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<document id="41317C7A82FCE80A03AE068973AFA7C9" ID-DOI="10.11865/zs.2024101" ID-ISSN="2095-6827" ID-Zenodo-Dep="10941018" ID-ZooBank="CA760FC0-4A28-4FEC-8D5F-766024ACF490" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712576729003" checkinUser="felipe" docAuthor="Lin, Yejie, Li, Shuqiang, Mo, Haolin &amp; Wang, Xihao" docDate="2024" docId="81114138FF9EFFFE7D9FFE6EC099FE4C" docLanguage="en" docName="ZoolSyst.49.1.4-98.pdf" docOrigin="Zoological Systematics 49 (1)" docSource="https://www.mendeley.com/catalogue/be4c04f9-10f9-351c-8fd7-9c0289bee230/" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Raveniola shixiu Lin, Wang &amp; Li 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="32" masterDocId="7D283940FF84FFE27D17FFD2C25AFFC3" masterDocTitle="Thirty-eight spider species (Arachnida: Araneae) from China, Indonesia, Japan and Vietnam" masterLastPageNumber="98" masterPageNumber="4" pageNumber="30" updateTime="1712577654997" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1805922008CCED2BEFB24C48B407F14D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DEEE31ED9B47AF05A37FF624A37D1AEC">
<mods:title id="1D102742DAC2C067BE4F1549BDE4DA3F">Thirty-eight spider species (Arachnida: Araneae) from China, Indonesia, Japan and Vietnam</mods:title>
</mods:titleInfo>
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<mods:namePart id="BFE3952C09EB733D44438DCA187FEC8A">Lin, Yejie</mods:namePart>
<mods:affiliation id="541C9CD1EB453C226DFCDA2458D06FB1">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
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<mods:namePart id="F47221634DC5CE0ECCDA3168AE1211B8">Li, Shuqiang</mods:namePart>
<mods:affiliation id="BA12FE7D8337FA462821ACD55F6AC4B9">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name id="1BB3DBA5D2D54135D1B3A8636E57897A" type="personal">
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<mods:roleTerm id="437863AB17A80EE02B00BC196C08837F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7AF7DC2134680875D489C7D44A6E109D">Mo, Haolin</mods:namePart>
<mods:affiliation id="ADE7D3DC6CB9B8F34D070C4F3251EB81">Key Laboratory of Wildlife Evolution and Conservation in Mountain Ecosystem of Guangxi, Nanning Normal University, Nanning 530001, China</mods:affiliation>
</mods:name>
<mods:name id="1F29D0F4B096756EB3BEBD1D2E31EBE8" type="personal">
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</mods:role>
<mods:namePart id="18D33C6C68A5B8948F263E7AED6BACB2">Wang, Xihao</mods:namePart>
<mods:affiliation id="E08B0970E6AF757FDEAD7270E6800209">Jinan Licheng No. 2 High School, Jinan 250109, China Corresponding author, E-mail: lisq @ ioz. ac. cn</mods:affiliation>
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<mods:title id="458A3C3FC93E0D01D9B922B248782B20">Zoological Systematics</mods:title>
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<mods:part id="AE396E82881D9352D18E5D926A24069B">
<mods:date id="9865EA81265CB7E672F9554110B7D6C1">2024</mods:date>
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<mods:number id="B4F595835DA4E1ABF48279092A10CACF">49</mods:number>
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<mods:start id="426C8CAA0F65EEFC60699C8EF53D02C6">4</mods:start>
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<mods:url id="0BE3250DAEFD054862AED1272696F310">https://www.mendeley.com/catalogue/be4c04f9-10f9-351c-8fd7-9c0289bee230/</mods:url>
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<mods:identifier id="77516CFDD5096BD5B4CDECAD69CB9C4A" type="DOI">10.11865/zs.2024101</mods:identifier>
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<treatment id="81114138FF9EFFFE7D9FFE6EC099FE4C" ID-DOI="http://doi.org/10.5281/zenodo.10941172" ID-Zenodo-Dep="10941172" LSID="urn:lsid:plazi:treatment:81114138FF9EFFFE7D9FFE6EC099FE4C" httpUri="http://treatment.plazi.org/id/81114138FF9EFFFE7D9FFE6EC099FE4C" lastPageId="28" lastPageNumber="32" pageId="26" pageNumber="30">
<subSubSection id="41A2A3A5FF9EFFF87D9FFE6EC148FE16" box="[136,786,444,469]" pageId="26" pageNumber="30" type="nomenclature">
<paragraph id="0907F02EFF9EFFF87D9FFE6EC148FE16" blockId="26.[136,786,444,469]" box="[136,786,444,469]" pageId="26" pageNumber="30">
<heading id="524F4742FF9EFFF87D9FFE6EC148FE16" box="[136,786,444,469]" centered="true" fontSize="10" level="2" pageId="26" pageNumber="30" reason="2">
<emphasis id="3BCC2C3CFF9EFFF87D9FFE6EC03AFE16" bold="true" box="[136,608,444,469]" pageId="26" pageNumber="30">
<taxonomicName id="CEB88BADFF9EFFF87D9FFE6EC3A5FE17" authority="Lin, Wang &amp; Li" authorityName="Lin, Wang &amp; Li" authorityYear="2024" box="[136,511,444,469]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="30" phylum="Arthropoda" rank="species" species="shixiu" status="sp. nov.">
<emphasis id="3BCC2C3CFF9EFFF87D9FFE6EC318FE17" bold="true" box="[136,322,444,468]" italics="true" pageId="26" pageNumber="30">Raveniola shixiu</emphasis>
Lin, Wang &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="20FF9147FF9EFFF87F1CFE6FC03AFE16" box="[523,608,445,469]" pageId="26" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="9183ECABFF9EFFF87F6AFE6EC08AFE16" box="[637,720,444,469]" captionStart="Figure 21" captionStartId="26.[155,218,2024,2046]" captionTargetBox="[189,1399,542,1988]" captionTargetId="figure-63@26.[189,1399,542,1989]" captionTargetPageId="26" captionText="Figure 21. Raveniola shixiu Lin, Wang &amp; Li, sp. nov., holotype female. A. Vulva, ventral view; B. Same, dorsal view. Abbreviation: S—spermatheca. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10941060" httpUri="https://zenodo.org/record/10941060/files/figure.png" pageId="26" pageNumber="30">Figs 21</figureCitation>
,
<figureCitation id="9183ECABFF9EFFF87FCCFE6FC151FE16" box="[731,779,445,469]" captionStart="Figure 24" captionStartId="29.[155,218,1701,1723]" captionTargetBox="[189,1397,898,1652]" captionTargetId="figure-366@29.[189,1399,898,1652]" captionTargetPageId="29" captionText="Figure 24. Raveniola spp., holotype female (A) and male (B, D) and paratype female (C), habitus. A. R. shixiu Lin, Wang &amp; Li, sp. nov., dorsal view; B. R. xiezhen Lin, Wang &amp; Li, sp. nov., dorsal view; C. Same, dorsal view; D. Same, leg I, lateral view. Scale bars: A–B=2.0 mm; C–D=3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10941066" httpUri="https://zenodo.org/record/10941066/files/figure.png" pageId="26" pageNumber="30">24A</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<caption id="5DC7A0A6FF9EFFF87D8CF83AC05DF7DD" ID-DOI="http://doi.org/10.5281/zenodo.10941060" ID-Zenodo-Dep="10941060" httpUri="https://zenodo.org/record/10941060/files/figure.png" pageId="26" pageNumber="30" startId="26.[155,218,2024,2046]" targetBox="[189,1399,542,1988]" targetPageId="26" targetType="figure">
<paragraph id="0907F02EFF9EFFF87D8CF83AC05DF7DD" blockId="26.[155,1434,2024,2078]" pageId="26" pageNumber="30">
Figure 21.
<taxonomicName id="CEB88BADFF9EFFF87C12F83AC013F83D" authority="Lin, Wang &amp; Li" authorityName="Lin, Wang &amp; Li" authorityYear="2024" box="[261,585,2024,2046]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="30" phylum="Arthropoda" rank="species" species="shixiu" status="sp. nov.">
<emphasis id="3BCC2C3CFF9EFFF87C12F83AC3FCF83E" box="[261,422,2024,2045]" italics="true" pageId="26" pageNumber="30">Raveniola shixiu</emphasis>
Lin, Wang &amp; Li
</taxonomicName>
,
<emphasis id="3BCC2C3CFF9EFFF87F44F83BC0FAF83D" bold="true" box="[595,672,2025,2046]" pageId="26" pageNumber="30">
<taxonomicNameLabel id="20FF9147FF9EFFF87F44F83BC0FAF83D" box="[595,672,2025,2046]" pageId="26" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female. A. Vulva, ventral view; B. Same, dorsal view. Abbreviation: S—spermatheca. Scale bars =0.1 mm.
</paragraph>
</caption>
<subSubSection id="41A2A3A5FF9FFFFE7DAAFF3EC099FE4C" lastPageId="28" lastPageNumber="32" pageId="27" pageNumber="31" type="etymology">
<paragraph id="0907F02EFF9FFFF97DAAFF3EC2A5FEE4" blockId="27.[136,1452,236,676]" pageId="27" pageNumber="31">
Etymology. The species is named after Shi Xiu, a character in the classic Chinese novel
<emphasis id="3BCC2C3CFF9FFFF97971FF3EC714FEC7" box="[1126,1358,236,260]" italics="true" pageId="27" pageNumber="31">Outlaws of the Marsh</emphasis>
; noun in apposition.
</paragraph>
<paragraph id="0907F02EFF9FFFF97DAAFEE0C6E1FE4C" blockId="27.[136,1452,236,676]" pageId="27" pageNumber="31">
Diagnosis. Females of the new species are similar to
<taxonomicName id="CEB88BADFF9FFFF97E17FEE0C67FFE89" authority="Lin &amp; Li, 2022" authorityName="Lin &amp; Li" authorityYear="2022" box="[768,1061,306,330]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="31" phylum="Arthropoda" rank="species" species="yangren">
<emphasis id="3BCC2C3CFF9FFFF97E17FEE0C12CFE8A" box="[768,886,306,329]" italics="true" pageId="27" pageNumber="31">R. yangren</emphasis>
Lin &amp; Li, 2022
</taxonomicName>
and
<taxonomicName id="CEB88BADFF9FFFF9794CFEE0C7D3FE89" authority="Yu &amp; Zhang, 2021" authorityName="Yu &amp; Zhang" authorityYear="2021" box="[1115,1417,306,330]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="31" phylum="Arthropoda" rank="species" species="lamia">
<emphasis id="3BCC2C3CFF9FFFF9794CFEE0C6EDFE89" box="[1115,1207,306,330]" italics="true" pageId="27" pageNumber="31">R. lamia</emphasis>
Yu &amp; Zhang, 2021
</taxonomicName>
by having only 1 pair of spermathecae. However, the new species can be distinguished by the straight spermathecae (
<emphasis id="3BCC2C3CFF9FFFF97857FE87C707FEAF" box="[1344,1373,341,364]" italics="true" pageId="27" pageNumber="31">vs.</emphasis>
curved in
<taxonomicName id="CEB88BADFF9FFFF97DB3FEAAC2A4FE4C" authorityName="Yu &amp; Zhang" authorityYear="2021" box="[164,254,375,399]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="31" phylum="Arthropoda" rank="species" species="lamia">
<emphasis id="3BCC2C3CFF9FFFF97DB3FEAAC2A4FE4C" box="[164,254,375,399]" italics="true" pageId="27" pageNumber="31">R. lamia</emphasis>
</taxonomicName>
) and spermathecae 2 times longer than wide (
<emphasis id="3BCC2C3CFF9FFFF97FFDFEAAC15DFE4C" box="[746,775,376,399]" italics="true" pageId="27" pageNumber="31">vs.</emphasis>
4 times longer than wide in
<taxonomicName id="CEB88BADFF9FFFF97920FEAAC6F1FE4C" authorityName="Lin &amp; Li" authorityYear="2022" box="[1079,1195,376,399]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="31" phylum="Arthropoda" rank="species" species="yangren">
<emphasis id="3BCC2C3CFF9FFFF97920FEAAC6F1FE4C" box="[1079,1195,376,399]" italics="true" pageId="27" pageNumber="31">R. yangren</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="0907F02EFF9FFFF97DAAFE48C7F6FDDA" blockId="27.[136,1452,236,676]" pageId="27" pageNumber="31">
Description. Female (
<typeStatus id="D6034E8CFF9FFFF97CB4FE48C05CFE71" box="[419,518,410,434]" pageId="27" pageNumber="31" type="holotype">holotype</typeStatus>
). Total length 17.63, carapace 6.79 long, 5.77 wide, opisthosoma 10.77 long, 5.58 wide. Eye sizes and interdistances: AME 0.11, ALE 0.29, PME 0.13, PLE 0.15, AME–AME 0.13, AME–ALE 0.08, PME–PME 0.40, PME–PLE 0.04, AME–PME 0.14, ALE–PLE 0.04. Chelicerae with 9 retromarginal teeth. Leg measurements: I 19.25 (4.94, 7.62, 3.59, 3.10), II 18.47 (4.68, 6.99, 3.75, 3.05), III 18.40 (4.49, 6.15, 4.68, 3.08), IV 24.04 (5.26, 8.21, 6.98, 3.59).
</paragraph>
<paragraph id="0907F02EFF9FFFF97DAAFDF6C3D9FD41" blockId="27.[136,1452,236,676]" pageId="27" pageNumber="31">
Coloration (
<figureCitation id="9183ECABFF9FFFF97C29FDF6C3FBFDFF" box="[318,417,548,572]" captionStart="Figure 24" captionStartId="29.[155,218,1701,1723]" captionTargetBox="[189,1397,898,1652]" captionTargetId="figure-366@29.[189,1399,898,1652]" captionTargetPageId="29" captionText="Figure 24. Raveniola spp., holotype female (A) and male (B, D) and paratype female (C), habitus. A. R. shixiu Lin, Wang &amp; Li, sp. nov., dorsal view; B. R. xiezhen Lin, Wang &amp; Li, sp. nov., dorsal view; C. Same, dorsal view; D. Same, leg I, lateral view. Scale bars: A–B=2.0 mm; C–D=3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10941066" httpUri="https://zenodo.org/record/10941066/files/figure.png" pageId="27" pageNumber="31">Fig. 24A</figureCitation>
). Chelicerae red-brown. Carapace, endites, labium and sternum yellowish brown. Legs yellowish brown with black setae. Opisthosoma oval, brown, without any pattern. Heart mark pale yellow, sword shaped. Four spinnerets, pale yellow.
</paragraph>
<paragraph id="0907F02EFF9FFFFE7DAAFD5EC6B9FEC7" blockId="27.[136,1452,236,676]" lastBlockId="28.[136,1452,236,399]" lastPageId="28" lastPageNumber="32" pageId="27" pageNumber="31">
Epigyne (
<figureCitation id="9183ECABFF9FFFF97C30FD5EC322FD67" box="[295,376,652,676]" captionStart="Figure 21" captionStartId="26.[155,218,2024,2046]" captionTargetBox="[189,1399,542,1988]" captionTargetId="figure-63@26.[189,1399,542,1989]" captionTargetPageId="26" captionText="Figure 21. Raveniola shixiu Lin, Wang &amp; Li, sp. nov., holotype female. A. Vulva, ventral view; B. Same, dorsal view. Abbreviation: S—spermatheca. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10941060" httpUri="https://zenodo.org/record/10941060/files/figure.png" pageId="27" pageNumber="31">Fig. 21</figureCitation>
). Spermathecae white, 5 times longer than wide, weakly sclerotized. Pore glands denser anteriorly, becoming less numerous proximally. Apices blunt, unexpanded. Spermathecae separated by their length.
</paragraph>
<caption id="5DC7A0A6FF9FFFF97D8CF83EC62DF7E2" ID-DOI="http://doi.org/10.5281/zenodo.10941062" ID-Zenodo-Dep="10941062" httpUri="https://zenodo.org/record/10941062/files/figure.png" pageId="27" pageNumber="31" startId="27.[155,218,2028,2050]" targetBox="[189,1399,714,1991]" targetPageId="27" targetType="figure">
<paragraph id="0907F02EFF9FFFF97D8CF83EC62DF7E2" blockId="27.[155,1433,2028,2082]" pageId="27" pageNumber="31">
Figure 22.
<taxonomicName id="CEB88BADFF9FFFF97C1CF83EC036F7C1" authority="Lin, Wang &amp; Li" authorityName="Lin, Wang &amp; Li" authorityYear="2024" box="[267,620,2028,2050]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="31" phylum="Arthropoda" rank="species" species="xiezhen" status="sp. nov.">
<emphasis id="3BCC2C3CFF9FFFF97C1CF83EC3E4F7C2" box="[267,446,2028,2049]" italics="true" pageId="27" pageNumber="31">Raveniola xiezhen</emphasis>
Lin, Wang &amp; Li
</taxonomicName>
,
<emphasis id="3BCC2C3CFF9FFFF97F6DF83EC091F7C2" bold="true" box="[634,715,2028,2049]" pageId="27" pageNumber="31">
<taxonomicNameLabel id="20FF9147FF9FFFF97F6DF83EC091F7C2" box="[634,715,2028,2049]" pageId="27" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A. Left palp, ventral view; B. Same, retrolateral view; C. Psembolus, retrolateral view. Abbreviation: Pb—psembolus. Scale bars: A–B= 0.5 mm; C =0.1 mm.
</paragraph>
</caption>
<paragraph id="0907F02EFF98FFFE7DAAFEDDC328FEE4" blockId="28.[136,1452,236,399]" box="[189,370,271,295]" pageId="28" pageNumber="32">Male. Unknown.</paragraph>
<paragraph id="0907F02EFF98FFFE7DAAFEE0C1AFFEAF" blockId="28.[136,1452,236,399]" pageId="28" pageNumber="32">
<materialsCitation id="B9D0FA73FF98FFFE7DAAFEE0C1ABFEAF" collectingDate="2023-06-15" collectorName="Ming Yi" country="China" location="Material" pageId="28" pageNumber="32" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<location id="0C67A6F5FF98FFFE7DAAFEE0C34DFE89" LSID="urn:lsid:plazi:treatment:81114138FF9EFFFE7D9FFE6EC099FE4C:0C67A6F5FF98FFFE7DAAFEE0C34DFE89" box="[189,279,306,330]" country="China" name="Material" pageId="28" pageNumber="32">Material</location>
examined.
<typeStatus id="D6034E8CFF98FFFE7C85FEE0C3AFFE89" box="[402,501,306,330]" pageId="28" pageNumber="32" type="holotype">Holotype</typeStatus>
<specimenCount id="1FBE3BA7FF98FFFE7CECFEE0C055FE8A" box="[507,527,306,329]" count="1" pageId="28" pageNumber="32" type="female">♀</specimenCount>
(IZCAS-Ar44817),
<collectingCountry id="71AFB0BEFF98FFFE7FFFFEE0C176FE89" box="[744,812,306,330]" name="China" pageId="28" pageNumber="32">China</collectingCountry>
:
<taxonomicName id="CEB88BADFF98FFFE7E22FEE0C7F2FE89" authority=", Guilin City, Lingui Dist., Huixian Town, Douyan" authorityName="Guilin City, Lingui Dist., Huixian Town, Douyan" box="[821,1448,306,330]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="32" phylum="Arthropoda" rank="species" species="guangxi">Guangxi, Guilin City, Lingui Dist., Huixian Town, Douyan</taxonomicName>
,
<date id="7D06D6EEFF98FFFE7D9FFE86C359FEAF" box="[136,259,340,364]" pageId="28" pageNumber="32" value="2023-06-15">
<collectingDate id="6D422F06FF98FFFE7D9FFE86C359FEAF" box="[136,259,340,364]" pageId="28" pageNumber="32" value="2023-06-15">15.VI.2023</collectingDate>
</date>
,
<collectorName id="A44D95F8FF98FFFE7C18FE86C332FEAF" box="[271,360,340,364]" pageId="28" pageNumber="32">Ming Yi</collectorName>
leg.
<typeStatus id="D6034E8CFF98FFFE7C8BFE87C3A7FEAE" box="[412,509,341,365]" pageId="28" pageNumber="32" type="paratype">Paratype</typeStatus>
.
<specimenCount id="1FBE3BA7FF98FFFE7F10FE86C073FEAF" box="[519,553,340,364]" count="1" pageId="28" pageNumber="32" type="female">1♀</specimenCount>
(IZCAS-Ar44818), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="0907F02EFF98FFFE7DAAFEA5C099FE4C" blockId="28.[136,1452,236,399]" box="[189,707,375,399]" pageId="28" pageNumber="32">
Distribution. Known only from the
<typeStatus id="D6034E8CFF98FFFE7F2FFEA5C03CFE4C" box="[568,614,375,399]" pageId="28" pageNumber="32">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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