54830DDAFF9DFF9FFE1F75D5FDEEFAA7" box="[503,602,1257,1281]" pageId="52" pageNumber="253" type="holotype">Holotype</typeStatus>
♂: “
<collectingMunicipality id="6BE32902FF9DFF9FFD4C75D5FD4AFAA7" box="[676,766,1257,1281]" pageId="52" pageNumber="253">Tokmak</collectingMunicipality>
/
<collectorName id="26CDD6AEFF9DFF9FFF557435FEB6FA87" box="[189,258,1289,1313]" pageId="52" pageNumber="253">R.I.Sc.</collectorName>
N.B. 17.479,
<location id="8EE7E5A3FF9DFF9FFE627435FE0EFA87" LSID="urn:lsid:plazi:treatment:0391026EFF9DFF9FFF0B756FFB78F867:8EE7E5A3FF9DFF9FFE627435FE0EFA87" box="[394,442,1289,1313]" municipality="Tokmak" name="Coll" pageId="52" pageNumber="253">Coll</location>
. et det.
<determinerName id="7E5B5FBCFF9DFF9FFDE47435FDC4FA87" box="[524,624,1289,1313]" pageId="52" pageNumber="253">A. Fauvel</determinerName>
/ /
<determinerName id="7E5B5FBCFF9DFF9FFD7D7435FD5FFA87" box="[661,747,1289,1313]" pageId="52" pageNumber="253">G. Fagel</determinerName>
det. 1969,
<taxonomicName id="4C38C8FBFF9DFF9FFEE87415FE6BFAE7" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[256,479,1321,1345]" class="Insecta" family="Staphylinidae" genus="Melagria" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="inexpectata" status="sp. nov.">
<emphasis id="B94C6F6AFF9DFF9FFEE87415FE6BFAE7" box="[256,479,1321,1345]" italics="true" pageId="52" pageNumber="253">
<location id="8EE7E5A3FF9DFF9FFEE87415FEEAFAE7" LSID="urn:lsid:plazi:treatment:0391026EFF9DFF9FFF0B756FFB78F867:8EE7E5A3FF9DFF9FFEE87415FEEAFAE7" box="[256,350,1321,1345]" municipality="Tokmak" name="Melagria" pageId="52" pageNumber="253">Melagria</location>
inexpectata
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27FD211FF9DFF9FFE027416FD96FAE7" box="[490,546,1322,1345]" pageId="52" pageNumber="253" rank="species">
n.
<specimenCount id="9D3E78F1FF9DFF9FFDEC7416FD96FAE7" box="[516,546,1322,1345]" pageId="52" pageNumber="253" type="generic">sp.</specimenCount>
</taxonomicNameLabel>
/
<typeStatus id="54830DDAFF9DFF9FFDD67416FDC6FAE4" box="[574,626,1322,1346]" pageId="52" pageNumber="253">Type</typeStatus>
/
<taxonomicName id="4C38C8FBFF9DFF9FFD787415FE3CFAC7" authority="(Fagel)" baseAuthorityName="Fagel" baseAuthorityYear="1969" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="inexpectata">
<emphasis id="B94C6F6AFF9DFF9FFD787415FE80FAC7" italics="true" pageId="52" pageNumber="253">
<location id="8EE7E5A3FF9DFF9FFD787415FCAEFAE7" LSID="urn:lsid:plazi:treatment:0391026EFF9DFF9FFF0B756FFB78F867:8EE7E5A3FF9DFF9FFD787415FCAEFAE7" box="[656,794,1321,1345]" municipality="Tokmak" name="Anaulacaspis" pageId="52" pageNumber="253">Anaulacaspis</location>
inexpectata
</emphasis>
(
<location id="8EE7E5A3FF9DFF9FFEAC7475FE35FAC7" LSID="urn:lsid:plazi:treatment:0391026EFF9DFF9FFF0B756FFB78F867:8EE7E5A3FF9DFF9FFEAC7475FE35FAC7" box="[324,385,1353,1377]" municipality="Tokmak" name="Fagel" pageId="52" pageNumber="253">Fagel</location>
)
</taxonomicName>
, det.
<determinerName id="7E5B5FBCFF9DFF9FFE2D7476FD93FAC7" box="[453,551,1353,1377]" pageId="52" pageNumber="253">V. Assing</determinerName>
2016” (
<collectionCode id="ED292BBDFF9DFF9FFD967475FD7AFAC7" box="[638,718,1353,1377]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="52" pageNumber="253">IRSNB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B50B925FF9DFF9FFD097476FDEBFA27" ID-GBIF-Occurrence="3449483518" collectionCode="IRSNB" collectorName="R. I. Sc." determinerName="A. Fauvel &amp; G. Fagel &amp; V. Assing" pageId="52" pageNumber="253" specimenCount="1" typeStatus="paratype">
<typeStatus id="54830DDAFF9DFF9FFD097476FF5FFA24" pageId="52" pageNumber="253" type="paratype">Paratype</typeStatus>
♀: same data as
<typeStatus id="54830DDAFF9DFF9FFE757455FE4FFA27" box="[413,507,1385,1409]" pageId="52" pageNumber="253" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED292BBDFF9DFF9FFDE07455FDECFA27" box="[520,600,1385,1409]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="52" pageNumber="253">IRSNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FF9DFF9FFF557495FE16F947" pageId="52" pageNumber="253" type="discussion">
<paragraph id="8B87B378FF9DFF9FFF557495FEB4F9A7" blockId="52.[189,794,1449,1537]" pageId="52" pageNumber="253">
<emphasis id="B94C6F6AFF9DFF9FFF557495FE98FA67" bold="true" box="[189,300,1449,1473]" pageId="52" pageNumber="253">Comment</emphasis>
: The original description is based on a male
<typeStatus id="54830DDAFF9DFF9FFF5574F5FEAFFA47" box="[189,283,1481,1505]" pageId="52" pageNumber="253" type="holotype">holotype</typeStatus>
and a female
<typeStatus id="54830DDAFF9DFF9FFE5F74F6FDA0FA44" box="[439,532,1482,1506]" pageId="52" pageNumber="253" type="paratype">paratype</typeStatus>
from “Tokmak” (
<bibRefCitation id="EFA9CE89FF9DFF9FFD3F74F5FF40F9A7" author="FAGEL, G." pageId="52" pageNumber="253" pagination="1 - 43" refId="ref48838" refString="FAGEL, G. 1969: Contribution a la connaissance des Staphylinidae. XIX. Les Melagria CASEY de la region palearctique occidentale. - Bulletin Institut Royal des Sciences Naturelles de Belgique 45 (34): 1 - 43." type="journal article" year="1969">FAGEL 1969</bibRefCitation>
).
</paragraph>
<paragraph id="8B87B378FF9DFF9FFF557715FE16F947" blockId="52.[189,794,1577,1761]" pageId="52" pageNumber="253">
<emphasis id="B94C6F6AFF9DFF9FFF557715FD97F9E7" bold="true" box="[189,547,1577,1601]" pageId="52" pageNumber="253">Additional material examined</emphasis>
:
<materialsCitation id="3B50B925FF9DFF9FFDD67715FE60F907" ID-GBIF-Occurrence="3449483618" collectingDate="2008-08-31" collectingDateMax="2008-09-08" collectingDateMin="2008-08-31" collectionCode="MNB" country="Iran" elevation="2100" latitude="36.633335" location="Salmas" longLatPrecision="1186" longitude="47.233334" pageId="52" pageNumber="253" specimenCount="1" stateProvince="Azarbayjan-e Gharbi">
<emphasis id="B94C6F6AFF9DFF9FFDD67715FDD9F9E7" bold="true" box="[574,621,1577,1601]" pageId="52" pageNumber="253">
<collectingCountry id="F32FF3E8FF9DFF9FFDD67715FDD9F9E7" box="[574,621,1577,1601]" name="Iran" pageId="52" pageNumber="253">Iran</collectingCountry>
</emphasis>
:
<emphasis id="B94C6F6AFF9DFF9FFD6F7715FEBFF9C7" bold="true" pageId="52" pageNumber="253">
<collectingRegion id="49FC7D9AFF9DFF9FFD6F7715FEBFF9C7" country="Iran" name="Azarbayjan-e Gharbi" pageId="52" pageNumber="253">Azarbayjan-e Gharbi</collectingRegion>
</emphasis>
: 2 ♂♂ [1 teneral], 1 ♀,
<quantity id="4CC01E9DFF9DFF9FFDE77775FDE6F9C7" box="[527,594,1609,1633]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="52" pageNumber="253" unit="km" value="20.0">
<locationDeviation id="9A35AA5AFF9DFF9FFDE77775FDE6F9C7" box="[527,594,1609,1633]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="52" pageNumber="253" unit="km" value="20.0">20 km</locationDeviation>
</quantity>
W
<location id="8EE7E5A3FF9DFF9FFD967775FD7FF9C7" LSID="urn:lsid:plazi:treatment:0391026EFF9DFF9FFF0B756FFB78F867:8EE7E5A3FF9DFF9FFD967775FD7FF9C7" box="[638,715,1609,1633]" country="Iran" latitude="36.633335" longLatPrecision="1186" longitude="47.233334" name="Salmas" pageId="52" pageNumber="253" stateProvince="Azarbayjan-e Gharbi">Salmas</location>
,
<locationDeviation id="9A35AA5AFF9DFF9FFD3F7775FED6F927" location="Kuzeh Rash" pageId="52" pageNumber="253">
<quantity id="4CC01E9DFF9DFF9FFD3F7775FCADF9C7" box="[727,793,1609,1633]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="52" pageNumber="253" unit="km" value="10.0">10 km</quantity>
W Kuzeh Rash
</locationDeviation>
,
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,
<geoCoordinate id="EE0CD5BFFF9DFF9FFE397755FD96F927" box="[465,546,1641,1665]" degrees="44" direction="east" minutes="31" orientation="longitude" pageId="52" pageNumber="253" precision="925" value="44.516666">44°31'E</geoCoordinate>
,
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<elevation id="0015544BFF9DFF9FFDC77755FD34F927" box="[559,640,1641,1665]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="52" pageNumber="253" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<date id="FF8695B8FF9DFF9FFD657755FCA1F927" box="[653,789,1641,1665]" pageId="52" pageNumber="253" value="2008-08-31">
<collectingDate id="EFC26C50FF9DFF9FFD657755FCA1F927" box="[653,789,1641,1665]" pageId="52" pageNumber="253" value="2008-08-31">31.VIII.2008</collectingDate>
</date>
, leg.
<collectorName id="26CDD6AEFF9DFF9FFF0F77B5FE9DF907" box="[231,297,1673,1697]" pageId="52" pageNumber="253">Frisch</collectorName>
&amp;
<collectorName id="26CDD6AEFF9DFF9FFEAF77B5FECFF907" box="[327,379,1673,1697]" pageId="52" pageNumber="253">Serri</collectorName>
(
<collectionCode id="ED292BBDFF9DFF9FFE6077B5FE7DF907" box="[392,457,1673,1697]" pageId="52" pageNumber="253">MNB</collectionCode>
);
</materialsCitation>
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1 ♂,
<collectorName id="26CDD6AEFF9DFF9FFDE277B6FDD2F907" box="[522,614,1673,1697]" pageId="52" pageNumber="253">N Takab</collectorName>
,
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<quantity id="4CC01E9DFF9DFF9FFD8777B5FD11F907" box="[623,677,1673,1697]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="52" pageNumber="253" unit="km" value="3.0">3 km</quantity>
NE Takhte-Soleyman
</locationDeviation>
,
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,
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,
<quantity id="4CC01E9DFF9DFF9FFDF17795FDDEF967" box="[537,618,1705,1729]" metricMagnitude="3" metricUnit="m" metricValue="2.27" pageId="52" pageNumber="253" unit="m" value="2270.0">
<elevation id="0015544BFF9DFF9FFDF17795FDDEF967" box="[537,618,1705,1729]" metricMagnitude="3" metricUnit="m" metricValue="2.27" pageId="52" pageNumber="253" unit="m" value="2270.0">2270 m</elevation>
</quantity>
,
<date id="FF8695B8FF9DFF9FFD937795FD50F967" box="[635,740,1705,1729]" pageId="52" pageNumber="253" value="2008-09-08">
<collectingDate id="EFC26C50FF9DFF9FFD937795FD50F967" box="[635,740,1705,1729]" pageId="52" pageNumber="253" value="2008-09-08">8.IX.2008</collectingDate>
</date>
, leg.
<collectorName id="26CDD6AEFF9DFF9FFF5577F5FF4BF947" box="[189,255,1737,1761]" pageId="52" pageNumber="253">Frisch</collectorName>
&amp;
<collectorName id="26CDD6AEFF9DFF9FFEF677F5FEE6F947" box="[286,338,1737,1761]" pageId="52" pageNumber="253">Serri</collectorName>
(cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FF9DFF9FFF557635FCD3F967" pageId="52" pageNumber="253" type="description">
<paragraph id="8B87B378FF9DFF9FFF557635FED2F867" blockId="52.[189,794,1801,2082]" pageId="52" pageNumber="253">
<emphasis id="B94C6F6AFF9DFF9FFF557635FEE2F887" bold="true" box="[189,342,1801,1825]" pageId="52" pageNumber="253">Redescription</emphasis>
: Body length 2.0–
<quantity id="4CC01E9DFF9DFF9FFDF67635FDC6F887" box="[542,626,1801,1825]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="52" pageNumber="253" unit="mm" value="2.3">2.3 mm</quantity>
; length of forebody
<quantity id="4CC01E9DFF9DFF9FFF177615FE2EF8E7" box="[255,410,1833,1857]" metricMagnitude="-4" metricUnit="m" metricValue="9.75" metricValueMax="10.5" metricValueMin="9.0" pageId="52" pageNumber="253" unit="mm" value="0.975" valueMax="1.05" valueMin="0.9">0.90–1.05 mm</quantity>
. Coloration: body black, with the postero-sutural portion of the elytra sometimes slightly paler; legs yellowish-brown to dark-brown; antennae blackish-brown to black. Whole body without microsculpture.
</paragraph>
<paragraph id="8B87B378FF9DFF9FFF5576F5FC25FE27" blockId="52.[189,794,1801,2082]" lastBlockId="52.[831,1436,233,1345]" pageId="52" pageNumber="253">
Head weakly transverse, approximately 1.1 times as broad as long, without appreciable sexual dimorphism; punctation fine and moderately dense. Eyes slightly longer than postocular region in dorsal view. Antenna short, approximately
<quantity id="4CC01E9DFF9DFF9FFC657035FC6BFE87" box="[909,991,265,289]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="52" pageNumber="253" unit="mm" value="0.7">0.7 mm</quantity>
long; antennomere IV weakly transverse; antennomeres V–X of gradually increasing width and increasingly transverse, X approximately twice as broad as long.
</paragraph>
<paragraph id="8B87B378FF9DFF9FFCD770B6FC56FDE7" blockId="52.[831,1436,233,1345]" pageId="52" pageNumber="253">Pronotum weakly transverse, 1.10–1.15 times as broad as long, and approximately as broad as, or slightly narrower than head, without appreciable sexual dimorphism; punctation dense and fine, denser and more distinct than that of head; midline with fine and shallow median sulcus.</paragraph>
<paragraph id="8B87B378FF9DFF9FFCD77375FC07FD67" blockId="52.[831,1436,233,1345]" pageId="52" pageNumber="253">Elytra approximately as long as pronotum; punctation moderately dense and fine, but distinct. Scutellum coarsely and rugosely sculptured. Hind wings fully developed.</paragraph>
<paragraph id="8B87B378FF9DFF9FFCD773F5FCC8FBA7" blockId="52.[831,1436,233,1345]" pageId="52" pageNumber="253">
Abdomen as broad as, or slightly narrower than elytra; anterior impressions of tergites III–V deep and with a transverse row of very coarse punctures separated by narrow ridges; punctation of remainder of tergal surfaces distinct and moderately dense; tergite VIII (
<figureCitation id="1303AFFDFF9DFF9FFCAF7255FC6FFC27" box="[839,987,873,897]" captionStart="Figs 233–236" captionStartId="21.[151,191,725,747]" captionTargetBox="[160,1374,246,704]" captionTargetId="figure-716@21.[151,1398,232,724]" captionTargetPageId="21" captionText="Figs 233–236: Anaulacaspsis inexpectata: male tergite VIII (233); posterior margin of male tergite VIII (234); male sternite VIII (235); female sternite VIII (236). Scale bars: 233, 235–236: 0.1 mm; 234: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5212626" httpUri="https://zenodo.org/record/5212626/files/figure.png" pageId="52" pageNumber="253">Figs 233–234</figureCitation>
) without a cluster of long postero-lateral setae, anterior margin without the usual median excision, posterior margin convex and with conspicuous dense and strongly modified, short and stout marginal setae.
</paragraph>
<paragraph id="8B87B378FF9DFF9FFCD77536FC20FB47" blockId="52.[831,1436,233,1345]" pageId="52" pageNumber="253">
♂: head indistinctly depressed in postero-median portion; sternite VIII (
<figureCitation id="1303AFFDFF9DFF9FFBB07515FB00FBE7" box="[1112,1204,1065,1089]" captionStart="Figs 233–236" captionStartId="21.[151,191,725,747]" captionTargetBox="[160,1374,246,704]" captionTargetId="figure-716@21.[151,1398,232,724]" captionTargetPageId="21" captionText="Figs 233–236: Anaulacaspsis inexpectata: male tergite VIII (233); posterior margin of male tergite VIII (234); male sternite VIII (235); female sternite VIII (236). Scale bars: 233, 235–236: 0.1 mm; 234: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5212626" httpUri="https://zenodo.org/record/5212626/files/figure.png" pageId="52" pageNumber="253">Fig. 235</figureCitation>
) smoothly convex posteriorly; median lobe of aedeagus (
<figureCitation id="1303AFFDFF9DFF9FFB167575FA20FBC7" box="[1278,1428,1097,1121]" captionStart="Figs 211–232" captionStartId="20.[189,229,1941,1963]" captionTargetBox="[201,1414,238,1920]" captionTargetId="figure-41@20.[200,1426,232,1940]" captionText="Figs 211–232: Anaulacaspsis formosa (211), A. elegans (212–220), A. beesoni (221–228; 224–225: lectotype), and A. inexpectata (229–232; 231–232: holotype): habitus (211; photo by Marc Tronquet); male tergite VIII (212, 221); male sternite VIII (213, 222); median lobe of aedeagus in lateral and in ventral view (214–217, 224–227, 229–232); female tergite VIII (218); posterior margin of female tergite VIII (219); female sternite VIII (220, 228); postero-lateral portion of sternite VIII (223). Scale bars: 211: 1.0 mm; 212–218, 220–222, 224–232: 0.1 mm; 219, 223: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884779" httpUri="https://zenodo.org/record/5884779/files/figure.png" pageId="52" pageNumber="253">Figs 229–232</figureCitation>
) approximately
<quantity id="4CC01E9DFF9DFF9FFC1C7555FBE1FB27" box="[1012,1109,1129,1153]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="52" pageNumber="253" unit="mm" value="0.25">0.25 mm</quantity>
long, ventral process in lateral view straight and without median membranous projection at base, in ventral view narrow and apically convex.
</paragraph>
<paragraph id="8B87B378FF9DFF9FFCD775D6FB16FAE7" blockId="52.[831,1436,233,1345]" pageId="52" pageNumber="253">
♀: head not depressed in postero-median portion; posterior margin of sternite VIII (
<figureCitation id="1303AFFDFF9DFF9FFB6C7435FB68FA87" box="[1156,1244,1289,1313]" captionStart="Figs 233–236" captionStartId="21.[151,191,725,747]" captionTargetBox="[160,1374,246,704]" captionTargetId="figure-716@21.[151,1398,232,724]" captionTargetPageId="21" captionText="Figs 233–236: Anaulacaspsis inexpectata: male tergite VIII (233); posterior margin of male tergite VIII (234); male sternite VIII (235); female sternite VIII (236). Scale bars: 233, 235–236: 0.1 mm; 234: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5212626" httpUri="https://zenodo.org/record/5212626/files/figure.png" pageId="52" pageNumber="253">Fig. 236</figureCitation>
) broadly convex; spermatheca not comma-shaped.
</paragraph>
<paragraph id="8B87B378FF9DFF9FFCD77455FCD3F967" blockId="52.[831,1436,1385,1729]" pageId="52" pageNumber="253">
<emphasis id="B94C6F6AFF9DFF9FFCD77455FBA6FA27" bold="true" box="[831,1042,1385,1409]" pageId="52" pageNumber="253">Comparative notes</emphasis>
: This species is readily distinguished from all its West Palaearctic congeners by several unique character conditions: the coarsely and rugosely sculptured scutellum, the coarse punctation of the anterior impressions of tergites III–V, the conspicuous chaetotaxy of the abdominal tergite VIII, the absence of a median excision of the anterior margin of tergite VIII, the morphology of the median lobe of the aedeagus, and the different shape of the spermatheca. For additional comments see the section on species groups in chapter 3.1.
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FF9DFF9FFCD777D5FB78F867" pageId="52" pageNumber="253" type="distribution">
<paragraph id="8B87B378FF9DFF9FFCD777D5FB78F867" blockId="52.[831,1436,1769,1985]" pageId="52" pageNumber="253">
<emphasis id="B94C6F6AFF9DFF9FFCD777D5FC7FF8A7" bold="true" box="[831,971,1769,1793]" pageId="52" pageNumber="253">Distribution</emphasis>
: The
<typeStatus id="54830DDAFF9DFF9FFBFC77D6FBF7F8A4" box="[1044,1091,1770,1794]" pageId="52" pageNumber="253">type</typeStatus>
locality “Tokmak” is ambiguous; there are several geographic entities with that name in
<collectingCountry id="F32FF3E8FF9DFF9FFCB67615FC0DF8E7" box="[862,953,1833,1857]" name="Ukraine" pageId="52" pageNumber="253">Ukraine</collectingCountry>
,
<collectingCountry id="F32FF3E8FF9DFF9FFC2E7615FBBAF8E7" box="[966,1038,1833,1857]" name="Russia" pageId="52" pageNumber="253">Russia</collectingCountry>
, and Middle Asia. According to the Palaearctic Catalogue (
<bibRefCitation id="EFA9CE89FF9DFF9FFBA47676FA24F8C7" author="SCHULKE, M. &amp; SMETANA, A." box="[1100,1424,1865,1893]" pageId="52" pageNumber="253" pagination="304 - 1134" refId="ref49967" refString="SCHULKE, M. &amp; SMETANA, A. 2015: Staphylinidae, pp. 304 - 1134. - In: LOBL, I. &amp; LOBL, D. (eds), Catalogue of Palaearctic Coleoptera. New, updated Edition. Volume 2. Hydrophiloidea - Staphylinoidea. Revised and updated edition. - Leiden: Brill: xxvi + 1702 pp." type="book chapter" year="2015">SCHÜLKE &amp; SMETANA 2015</bibRefCitation>
), Tokmak is in
<collectingCountry id="F32FF3E8FF9DFF9FFC337656FBE0F824" box="[987,1108,1898,1922]" name="Kyrgyzstan" pageId="52" pageNumber="253">Kyrgyzstan</collectingCountry>
and thus identical to what is Tokmok today. The additional material was collected in two localities in North
<collectingCountry id="F32FF3E8FF9DFF9FFBD37696FBDCF867" box="[1083,1128,1962,1985]" name="Iran" pageId="52" pageNumber="253">Iran</collectingCountry>
(
<figureCitation id="1303AFFDFF9DFF9FFB9E7695FB0BF867" box="[1142,1215,1961,1985]" captionStart="Map 5" captionStartId="38.[189,234,2025,2046]" captionTargetBox="[189,1436,1328,2016]" captionTargetId="figure-506@38.[189,1436,1328,2016]" captionTargetPageId="38" captionText="Map 5: Distributions of Anaulacaspis flavomarginata (white diamonds), A. pamphylica (black star), A. inexpectata (white circles), and A. persica (black circles) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212639" httpUri="https://zenodo.org/record/5212639/files/figure.png" pageId="52" pageNumber="253">Map 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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