treatments-xml/data/F0/7D/52/F07D525F1B3D5ED18F61F19DE43D5D0E.xml

<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.34.50673" ID-GBIF-Dataset="bfacffac-2241-41a0-af08-9d8066f710b3" ID-Pensoft-Pub="1314-2615-34-109" ID-Pensoft-UUID="81A3A37D1191535A9AF7B8BFA11CD996" ID-ZooBank="522749328D7D4DAC94D89DF52EB60ACA" ModsDocID="1314-2615-34-109" checkinTime="1590646549751" checkinUser="pensoft" docAuthor="Palacios-Vargas, Jose G." docDate="2020" docId="F07D525F1B3D5ED18F61F19DE43D5D0E" docLanguage="en" docName="SubterBiol 34: 109-119" docOrigin="Subterranean Biology 34" docSource="http://dx.doi.org/10.3897/subtbiol.34.50673" docTitle="Acheroxenylla lipsae Palacios-Vargas 2020, sp. nov." docType="treatment" docUuid="3EF8532D-FD4F-4DB2-A547-5C60CE1107C5" docUuidSource="ZooBank" docVersion="4" id="81A3A37D1191535A9AF7B8BFA11CD996" lastPageNumber="109" masterDocId="81A3A37D1191535A9AF7B8BFA11CD996" masterDocTitle="Acheroxenylla (Collembola, Hypogastruridae), first record from the Americas with description of a new species from a Peruvian cave" masterLastPageNumber="119" masterPageNumber="109" pageNumber="109" updateTime="1643443090584" updateUser="ExternalLinkService">
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<mods:title>Acheroxenylla (Collembola, Hypogastruridae), first record from the Americas with description of a new species from a Peruvian cave</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Palacios-Vargas, Jose G.</mods:namePart>
<mods:affiliation>Laboratorio de Ecologia y Sistematica de Microartropodos, Depto. Ecologia y Recursos Naturales, Facultad de Ciencias, UNAM, Coyoacan 04510, Mexico</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-9097-6813</mods:nameIdentifier>
<mods:nameIdentifier type="email">troglolaphysa@hotmail.com</mods:nameIdentifier>
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<mods:title>Subterranean Biology</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>34</mods:number>
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<mods:extent unit="page">
<mods:start>109</mods:start>
<mods:end>119</mods:end>
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<mods:url>http://dx.doi.org/10.3897/subtbiol.34.50673</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.34.50673</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2615-34-109</mods:identifier>
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<treatment ID-GBIF-Taxon="164363168" LSID="urn:lsid:zoobank.org:act:3EF8532D-FD4F-4DB2-A547-5C60CE1107C5" httpUri="http://treatment.plazi.org/id/F07D525F1B3D5ED18F61F19DE43D5D0E" lastPageNumber="109" pageId="0" pageNumber="109">
<subSubSection pageId="0" pageNumber="109" type="nomenclature">
<paragraph pageId="0" pageNumber="109">
<taxonomicName LSID="F07D525F-1B3D-5ED1-8F61-F19DE43D5D0E" authority="Palacios-Vargas, 2020" authorityName="Palacios-Vargas" authorityYear="2020" class="Collembola" family="Hypogastruridae" genus="Acheroxenylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acheroxenylla lipsae" order="Collembola" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Lipsae" status="sp. nov.">Acheroxenylla lipsae</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="109">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. 1 Acheroxenylla lipsae sp. nov. 1, group of specimens floating on water close to the guano of oil birds 2 two specimens dorsal view just collected in ethanol 96 %; photos 1 and 2 by Josiane Lips 3 dorsal view 4 lateral view, photos 3 and 4 by Maira Montejo and Angela Arango, scale in photos 3 and 4 is 150 μm." figureDoi="10.3897/subtbiol.34.50673.figures1-4" httpUri="https://binary.pensoft.net/fig/414551" pageId="0" pageNumber="109">Figures 1-4</figureCitation>
<figureCitation captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Acheroxenylla lipsae sp. nov. 5 ant IV dorsal view 6 head chaetotaxy 7 Tita I." figureDoi="10.3897/subtbiol.34.50673.figures5-7" httpUri="https://binary.pensoft.net/fig/414552" pageId="0" pageNumber="109">, 5-7</figureCitation>
<figureCitation captionStart="Figures 8, 9" captionStartId="F3" captionText="Figures 8, 9. Acheroxenylla lipsae sp. nov. 8 dorsal chaetotaxy of Th. I - III 9 Tita III." figureDoi="10.3897/subtbiol.34.50673.figures8-9" httpUri="https://binary.pensoft.net/fig/414553" pageId="0" pageNumber="109">, 8, 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F4" captionText="Figure 10. Acheroxenylla lipsae sp. nov. Dorsal abdominal chaetotaxy." figureDoi="10.3897/subtbiol.34.50673.figure10" httpUri="https://binary.pensoft.net/fig/414554" pageId="0" pageNumber="109">, 10</figureCitation>
<figureCitation captionStart="Figures 11, 12" captionStartId="F5" captionText="Figures 11, 12. Acheroxenylla lipsae sp. nov. 11. Ventral abdominal chaetotaxy from Abd. III to V, with retinaculum and furcula; 12. two mucrodens in lateral position." figureDoi="10.3897/subtbiol.34.50673.figures11-12" httpUri="https://binary.pensoft.net/fig/414555" pageId="0" pageNumber="109">, 11-12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="description">
<paragraph pageId="0" pageNumber="109">Description.</paragraph>
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" italics="true" pageId="0" pageNumber="109">Holotype female</emphasis>
(number FC-UNAM 22501) and one paratype female (numbers FC- UNAM 22502) are kept at Dept. Entomology; Museo de Historia Natural, Universidad Mayor de San Marcos, 
<normalizedToken originalValue="Perú">Peru</normalizedToken>
; two paratypes females one male and one juvenile (numbers FC-UNAM 22500, 22503 to 22325) are kept at Mexican 
<taxonomicName authorityName="Ellis" authorityYear="1976" lsidName="" order="Collembola" pageId="0" pageNumber="109" rank="order">Collembola</taxonomicName>
collection at Facultad de Ciencias, UNAM.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="type locality">
<paragraph pageId="0" pageNumber="109">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="109">
<normalizedToken originalValue="Perú">Peru</normalizedToken>
: Region San 
<normalizedToken originalValue="Martín">Martin</normalizedToken>
; Province Rioja, Cueva de Samuel (6°06'92&quot;S, 
<geoCoordinate degrees="77" direction="west" minutes="31" orientation="longitude" precision="15" seconds="58" value="-77.53278">77°31'58&quot;W</geoCoordinate>
) 1,720 m a.s.l. About 5.5 Km North-West of town Naciente de Rio Negro. 16-viii-2017, sample 14470, J. Lips col.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="diagnosis">
<paragraph pageId="0" pageNumber="109">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName authorityName="Palacios-Vargas" authorityYear="2020" class="Collembola" family="Hypogastruridae" genus="Acheroxenylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acheroxenylla lipsae" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Lipsae">
<emphasis italics="true" pageId="0" pageNumber="109">Acheroxenylla lipsae</emphasis>
</taxonomicName>
sp. nov. is characterized by the presence of a well-developed furcula with mucro, long dente with two long dental setae each, three manubrial setae and a retinaculum with three teeth. Tibiotarsi are longer than in other species known.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="description">
<paragraph pageId="0" pageNumber="109">Description.</paragraph>
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" italics="true" pageId="0" pageNumber="109">Body length</emphasis>
(average of 7 specimens) = 1.25 mm. Setae not differentiated in macro and microsetae, all smooth and sharp mesosetae about 11 
<normalizedToken originalValue="μm">μm</normalizedToken>
with small barbulations. Sensorial setae longer than regular setae, about 30 
<normalizedToken originalValue="μm">μm</normalizedToken>
. Sensorial formula as 022/11111. Color, some specimens (Figures 
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. 1 Acheroxenylla lipsae sp. nov. 1, group of specimens floating on water close to the guano of oil birds 2 two specimens dorsal view just collected in ethanol 96 %; photos 1 and 2 by Josiane Lips 3 dorsal view 4 lateral view, photos 3 and 4 by Maira Montejo and Angela Arango, scale in photos 3 and 4 is 150 μm." figureDoi="10.3897/subtbiol.34.50673.figures1-4" httpUri="https://binary.pensoft.net/fig/414551" pageId="0" pageNumber="109">1</figureCitation>
, 
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. 1 Acheroxenylla lipsae sp. nov. 1, group of specimens floating on water close to the guano of oil birds 2 two specimens dorsal view just collected in ethanol 96 %; photos 1 and 2 by Josiane Lips 3 dorsal view 4 lateral view, photos 3 and 4 by Maira Montejo and Angela Arango, scale in photos 3 and 4 is 150 μm." figureDoi="10.3897/subtbiol.34.50673.figures1-4" httpUri="https://binary.pensoft.net/fig/414551" pageId="0" pageNumber="109">2</figureCitation>
) with very dark eyespots; others gray with small patches of blue color on body and black eyespots (Figures 
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. 1 Acheroxenylla lipsae sp. nov. 1, group of specimens floating on water close to the guano of oil birds 2 two specimens dorsal view just collected in ethanol 96 %; photos 1 and 2 by Josiane Lips 3 dorsal view 4 lateral view, photos 3 and 4 by Maira Montejo and Angela Arango, scale in photos 3 and 4 is 150 μm." figureDoi="10.3897/subtbiol.34.50673.figures1-4" httpUri="https://binary.pensoft.net/fig/414551" pageId="0" pageNumber="109">3</figureCitation>
, 
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. 1 Acheroxenylla lipsae sp. nov. 1, group of specimens floating on water close to the guano of oil birds 2 two specimens dorsal view just collected in ethanol 96 %; photos 1 and 2 by Josiane Lips 3 dorsal view 4 lateral view, photos 3 and 4 by Maira Montejo and Angela Arango, scale in photos 3 and 4 is 150 μm." figureDoi="10.3897/subtbiol.34.50673.figures1-4" httpUri="https://binary.pensoft.net/fig/414551" pageId="0" pageNumber="109">4</figureCitation>
). Cuticular granulation strong, 
<normalizedToken originalValue="Yossi´s">Yossi's</normalizedToken>
parameter 5 or 6. Ratio of head: antenna = 1:0.8 labrum formula: 2/5,5,4.
</paragraph>
<caption doi="10.3897/subtbiol.34.50673.figures1-4" httpUri="https://binary.pensoft.net/fig/414551" pageId="0" pageNumber="109" start="Figures 1–4" startId="F1">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figures 1-4.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="109">1</emphasis>
<taxonomicName authorityName="Palacios-Vargas" authorityYear="2020" class="Collembola" family="Hypogastruridae" genus="Acheroxenylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acheroxenylla lipsae" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Lipsae">
<emphasis italics="true" pageId="0" pageNumber="109">Acheroxenylla lipsae</emphasis>
</taxonomicName>
sp. nov. 1, group of specimens floating on water close to the guano of oil birds 
<emphasis bold="true" pageId="0" pageNumber="109">2</emphasis>
two specimens dorsal view just collected in ethanol 96%; photos 1 and 2 by Josiane Lips 
<emphasis bold="true" pageId="0" pageNumber="109">3</emphasis>
dorsal view 
<emphasis bold="true" pageId="0" pageNumber="109">4</emphasis>
lateral view, photos 3 and 4 by Maira Montejo and Angela Arango, scale in photos 3 and 4 is 150 
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" italics="true" pageId="0" pageNumber="109">Ant I</emphasis>
with seven dorsal setae, Ant II with 12 setae. Ant III with 17 setae in two whorls, sense organ with two free club-shaped microsensilla, not covered by tegumentary fold; two short guard sensilla (Sgd and Sgv) of same shape and size, and one ventral microsensillum. No eversible sac between Ant III-IV. Ant IV with four cylindrical sensilla, one dorsal and three latero-external; subapical organite, lateral microsensillum and simple subapical bulb (Figure 
<figureCitation captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Acheroxenylla lipsae sp. nov. 5 ant IV dorsal view 6 head chaetotaxy 7 Tita I." figureDoi="10.3897/subtbiol.34.50673.figures5-7" httpUri="https://binary.pensoft.net/fig/414552" pageId="0" pageNumber="109">5</figureCitation>
), no sensory file on ventral side. Ratio of Ant I: II; III-IV = 1:1.1; 1.4; 3.8.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" italics="true" pageId="0" pageNumber="109">Head</emphasis>
with typical chaetotaxy for the genus, similar to 
<taxonomicName authorityName="Tullberg" authorityYear="1869" class="Collembola" family="Hypogastruridae" genus="Xenylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenylla" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Xenylla</emphasis>
</taxonomicName>
, only 3 subdorsal setae, seta c1 and only 1 setae v (v1), and 3 subequal setae in ocular area (Figure 
<figureCitation captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Acheroxenylla lipsae sp. nov. 5 ant IV dorsal view 6 head chaetotaxy 7 Tita I." figureDoi="10.3897/subtbiol.34.50673.figures5-7" httpUri="https://binary.pensoft.net/fig/414552" pageId="0" pageNumber="109">6</figureCitation>
). 2 + 2 eyes of about equal diameter with very strong granulations of dark pigment. PAO absent. Labium with 4 + 4 setae (one longer than others); 3 pairs of postlabial setae. Mandible with 3-4 apical teeth, and normal molar plate. Maxilla with six lamellae. Th I with 3 + 3 dorsal setae and 1 + 1 lateral on upper subcoxae. Each Th II and III with 3 irregular rows of setae (Figure 
<figureCitation captionStart="Figures 8, 9" captionStartId="F3" captionText="Figures 8, 9. Acheroxenylla lipsae sp. nov. 8 dorsal chaetotaxy of Th. I - III 9 Tita III." figureDoi="10.3897/subtbiol.34.50673.figures8-9" httpUri="https://binary.pensoft.net/fig/414553" pageId="0" pageNumber="109">8</figureCitation>
), sensorial setae m6 and p4 as usual.
</paragraph>
<caption doi="10.3897/subtbiol.34.50673.figures5-7" httpUri="https://binary.pensoft.net/fig/414552" pageId="0" pageNumber="109" start="Figures 5–7" startId="F2">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figures 5-7.</emphasis>
<taxonomicName authorityName="Palacios-Vargas" authorityYear="2020" class="Collembola" family="Hypogastruridae" genus="Acheroxenylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acheroxenylla lipsae" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Lipsae">
<emphasis italics="true" pageId="0" pageNumber="109">Acheroxenylla lipsae</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="109">5</emphasis>
ant IV dorsal view 
<emphasis bold="true" pageId="0" pageNumber="109">6</emphasis>
head chaetotaxy 
<emphasis bold="true" pageId="0" pageNumber="109">7</emphasis>
Tita I.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" italics="true" pageId="0" pageNumber="109">Leg</emphasis>
chaetotaxy from I to III: precoxae 0,1,2; coxae 3,6-8,5-8; trochanters 5,5,4; femora 12,11,10 one ventral seta very long, as acuminate tenent hair; Tita 19,19,18 (Figures 
<figureCitation captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Acheroxenylla lipsae sp. nov. 5 ant IV dorsal view 6 head chaetotaxy 7 Tita I." figureDoi="10.3897/subtbiol.34.50673.figures5-7" httpUri="https://binary.pensoft.net/fig/414552" pageId="0" pageNumber="109">7</figureCitation>
, 
<figureCitation captionStart="Figures 8, 9" captionStartId="F3" captionText="Figures 8, 9. Acheroxenylla lipsae sp. nov. 8 dorsal chaetotaxy of Th. I - III 9 Tita III." figureDoi="10.3897/subtbiol.34.50673.figures8-9" httpUri="https://binary.pensoft.net/fig/414553" pageId="0" pageNumber="109">9</figureCitation>
); pretarsi 2,2,2. Two dorsal tenent hairs weakly clavate on dorso-distal whorl on Tita I and II; one on Tita III. Unguis thin, elongated, curving slightly, without any tooth (Figures 
<figureCitation captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Acheroxenylla lipsae sp. nov. 5 ant IV dorsal view 6 head chaetotaxy 7 Tita I." figureDoi="10.3897/subtbiol.34.50673.figures5-7" httpUri="https://binary.pensoft.net/fig/414552" pageId="0" pageNumber="109">7</figureCitation>
- 
<figureCitation captionStart="Figures 8, 9" captionStartId="F3" captionText="Figures 8, 9. Acheroxenylla lipsae sp. nov. 8 dorsal chaetotaxy of Th. I - III 9 Tita III." figureDoi="10.3897/subtbiol.34.50673.figures8-9" httpUri="https://binary.pensoft.net/fig/414553" pageId="0" pageNumber="109">9</figureCitation>
). No unguiculus. Ratio Tita/unguis = 1: 1.
</paragraph>
<caption doi="10.3897/subtbiol.34.50673.figures8-9" httpUri="https://binary.pensoft.net/fig/414553" pageId="0" pageNumber="109" start="Figures 8, 9" startId="F3">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figures 8, 9.</emphasis>
<taxonomicName authorityName="Palacios-Vargas" authorityYear="2020" class="Collembola" family="Hypogastruridae" genus="Acheroxenylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acheroxenylla lipsae" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Lipsae">
<emphasis italics="true" pageId="0" pageNumber="109">Acheroxenylla lipsae</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="109">8</emphasis>
dorsal chaetotaxy of Th. I - III 
<emphasis bold="true" pageId="0" pageNumber="109">9</emphasis>
Tita III.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" italics="true" pageId="0" pageNumber="109">Dorsal</emphasis>
chaetotaxy of abdomen as in Figure 
<figureCitation captionStart="Figure 10" captionStartId="F4" captionText="Figure 10. Acheroxenylla lipsae sp. nov. Dorsal abdominal chaetotaxy." figureDoi="10.3897/subtbiol.34.50673.figure10" httpUri="https://binary.pensoft.net/fig/414554" pageId="0" pageNumber="109">10</figureCitation>
. Abd I-III with 2 irregular rows of dorsal setae, 1 sensorial seta on P5, except Abd V with p3 as sensorial seta. Number of axial setae from Abd I to III is 2 + 2: Abd. IV with 3 + 3; Abd. V with a1 and a2, p3 is ss. Abd VI with 2 rows of setae, a1-3, p1 modified as spine, p 2-3 normal setae. Two small anal spines, as short as their tubercle.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Ventral chaetotaxy.</emphasis>
Thoracic sternites and Abd I without setae. Ventral tube with 4 + 4 setae. Abd II with 9 - 12 setae, one of them (p3) very long; Abd III with 6 setae, p3 slightly longer than others. Abd IV dorsolateral with 5 setae, one of them very long (Figure 
<figureCitation captionStart="Figure 10" captionStartId="F4" captionText="Figure 10. Acheroxenylla lipsae sp. nov. Dorsal abdominal chaetotaxy." figureDoi="10.3897/subtbiol.34.50673.figure10" httpUri="https://binary.pensoft.net/fig/414554" pageId="0" pageNumber="109">10</figureCitation>
). Retinaculum with 3 + 3 teeth, without seta on corpus (Figure 
<figureCitation captionStart="Figure 10" captionStartId="F4" captionText="Figure 10. Acheroxenylla lipsae sp. nov. Dorsal abdominal chaetotaxy." figureDoi="10.3897/subtbiol.34.50673.figure10" httpUri="https://binary.pensoft.net/fig/414554" pageId="0" pageNumber="109">10</figureCitation>
). Furcula well developed. Manubrium with 3 pairs of setae of same length. Dens dorsally with moderate granulation and with 2 subequal setae, with a smooth elongated area on anterior part of dens. Mucro more than the half-length of dens, long and narrow with one small outer lamella, apex curved and more sclerotized (Figure 
<figureCitation captionStart="Figures 11, 12" captionStartId="F5" captionText="Figures 11, 12. Acheroxenylla lipsae sp. nov. 11. Ventral abdominal chaetotaxy from Abd. III to V, with retinaculum and furcula; 12. two mucrodens in lateral position." figureDoi="10.3897/subtbiol.34.50673.figures11-12" httpUri="https://binary.pensoft.net/fig/414555" pageId="0" pageNumber="109">11</figureCitation>
). (Ratio Manubrium: dens; mucro = 1:0.8; 0.5). Mucro better delimited on anterior part by a clear notch (Figure 
<figureCitation captionStart="Figures 11, 12" captionStartId="F5" captionText="Figures 11, 12. Acheroxenylla lipsae sp. nov. 11. Ventral abdominal chaetotaxy from Abd. III to V, with retinaculum and furcula; 12. two mucrodens in lateral position." figureDoi="10.3897/subtbiol.34.50673.figures11-12" httpUri="https://binary.pensoft.net/fig/414555" pageId="0" pageNumber="109">12</figureCitation>
). Genital plate of female with 3+3 pregenital, 10-13 circumgenital and 1 + 1 eugenital setae. Genital plate of male with 3 + 3 pregenital, 44 circumgenital and 4 + 4 eugenital setae. Each anal valve with 13 regular and 1 hr setae.
</paragraph>
<caption doi="10.3897/subtbiol.34.50673.figure10" httpUri="https://binary.pensoft.net/fig/414554" pageId="0" pageNumber="109" start="Figure 10" startId="F4">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 10.</emphasis>
<taxonomicName authorityName="Palacios-Vargas" authorityYear="2020" class="Collembola" family="Hypogastruridae" genus="Acheroxenylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acheroxenylla lipsae" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Lipsae">
<emphasis italics="true" pageId="0" pageNumber="109">Acheroxenylla lipsae</emphasis>
</taxonomicName>
sp. nov. Dorsal abdominal chaetotaxy.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.34.50673.figures11-12" httpUri="https://binary.pensoft.net/fig/414555" pageId="0" pageNumber="109" start="Figures 11, 12" startId="F5">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figures 11, 12.</emphasis>
<taxonomicName authorityName="Palacios-Vargas" authorityYear="2020" class="Collembola" family="Hypogastruridae" genus="Acheroxenylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acheroxenylla lipsae" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Lipsae">
<emphasis italics="true" pageId="0" pageNumber="109">Acheroxenylla lipsae</emphasis>
</taxonomicName>
sp. nov. 11. Ventral abdominal chaetotaxy from Abd. III to V, with retinaculum and furcula; 12. two mucrodens in lateral position.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="variation">
<paragraph pageId="0" pageNumber="109">Variation.</paragraph>
<paragraph pageId="0" pageNumber="109">Some asymmetries on body chaetotaxy were observed. Several specimens have setae somehow displaced, giving the appearance of asymmetries. One case of supernumerary setae on left side of abdominal segment III was observed on one paratype, where there were 3 setae &quot;p4&quot;, giving the appearance of sensorial seta to be on position &quot;p7&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="etymology">
<paragraph pageId="0" pageNumber="109">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="109">
This species is dedicated to Josiane Lips, for her contribution to the knowledge of cave fauna from 
<normalizedToken originalValue="Perú">Peru</normalizedToken>
, Mexico and many other places.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="discussion">
<paragraph pageId="0" pageNumber="109">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="109">
This new species is the largest and most pigmented member of this genus. The main differences of 
<taxonomicName authorityName="Palacios-Vargas" authorityYear="2020" class="Collembola" family="Hypogastruridae" genus="Acheroxenylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acheroxenylla lipsae" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Lipsae">
<emphasis italics="true" pageId="0" pageNumber="109">Acheroxenylla lipsae</emphasis>
</taxonomicName>
sp. nov. and the other species is the presence of a well-developed furcula with mucro, long dens and two long dental setae, 3 + 3 manubrial setae and retinaculum with 3 + 3 teeth. 
<taxonomicName lsidName="A. furcata" pageId="0" pageNumber="109" rank="species" species="Furcata">
<emphasis italics="true" pageId="0" pageNumber="109">A. furcata</emphasis>
</taxonomicName>
has a reduced furcula, with no manubrial setae, very short dens with one seta each and mucro absent; its retinaculum has only two teeth on each ramus. The type species 
<taxonomicName lsidName="A. cretensis" pageId="0" pageNumber="109" rank="species" species="Cretensis">
<emphasis italics="true" pageId="0" pageNumber="109">A. cretensis</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="A. canariensis" pageId="0" pageNumber="109" rank="species" species="Canariensis">
<emphasis italics="true" pageId="0" pageNumber="109">A. canariensis</emphasis>
</taxonomicName>
lack completely the furcula and retinaculum. All the species have only two eyes per side; nevertheless, in the new species 
<taxonomicName lsidName="A. lipsae" pageId="0" pageNumber="109" rank="species" species="Lipsae">
<emphasis italics="true" pageId="0" pageNumber="109">A. lipsae</emphasis>
</taxonomicName>
sp. nov., eyes are better developed and closer to each other, and that is why their position seems to be &quot;D&quot; and &quot;E&quot;. After the drawings of 
<bibRefCitation author="Ellis, WN" journalOrPublisher="Tijdschrift voor Entomologie," pageId="0" pageNumber="109" pagination="221 - 326" refId="B3" refString="Ellis, WN, 1976. Autumn fauna of Collembola from Central Crete. Tijdschrift voor Entomologie, 119 (8): 221 - 326" title="Autumn fauna of Collembola from Central Crete." volume="119" year="1976">Ellis (1976)</bibRefCitation>
, the eyes of 
<taxonomicName lsidName="A. cretensis" pageId="0" pageNumber="109" rank="species" species="Cretensis">
<emphasis italics="true" pageId="0" pageNumber="109">A. cretensis</emphasis>
</taxonomicName>
are &quot;B&quot; and &quot;E&quot;; as pointed by 
<bibRefCitation author="Ellis, WN" journalOrPublisher="Tijdschrift voor Entomologie," pageId="0" pageNumber="109" pagination="221 - 326" refId="B3" refString="Ellis, WN, 1976. Autumn fauna of Collembola from Central Crete. Tijdschrift voor Entomologie, 119 (8): 221 - 326" title="Autumn fauna of Collembola from Central Crete." volume="119" year="1976">Ellis (1976)</bibRefCitation>
: &quot;only 2 + 2 eyes small, widely separated ocelli&quot;. Another difference is that 
<taxonomicName lsidName="A. cretensis" pageId="0" pageNumber="109" rank="species" species="Cretensis">
<emphasis italics="true" pageId="0" pageNumber="109">A. cretensis</emphasis>
</taxonomicName>
has only 2 subdorsal cephalic setae, while the new species has 3 pairs, setae sd5, sd4 and sd3, similar to 
<taxonomicName lsidName="A. furcata" pageId="0" pageNumber="109" rank="species" species="Furcata">
<emphasis italics="true" pageId="0" pageNumber="109">A. furcata</emphasis>
</taxonomicName>
. There are small differences in the head chaetotaxy, 
<taxonomicName lsidName="A. lipsae" pageId="0" pageNumber="109" rank="species" species="Lipsae">
<emphasis italics="true" pageId="0" pageNumber="109">A. lipsae</emphasis>
</taxonomicName>
sp. nov. has cephalic setae a0, as 
<taxonomicName lsidName="A. cretensis" pageId="0" pageNumber="109" rank="species" species="Cretensis">
<emphasis italics="true" pageId="0" pageNumber="109">A. cretensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="109">
About the pigment, 
<taxonomicName lsidName="A. furcata" pageId="0" pageNumber="109" rank="species" species="Furcata">
<emphasis italics="true" pageId="0" pageNumber="109">A. furcata</emphasis>
</taxonomicName>
is white with small spots under each of the 2 + 2 or 1 + 1 eyes and sometimes a scattered bluish gray pigment is present all over the body, while 
<taxonomicName lsidName="A. lipsae" pageId="0" pageNumber="109" rank="species" species="Lipsae">
<emphasis italics="true" pageId="0" pageNumber="109">A. lipsae</emphasis>
</taxonomicName>
sp. nov. is more pigmented. Tibiotarsi of 
<taxonomicName lsidName="A. furcata" pageId="0" pageNumber="109" rank="species" species="Furcata">
<emphasis italics="true" pageId="0" pageNumber="109">A. furcata</emphasis>
</taxonomicName>
with one apical tenent hair (A1), sometimes weakly clavate, is similar to the new species. The ungues of the Peruvian species is more elongated than in any other species of the genus (ratio tibiotarsus: ungues: 1: 1.0), so this may be a troglomorphic character, and also in the other species tibiotarsus is about twice the length of ungues.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="molecular results">
<paragraph pageId="0" pageNumber="109">Molecular results.</paragraph>
<paragraph pageId="0" pageNumber="109">
DNA was successfully obtained from three specimens, sequences BCICL008-19 (length 620bp), BCICL009-19 (657bp) and BCICL010 (632bp), which were deposited in the project BCICL of the Barcode of Life Data System (http://www.barcodinglife.org/index.php). Cuticles of two specimens were recovered and mounted in 
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
solution which represent the vouchers and are kept at the 
<normalizedToken originalValue="author’s">author's</normalizedToken>
institution as type material.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="description">
<paragraph pageId="0" pageNumber="109">Description of the cave.</paragraph>
<paragraph pageId="0" pageNumber="109">
The type locality is Cueva de Samuel. Collecting was done 200 m from the entrance on 
<normalizedToken originalValue="Guácharo">Guacharo</normalizedToken>
guano, where was the new species found. It is an active cave with water flowing from the cave. The air temperature was 15 °C. Its entrance is about 1,720 m a.s.l. In the first part of the cave there is a gallery with a colony of oil birds (
<normalizedToken originalValue="Guácharos">Guacharos</normalizedToken>
, 
<taxonomicName authorityName="Humboldt" authorityYear="1817" class="Aves" family="Steatornithidae" genus="Steatornis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Steatornis caripensis" order="Steatornithiformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="Caripensis">
<emphasis italics="true" pageId="0" pageNumber="109">Steatornis caripensis</emphasis>
</taxonomicName>
Humboldt, 1817). 500 m deep in the cave there is a big room with many big stalagmites, named Chachapoyas room. There are two small waterfalls, a small at 800 m from entrance and more far another of 10 m. One of the tunnels finishes in a well of more than 40 m.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="other collembola in the area">
<paragraph pageId="0" pageNumber="109">
Other 
<taxonomicName lsidName="" order="Collembola" pageId="0" pageNumber="109" rank="order">Collembola</taxonomicName>
in the area.
</paragraph>
<paragraph pageId="0" pageNumber="109">
There were 20 springtails found in the region of Rioja. In two caves only one species was found in each: Cueva del Lobo Perdido (
<taxonomicName authorityName="Schaeffer" authorityYear="1897" class="Collembola" family="Entomobryidae" genus="Pseudosinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudosinella" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Pseudosinella</emphasis>
</taxonomicName>
sp.) and Tragadero de Bellavista (
<taxonomicName class="Collembola" family="Paronellidae" genus="Cyphoderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphoderus" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Cyphoderus</emphasis>
</taxonomicName>
sp.). Two other caves had three 
<taxonomicName lsidName="" order="Collembola" pageId="0" pageNumber="109" rank="order">Collembola</taxonomicName>
: Piedra Brillante (
<taxonomicName authorityName="Mills" authorityYear="1938" class="Collembola" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trogolaphysa" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Trogolaphysa</emphasis>
</taxonomicName>
sp., 
<taxonomicName authorityName="Schaeffer" authorityYear="1897" class="Collembola" family="Entomobryidae" genus="Pseudosinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudosinella" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Pseudosinella</emphasis>
</taxonomicName>
sp., 
<taxonomicName authorityName="Bonet &amp; Tellez" authorityYear="1947" class="Collembola" family="Sminthuridae" genus="Pararrhopalites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararrhopalites" order="Symphypleona" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Pararrhopalites</emphasis>
</taxonomicName>
) and Cueva de los Loros (
<taxonomicName authorityName="Schaeffer" authorityYear="1897" class="Collembola" family="Entomobryidae" genus="Pseudosinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudosinella" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Pseudosinella</emphasis>
</taxonomicName>
sp., 
<taxonomicName authorityName="Schott" authorityYear="1893" class="Collembola" family="Orchesellidae" genus="Dicranocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranocentrus" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Dicranocentrus</emphasis>
</taxonomicName>
sp., 
<taxonomicName authorityName="Mills" authorityYear="1938" class="Collembola" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trogolaphysa" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Trogolaphysa</emphasis>
</taxonomicName>
sp.). In Cueva Palestina there were four species: (
<taxonomicName authorityName="Schaeffer" authorityYear="1897" class="Collembola" family="Entomobryidae" genus="Pseudosinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudosinella" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Pseudosinella</emphasis>
</taxonomicName>
sp., 
<taxonomicName authorityName="Willem" authorityYear="1902" class="Collembola" family="Isotomidae" genus="Folsomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Folsomia" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Folsomia</emphasis>
</taxonomicName>
sp., 
<taxonomicName class="Collembola" family="Brachystomellidae" genus="Folsomiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Folsomiella caeca" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Caeca">
<emphasis italics="true" pageId="0" pageNumber="109">Folsomiella caeca</emphasis>
</taxonomicName>
(Folsom, 1927), 
<taxonomicName baseAuthorityName="Rapoport &amp; Mano" baseAuthorityYear="1969" class="Collembola" family="Isotomidae" genus="Folsomides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Folsomides troglobius" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Troglobius">
<emphasis italics="true" pageId="0" pageNumber="109">Folsomides troglobius</emphasis>
</taxonomicName>
(Rapoport &amp; 
<normalizedToken originalValue="Maño">Mano</normalizedToken>
, 1969) while Cueva de Samuel was the most diverse with a new species of 
<taxonomicName authorityName="Ellis" authorityYear="1976" class="Collembola" family="Hypogastruridae" genus="Acheroxenylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acheroxenylla" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Acheroxenylla</emphasis>
</taxonomicName>
, and also specimens of 
<taxonomicName authorityName="Denis" authorityYear="1931" class="Collembola" family="Isotomidae" genus="Folsomina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Folsomina" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Folsomina</emphasis>
</taxonomicName>
sp., 
<taxonomicName authorityName="Schaeffer" authorityYear="1897" class="Collembola" family="Entomobryidae" genus="Pseudosinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudosinella" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Pseudosinella</emphasis>
</taxonomicName>
sp., 
<taxonomicName authorityName="Mills" authorityYear="1938" class="Collembola" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trogolaphysa" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Trogolaphysa</emphasis>
</taxonomicName>
sp., 
<taxonomicName class="Collembola" family="Paronellidae" genus="Cyphoderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphoderus" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Cyphoderus</emphasis>
</taxonomicName>
sp., 
<taxonomicName class="Collembola" family="Isotomidae" genus="Isotomurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isotomurus" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Isotomurus</emphasis>
</taxonomicName>
sp., 
<taxonomicName class="Collembola" family="Sminthuridae" genus="Pararrhopalites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararrhopalites ecuadorensis" order="Symphypleona" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Ecuadorensis">
<emphasis italics="true" pageId="0" pageNumber="109">Pararrhopalites ecuadorensis</emphasis>
</taxonomicName>
Bretfeld et Trinklein, 2000.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Collembola" family="Brachystomellidae" genus="Folsomiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Folsomiella caeca" order="Poduromorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Caeca">
<emphasis italics="true" pageId="0" pageNumber="109">Folsomiella caeca</emphasis>
</taxonomicName>
(Folsom, 1927) was described from limestone caves in bat dung at Panama. Later it was found at Ecuador in guano from several caves by 
<bibRefCitation author="Najt, J" journalOrPublisher="Bulletin du Museum National d'histore Naturelle, Section A Zoologie Biologie et Ecologie Animales" pageId="0" pageNumber="109" pagination="201 - 209" refId="B5" refString="Najt, J, Thibaud, J-M, 1987. Collemboles (Insecta) de l'Equteur. 1. Hypogastruridae, Neanuridae et Isotomidae. Bulletin du Museum National d'histore Naturelle, Section A Zoologie Biologie et Ecologie Animales 9 (1): 201 - 209" title="Collemboles (Insecta) de l'Equteur. 1. Hypogastruridae, Neanuridae et Isotomidae." volume="9" year="1987">Najt and Thibaud (1987)</bibRefCitation>
. As it also was collected by one of them in Venezuela and Peru, they have considered this species to edaphic - troglofile. 
<taxonomicName baseAuthorityName="Rapoport &amp; Mano" baseAuthorityYear="1969" class="Collembola" family="Isotomidae" genus="Folsomides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Folsomides troglobius" order="Entomobryomorpha" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Troglobius">
<emphasis italics="true" pageId="0" pageNumber="109">Folsomides troglobius</emphasis>
</taxonomicName>
(Rapoport et 
<normalizedToken originalValue="Maño">Mano</normalizedToken>
, 1969) was originally described from one cave close to Araira, and Cueva del 
<normalizedToken originalValue="Guácharo">Guacharo</normalizedToken>
in the state of Miranda, Venezuela. Later was cited by 
<bibRefCitation author="Najt, J" journalOrPublisher="Bulletin du Museum National d'histore Naturelle, Section A Zoologie Biologie et Ecologie Animales" pageId="0" pageNumber="109" pagination="201 - 209" refId="B5" refString="Najt, J, Thibaud, J-M, 1987. Collemboles (Insecta) de l'Equteur. 1. Hypogastruridae, Neanuridae et Isotomidae. Bulletin du Museum National d'histore Naturelle, Section A Zoologie Biologie et Ecologie Animales 9 (1): 201 - 209" title="Collemboles (Insecta) de l'Equteur. 1. Hypogastruridae, Neanuridae et Isotomidae." volume="9" year="1987">Najt and Thibaud (1987)</bibRefCitation>
from the cave of Barberanes in Ecuador. This is the first time that both species has been cited from Cueva Palestina in 
<normalizedToken originalValue="Perú">Peru</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="109">
In Cueva Samuel, besides the new species, there were other six 
<taxonomicName authorityName="Stach" authorityYear="1922" lsidName="" order="Collembola" pageId="0" pageNumber="109" rank="order">Collembola</taxonomicName>
found in different samples. Among them, 
<taxonomicName class="Collembola" family="Sminthuridae" genus="Pararrhopalites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararrhopalites ecuadorensis" order="Symphypleona" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="Ecuadorensis">
<emphasis italics="true" pageId="0" pageNumber="109">Pararrhopalites ecuadorensis</emphasis>
</taxonomicName>
Bretfeld et Trinklein, 2000 was described from one cave from Otonga (Cotopaxi), Ecuador and now is found for the first time from Peru.
</paragraph>
</subSubSection>
</treatment>
</document>