92 lines
9.3 KiB
XML
92 lines
9.3 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6210769" ID-GBIF-Dataset="7485373c-0219-4ed7-bb5c-ac664acf6775" ID-GBIF-Taxon="114923677" ID-Zenodo-Dep="6210769" checkinTime="1427905504904" checkinUser="donat" docAuthor="Alberto Ballerio & Munetoshi Maruyama" docDate="2010" docId="038951095C0F3D1466B6FA90AAA1FDF7" docLanguage="en" docName="ZK_article_2140.pdf" docOrigin="Zookeys 34" docTitle="Madrasostes clypeale Paulian 1993" docType="treatment" docVersion="8" lastPageId="8" lastPageNumber="84" masterDocId="FFB029715C083D1C663AFFC4A802FFD6" masterDocTitle="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)" masterLastPageNumber="104" masterPageNumber="77" pageId="7" pageNumber="84" updateTime="1645477292090" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Alberto Ballerio</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Munetoshi Maruyama</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zookeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="volume">
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<mods:number>34</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>77</mods:start>
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<mods:end>104</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.34.268</mods:identifier>
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<mods:identifier type="ZooBank">553C91CE-FEA1-4867-929B-2865647E4B2E</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6210769" ID-GBIF-Taxon="114923677" ID-Zenodo-Dep="6210769" LSID="urn:lsid:plazi:treatment:038951095C0F3D1466B6FA90AAA1FDF7" httpUri="http://treatment.plazi.org/id/038951095C0F3D1466B6FA90AAA1FDF7" lastPageId="8" lastPageNumber="84" pageId="7" pageNumber="84">
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<subSubSection box="[140,810,1363,1390]" pageId="7" pageNumber="84" type="nomenclature">
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<paragraph box="[140,810,1363,1390]" pageId="7" pageNumber="84">
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<emphasis bold="true" box="[140,559,1363,1390]" pageId="7" pageNumber="84">
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<taxonomicName box="[140,559,1363,1390]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="clypeale">
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<emphasis bold="true" box="[140,385,1364,1390]" italics="true" pageId="7" pageNumber="84">Madrasostes clypeale</emphasis>
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Paulian, 1993
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</taxonomicName>
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</emphasis>
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(
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<figureCitation box="[574,641,1363,1390]" captionStart="Figure 2" captionStartId="15.[140,211,1558,1580]" captionTargetBox="[140,1107,172,1515]" captionTargetId="figure@15.[140,1108,165,1521]" captionTargetPageId="15" captionText="Figure 2. Ceratocanthinae of Ulu Gombak. A Madrasostes depressum habitus dorsal B Madrasostes boucomonti habitus dorsal C Madrasostes agostii sp. n. habitus dorsal D Madrasostes clypeale male habitus dorsal E Madrasostes malyanum habitus dorsal F Madrasostes mirificum sp. n. habitus dorsal G habitus lateral of male H fore body of female I male protibia dorsal view J male protibia lateral view." pageId="7" pageNumber="84">Fig. 2</figureCitation>
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D,
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<figureCitation box="[678,743,1363,1390]" captionStart="Figure 6" captionStartId="21.[140,211,1206,1228]" captionTargetBox="[140,1107,165,1186]" captionTargetId="figure@21.[140,1108,165,1187]" captionTargetPageId="21" captionText="Figure 6. Photos of alive specimens taken in habitat at Ulu Gombak (photos © M. Maruyama, 2008). A Madrasostes variolosum B Madrasostes agostii C Madrasostes clypeale male D idem female E Eusphaeropeltis sp. b F Eusphaeropeltis sp. c." pageId="7" pageNumber="84">Fig. 6</figureCitation>
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C, D)
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</paragraph>
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</subSubSection>
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<subSubSection box="[185,839,1398,1425]" pageId="7" pageNumber="84" type="description">
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<paragraph box="[185,839,1398,1425]" pageId="7" pageNumber="84">Known distribution: Indonesia (Sumatra) (Paulian 1993).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="85" pageId="7" pageNumber="84" type="materials_examined">
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<paragraph lastPageId="8" lastPageNumber="85" pageId="7" pageNumber="84">
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New material examined: West Malaysia, Perak, 25 km NE of Ipoh, Banjaran Titi Wangsa Mounts, Gunung Korbu, 1400–1800 mt., 11–31.I.1999, leg. P. Cechovsky (ABCB); West Malaysia, Pearak, Penang Island, above botanical garden, 250 mt., 12.XI.1999, #4a, leg. Cuccodoro & Loebl, sifting forest leaf litter (MHNG, ABCB). Remarks. First record for West Malaysia (Perak and Selangor). The amazing sexual dimorphism of this species has already been described (Ballerio 2006). An examination of the microsculpturing of pronotum through scanning electron microscopy revealed that the pronotal punctures bottom surface is areolate (
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<figureCitation box="[849,914,166,193]" captionStart="Figure 5" captionStartId="20.[140,211,1093,1115]" captionTargetBox="[140,1107,165,1070]" captionTargetId="figure@20.[140,1108,165,1073]" captionText="Figure 5. SEM photographs of: A Madrasostes clypeale: male pronotum B detail of male pronotum puncture C female pronotum D detail of female pronotum punctures E and F Madrasostes mirificum sp. n. mouthparts: labium G epipharynx H mandibles I detail of galear brush J maxilla." pageId="8" pageNumber="85">Fig. 5</figureCitation>
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A, B). The larger punctures of females have the areolate surface limited to a portion of the bottom and slightly raised (
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<figureCitation box="[310,376,237,263]" captionStart="Figure 5" captionStartId="20.[140,211,1093,1115]" captionTargetBox="[140,1107,165,1070]" captionTargetId="figure@20.[140,1108,165,1073]" captionText="Figure 5. SEM photographs of: A Madrasostes clypeale: male pronotum B detail of male pronotum puncture C female pronotum D detail of female pronotum punctures E and F Madrasostes mirificum sp. n. mouthparts: labium G epipharynx H mandibles I detail of galear brush J maxilla." pageId="8" pageNumber="85">Fig. 5</figureCitation>
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C, D, E). Each puncture is often preceded by a short fine seta. It was not possible to detect any pore inside the punctures, so that the dirt which usually fills the punctures comes probably from outside. The species closest to
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<emphasis box="[985,1107,307,334]" italics="true" pageId="8" pageNumber="85">M. clypeale</emphasis>
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is
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<taxonomicName authority="Paulian, 1987" authorityName="Paulian" authorityYear="1987" box="[167,521,342,369]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="thai">
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<emphasis box="[167,349,342,369]" italics="true" pageId="8" pageNumber="85">Madrasostes thai</emphasis>
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Paulian, 1987
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</taxonomicName>
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, known only by the male holotype from Thailand (Chanthaburi province, well north of the isthmus of Kra, which is the boundary between the Sundaland and Indochinese subregions, see Corlett 2009). Both
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<emphasis box="[986,1107,413,439]" italics="true" pageId="8" pageNumber="85">M. clypeale</emphasis>
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and
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<emphasis box="[190,274,448,475]" italics="true" pageId="8" pageNumber="85">
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<taxonomicName box="[190,274,448,475]" class="Insecta" family="Hybosoridae" genus="Madrasostes" genus.bestMatchDistance="47" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="thai">M. thai</taxonomicName>
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</emphasis>
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represent a distinctive group within
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<emphasis box="[691,819,448,475]" italics="true" pageId="8" pageNumber="85">
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<taxonomicName box="[691,819,448,475]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
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</emphasis>
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and their current generic placement in the genus
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<taxonomicName box="[411,539,483,510]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="genus">
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<emphasis box="[411,539,483,510]" italics="true" pageId="8" pageNumber="85">Madrasostes</emphasis>
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</taxonomicName>
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must be regarded as provisional in the framework of a revision of the
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<taxonomicName box="[358,505,519,545]" class="Insecta" family="Hybosoridae" genus="Perignamptus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="genus">
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<emphasis box="[358,505,519,545]" italics="true" pageId="8" pageNumber="85">Perignamptus</emphasis>
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</taxonomicName>
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generic group.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |