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<document id="F42D6A06C04175F2F5BF7C3D674AF3E6" ID-DOI="10.21248/contrib.entomol.69.1.091-173" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742724" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638269330072" checkinUser="felipe" docAuthor="Assing, Volker &amp; Schülke, Michael" docDate="2019" docId="F078561B726A7176FF2B6452FC00FA7E" docLanguage="en" docName="ContribEntomol.69.1.91-173.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 69 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Omalium kociani ZANETTI 2019, spec. nov." docType="treatment" docUuid="5F24413E-6AB4-4D1C-B567-AB93C38380C9" docUuidSource="ZooBank" docVersion="3" lastPageNumber="152" masterDocId="0C412E637256714BFFD26338F915FFC6" masterDocTitle="The Staphylinidae of Armenia and Nagorno-Karabakh (Coleoptera)" masterLastPageNumber="173" masterPageNumber="91" pageNumber="151" updateTime="1699437206044" updateUser="plazi">
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<mods:title id="B47FFB5DA2C790C3344773B81BA4D039">The Staphylinidae of Armenia and Nagorno-Karabakh (Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name id="19C71894E1EAC210ED580FF3A2D5EA2A" type="personal">
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<mods:namePart id="C66EBB88ABEAD76268563F0447658B01">Assing, Volker</mods:namePart>
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<mods:namePart id="BD66DA7FED9EC6F9636945013FAFACD3">Schülke, Michael</mods:namePart>
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<mods:title id="A641A67D74382318927D24F229D8E055">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="C8A4449D44045CE5D64559B4F7B56188">2019</mods:date>
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<mods:number id="1514E783A4F0D3F12637269A722C84EB">2019-06-24</mods:number>
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<mods:number id="4FE94DD2FB26BFBC9385708C201A8101">69</mods:number>
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<treatment id="F078561B726A7176FF2B6452FC00FA7E" LSID="urn:lsid:zoobank.org:act:5F24413E-6AB4-4D1C-B567-AB93C38380C9" httpUri="http://treatment.plazi.org/id/F078561B726A7176FF2B6452FC00FA7E" lastPageId="61" lastPageNumber="152" pageId="60" pageNumber="151">
<subSubSection id="30CBB486726A7177FF2B6452FA16F863" pageId="60" pageNumber="151" type="nomenclature">
<paragraph id="786EE70D726A7177FF2B6452FB7FF840" blockId="60.[249,771,1898,1989]" box="[249,618,1898,1926]" pageId="60" pageNumber="151">
<heading id="23265061726A7177FF2B6452FB7FF840" box="[249,618,1898,1926]" fontSize="12" level="2" pageId="60" pageNumber="151" reason="2">
<taxonomicName id="BFD19C8E726A7177FF2B6452F8EFF840" ID-CoL="8Y6XJ" authority="ZANETTI" authorityName="ZANETTI" authorityYear="2019" box="[249,506,1898,1926]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="151" phylum="Arthropoda" rank="species" species="kociani" status="sp. nov.">
<emphasis id="4AA53B1F726A7177FF2B6452F8B1F840" box="[249,420,1898,1926]" italics="true" pageId="60" pageNumber="151">Omalium kociani</emphasis>
ZANETTI
</taxonomicName>
<taxonomicNameLabel id="51968664726A7177FDD36452FB7FF840" box="[513,618,1898,1926]" pageId="60" pageNumber="151" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="30CBB486726A7177FF2B6489F875F803" box="[249,352,1969,1989]" pageId="60" pageNumber="151" type="description">
<paragraph id="786EE70D726A7177FF2B6489F875F803" blockId="60.[249,771,1898,1989]" box="[249,352,1969,1989]" pageId="60" pageNumber="151">
(
<figureCitation id="E0EAFB88726A7177FF2F6489F838F803" box="[253,301,1969,1989]" captionStart="Figs 1–6" captionStartId="56.[212,252,2003,2025]" captionTargetBox="[212,1459,271,1981]" captionTargetId="figure-27@56.[211,1459,254,1994]" captionTargetPageId="56" captionText="Figs 1–6: Type locality ofBellatheta khustupica in the peak region of Mount Khustup (1); microhabitat whereB. khustupica was" figureDoi="http://doi.org/10.5281/zenodo.5742740" httpUri="https://zenodo.org/record/5742740/files/figure.png" pageId="60" pageNumber="151">Figs 3</figureCitation>
,
<figureCitation id="E0EAFB88726A7177FEE66489F84EF803" box="[308,347,1969,1989]" captionStart="Figs 7–23" captionStartId="57.[178,218,1988,2010]" captionTargetBox="[184,1416,256,1980]" captionTargetId="figure-27@57.[177,1417,254,1983]" captionTargetPageId="57" captionText="Figs 7–23:Omalium kociani (7–10), Proteinus baculatus (11–14), Bryaxis armeniacus (15, 17), B. meghruicus (16, 18), Aleochara fugax (19–21), and Atheta meghruica (22–23): (median lobe of) aedeagus in lateral and in ventral view (7–8, 13–16); female accessory sclerites (9); spermatheca (10, 21); habitus (11, 19, 22); thorax in ventral view (12); male antennomeres I–III (17–18); posterior margin of female sternite VIII (20); forebody (23). Scale bars: 11, 19, 22: 1.0 mm; 12, 23: 0.5 mm; 7–10, 13–16, 20–21: 0.1 mm; 17–18: without scale." figureDoi="http://doi.org/10.5281/zenodo.5742742" httpUri="https://zenodo.org/record/5742742/files/figure.png" pageId="60" pageNumber="151">7–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="30CBB486726A7177FF0664D8FD31F7FE" pageId="60" pageNumber="151" type="materials_examined">
<paragraph id="786EE70D726A7177FF0664D8FD31F7FE" blockId="60.[212,823,2015,2104]" lastBlockId="60.[854,1465,2016,2104]" pageId="60" pageNumber="151">
<emphasis id="4AA53B1F726A7177FF0664D8F878F831" bold="true" box="[212,365,2015,2040]" pageId="60" pageNumber="151">Type material</emphasis>
:
<materialsCitation id="C8B9ED50726A7177FEAF64D8FAD2F7DE" collectingDate="2015-05-21" collectionCode="MCSNV" collectorName="M. Kocian" country="Armenia" determinerName="A. Zanetti" elevation="2400" latitude="39.839054" location="Jermuk" longLatPrecision="1" longitude="45.693497" pageId="60" pageNumber="151" specimenCount="2" typeStatus="holotype">
<typeStatus id="A76A59AF726A7177FEAF64D8F8F3F83E" box="[381,486,2016,2040]" pageId="60" pageNumber="151" type="holotype">Holotype</typeStatus>
: “
<collectingCountry id="00C6A79D726A7177FDC464D8FB9BF83E" box="[534,654,2016,2040]" name="Armenia" pageId="60" pageNumber="151">ARMENIA</collectingCountry>
, above
<location id="7D0EB1D6726A7177FD3064D8FA22F83E" LSID="urn:lsid:plazi:treatment:F078561B726A7176FF2B6452FC00FA7E:7D0EB1D6726A7177FD3064D8FA22F83E" box="[738,823,2016,2040]" country="Armenia" latitude="39.839054" longLatPrecision="1" longitude="45.693497" name="Jermuk" pageId="60" pageNumber="151">Jermuk</location>
sifting of plant leavings near snow residues,
<quantity id="BF294AE8726A7177FD336B38FA22F7D1" box="[737,823,2048,2072]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="60" pageNumber="151" unit="m" value="2400.0">
<elevation id="F3FC003E726A7177FD336B38FA22F7D1" box="[737,823,2048,2072]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="60" pageNumber="151" unit="m" value="2400.0">2400 m</elevation>
</quantity>
<geoCoordinate id="1DE581CA726A7177FF066B18F841F7FE" box="[212,340,2080,2104]" degrees="39,839053" direction="north" orientation="latitude" pageId="60" pageNumber="151" precision="1" value="39.839054">39,839053N</geoCoordinate>
<geoCoordinate id="1DE581CA726A7177FEB56B18F8F6F7FE" box="[359,483,2080,2104]" degrees="45,693496" direction="east" orientation="longitude" pageId="60" pageNumber="151" precision="1" value="45.693497">45,693496E</geoCoordinate>
<date id="0C6FC1CD726A7177FE246B18FB4BF7FE" box="[502,606,2080,2104]" pageId="60" pageNumber="151" value="2015-05-21">
<collectingDate id="1C2B3825726A7177FE246B18FB4BF7FE" box="[502,606,2080,2104]" pageId="60" pageNumber="151" value="2015-05-21">21.V.2015</collectingDate>
</date>
<collectorName id="D52482DB726A7177FDA26B18FBF5F7FE" box="[624,736,2080,2104]" pageId="60" pageNumber="151">M. Kocian</collectorName>
lgt. /
<taxonomicName id="BFD19C8E726A7177FC8464D8FD18F83E" authority="Assing &amp; Schülke, 2019" authorityName="Assing &amp; Schülke" authorityYear="2019" box="[854,1037,2016,2040]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="151" phylum="Arthropoda" rank="species" species="kociani" status="sp. nov.">
<emphasis id="4AA53B1F726A7177FC8464D8FD18F83E" box="[854,1037,2016,2040]" italics="true" pageId="60" pageNumber="151">Omalium kociani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51968664726A7177FBC164D8FD5EF831" box="[1043,1099,2016,2039]" pageId="60" pageNumber="151" rank="species">n. sp.</taxonomicNameLabel>
det.
<determinerName id="8DB20BC9726A7177FBAD64D8FDFEF83E" box="[1151,1259,2016,2040]" pageId="60" pageNumber="151">A. Zanetti</determinerName>
2016 /
<typeStatus id="A76A59AF726A7177FAE864D8FCBAF83E" box="[1338,1455,2016,2040]" pageId="60" pageNumber="151" type="holotype">Holotypus</typeStatus>
” (
<collectionCode id="1EC07FC8726A7177FC8F6B38FAD5F7DE" box="[861,960,2048,2072]" pageId="60" pageNumber="151">MCSNV</collectionCode>
)
</materialsCitation>
.
<typeStatus id="A76A59AF726A7177FC026B38FD2FF7DE" box="[976,1082,2048,2072]" pageId="60" pageNumber="151" type="paratype">Paratypes</typeStatus>
: 5, 3: same data as holotype (cKoc, cZan, cAss).
</paragraph>
</subSubSection>
<subSubSection id="30CBB486726B7176FF7C6238FBAAFEFE" pageId="61" pageNumber="152" type="etymology">
<paragraph id="786EE70D726B7176FF7C6238FA04FEDE" blockId="61.[174,785,256,312]" box="[174,785,256,280]" pageId="61" pageNumber="152">
<emphasis id="4AA53B1F726B7176FF7C6238F830FEDE" bold="true" box="[174,293,256,280]" pageId="61" pageNumber="152">Etymology</emphasis>
: This species is dedicated to the collector,
</paragraph>
<paragraph id="786EE70D726B7176FF7C6218FBAAFEFE" blockId="61.[174,785,256,312]" box="[174,703,288,312]" pageId="61" pageNumber="152">
Matúš Kocian (Praha), specialist of
<taxonomicName id="BFD19C8E726B7176FDF56218FBAEFEFE" authorityName="MacLeay" authorityYear="1825" box="[551,699,288,312]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="subFamily" subFamily="Tachyporinae">Tachyporinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="30CBB486726B7176FF7C6267FAB3FAD1" pageId="61" pageNumber="152" type="description">
<paragraph id="786EE70D726B7176FF7C6267F8BFFCDE" blockId="61.[174,785,351,2072]" pageId="61" pageNumber="152">
<emphasis id="4AA53B1F726B7176FF7C6267F825FEB1" bold="true" box="[174,304,351,375]" pageId="61" pageNumber="152">Description</emphasis>
: Measurements (n = 9): body length (with extended abdomen)
<quantity id="BF294AE8726B7176FE4F62B8FB0BFE51" box="[413,542,384,408]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.1" metricValueMin="2.4" pageId="61" pageNumber="152" unit="mm" value="2.75" valueMax="3.1" valueMin="2.4">2.4–3.1 mm</quantity>
; length of forebody from anterior margin of clypeus to posterior margin of elytra
<quantity id="BF294AE8726B7176FF2462F8F884FE11" box="[246,401,448,472]" metricMagnitude="-3" metricUnit="m" metricValue="1.705" metricValueMax="1.88" metricValueMin="1.53" pageId="61" pageNumber="152" unit="mm" value="1.705" valueMax="1.88" valueMin="1.53">1.53–1.88 mm</quantity>
; head width
<quantity id="BF294AE8726B7176FDFE62F8FBD2FE11" box="[556,711,448,472]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="61" pageNumber="152" unit="mm" value="0.485" valueMax="0.51" valueMin="0.46">0.46–0.51 mm</quantity>
; head length from clypeus to neck
<quantity id="BF294AE8726B7176FE2D62D8FB8FFE31" box="[511,666,480,504]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="4.2" metricValueMin="3.5" pageId="61" pageNumber="152" unit="mm" value="0.385" valueMax="0.42" valueMin="0.35">0.35–0.42 mm</quantity>
; width of pronotum
<quantity id="BF294AE8726B7176FEF96138F8D0FDD1" box="[299,453,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="7.0" metricValueMin="6.1" pageId="61" pageNumber="152" unit="mm" value="0.655" valueMax="0.7" valueMin="0.61">0.61–0.70 mm</quantity>
; length of pronotum
<quantity id="BF294AE8726B7176FD026138F81BFDF1" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.9" metricValueMin="4.2" pageId="61" pageNumber="152" unit="mm" value="0.45499999999999996" valueMax="0.49" valueMin="0.42">0.42– 0.49 mm</quantity>
; length of elytra from humeral angles to apex
<quantity id="BF294AE8726B7176FF7C6178F85CFD91" box="[174,329,576,600]" metricMagnitude="-4" metricUnit="m" metricValue="7.649999999999999" metricValueMax="8.299999999999999" metricValueMin="7.0" pageId="61" pageNumber="152" unit="mm" value="0.7649999999999999" valueMax="0.83" valueMin="0.7">0.70–0.83 mm</quantity>
; length of elytral suture
<quantity id="BF294AE8726B7176FDBE6178FA12FD91" box="[620,775,576,600]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="7.199999999999999" metricValueMin="6.1" pageId="61" pageNumber="152" unit="mm" value="0.665" valueMax="0.72" valueMin="0.61">0.61–0.72 mm</quantity>
; width of elytra
<quantity id="BF294AE8726B7176FE9E6158F8F2FDB1" box="[332,487,608,632]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="9.0" metricValueMin="7.5" pageId="61" pageNumber="152" unit="mm" value="0.825" valueMax="0.9" valueMin="0.75">0.75–0.90 mm</quantity>
. Habitus as in
<figureCitation id="E0EAFB88726B7176FDAC6158FBAEFDBE" box="[638,699,608,632]" captionStart="Figs 1–6" captionStartId="56.[212,252,2003,2025]" captionTargetBox="[212,1459,271,1981]" captionTargetId="figure-27@56.[211,1459,254,1994]" captionTargetPageId="56" captionText="Figs 1–6: Type locality ofBellatheta khustupica in the peak region of Mount Khustup (1); microhabitat whereB. khustupica was" figureDoi="http://doi.org/10.5281/zenodo.5742740" httpUri="https://zenodo.org/record/5742740/files/figure.png" pageId="61" pageNumber="152">Fig. 3</figureCitation>
. Coloration: head usually blackish, sometimes brown; pronotum reddish, sometimes with more or less extensive brownish discal area: elytra yellowish red; abdomen brownish with yellowish-brown paratergites; antennae, legs and mouth parts entirely yellowish.
</paragraph>
<paragraph id="786EE70D726B7176FF7C6018F8A8FA9E" blockId="61.[174,785,351,2072]" pageId="61" pageNumber="152">Head moderately transverse, 1.2–1.3 times as broad as long; temporal angles rounded but marked; two ridges extending from above insertion of antennae to compound eyes; surface without microsculpture, with dense and regular punctation forming some irregular striae between eyes and ante-ocellar pits, punctation anteriorly extending to the anterior margin of clypeus; sub-antennal impression rather wide, tentorial pits in front of ocelli long and linear, posteriorly extending to the rather small ocelli, the latter separated by a distance equal to that between ocellus and eye; space between tentorial pits and eyes elevated; neck clearly separated from the rest of the head, with dense punctation similar to that of head. Eyes small, slightly shorter than temples, without infra-orbital carina. Antenna rather thin and elongate, antennomere I elongate, twice as long as wide, II elongate ovoid, III–VIII longer than wide, gradually incrassate, IX–X subquadrate, XI longer than wide.</paragraph>
<paragraph id="786EE70D726B7176FF7C6658FB12F9BE" blockId="61.[174,785,351,2072]" pageId="61" pageNumber="152">Pronotum transverse, on average 1.4 times as broad as long and 1.4 times as broad as head, widest in anterior half; anterior margin wider than posterior margin, anterior and posterior angles marked and slightly obtuse; lateral margins with trace of crenulation, weakly sinuate posteriorly; surface of pronotum irregular with three longitudinal elevations and two pits near lateral margin; punctation dense, similar to that of head, ground surface smooth without microsculpture.</paragraph>
<paragraph id="786EE70D726B7176FF7C65B8F832F8FE" blockId="61.[174,785,351,2072]" pageId="61" pageNumber="152">Elytra small, 1.5 times as long (measured from humeral angles) and 1.3 times as broad as pronotum; punctation irregular and dense, coarser than on pronotum, interstices without microsculpture. Legs simple, with some spines in the apical portions of the meso- and metatibiae.</paragraph>
<paragraph id="786EE70D726B7176FF7C6478FB97F8BE" blockId="61.[174,785,351,2072]" pageId="61" pageNumber="152">Abdomen with isodiametric microsculpture, decumbent pubescence and small wing-folding patches.</paragraph>
<paragraph id="786EE70D726B7176FF1164B8FBF5F81E" blockId="61.[174,785,351,2072]" pageId="61" pageNumber="152">
: aedeagus as in
<figureCitation id="E0EAFB88726B7176FE5164B8F8C8F85E" box="[387,477,1920,1944]" captionStart="Figs 7–23" captionStartId="57.[178,218,1988,2010]" captionTargetBox="[184,1416,256,1980]" captionTargetId="figure-27@57.[177,1417,254,1983]" captionTargetPageId="57" captionText="Figs 7–23:Omalium kociani (7–10), Proteinus baculatus (11–14), Bryaxis armeniacus (15, 17), B. meghruicus (16, 18), Aleochara fugax (19–21), and Atheta meghruica (22–23): (median lobe of) aedeagus in lateral and in ventral view (7–8, 13–16); female accessory sclerites (9); spermatheca (10, 21); habitus (11, 19, 22); thorax in ventral view (12); male antennomeres I–III (17–18); posterior margin of female sternite VIII (20); forebody (23). Scale bars: 11, 19, 22: 1.0 mm; 12, 23: 0.5 mm; 7–10, 13–16, 20–21: 0.1 mm; 17–18: without scale." figureDoi="http://doi.org/10.5281/zenodo.5742742" httpUri="https://zenodo.org/record/5742742/files/figure.png" pageId="61" pageNumber="152">Figs 7–8</figureCitation>
, median lobe with narrow base, strongly enlarged subapically; apex weakly curved dorsad in lateral view and without ventral elevations.
</paragraph>
<paragraph id="786EE70D726B7176FF1164D8F9E9F7DE" blockId="61.[174,785,351,2072]" pageId="61" pageNumber="152">
: accessory sclerites as in
<figureCitation id="E0EAFB88726B7176FE2264D8FB39F83E" box="[496,556,2016,2040]" captionStart="Figs 7–23" captionStartId="57.[178,218,1988,2010]" captionTargetBox="[184,1416,256,1980]" captionTargetId="figure-27@57.[177,1417,254,1983]" captionTargetPageId="57" captionText="Figs 7–23:Omalium kociani (7–10), Proteinus baculatus (11–14), Bryaxis armeniacus (15, 17), B. meghruicus (16, 18), Aleochara fugax (19–21), and Atheta meghruica (22–23): (median lobe of) aedeagus in lateral and in ventral view (7–8, 13–16); female accessory sclerites (9); spermatheca (10, 21); habitus (11, 19, 22); thorax in ventral view (12); male antennomeres I–III (17–18); posterior margin of female sternite VIII (20); forebody (23). Scale bars: 11, 19, 22: 1.0 mm; 12, 23: 0.5 mm; 7–10, 13–16, 20–21: 0.1 mm; 17–18: without scale." figureDoi="http://doi.org/10.5281/zenodo.5742742" httpUri="https://zenodo.org/record/5742742/files/figure.png" pageId="61" pageNumber="152">Fig. 9</figureCitation>
; spermatheca as in
<figureCitation id="E0EAFB88726B7176FF7C6B38F9EDF7DE" box="[174,248,2048,2072]" captionStart="Figs 7–23" captionStartId="57.[178,218,1988,2010]" captionTargetBox="[184,1416,256,1980]" captionTargetId="figure-27@57.[177,1417,254,1983]" captionTargetPageId="57" captionText="Figs 7–23:Omalium kociani (7–10), Proteinus baculatus (11–14), Bryaxis armeniacus (15, 17), B. meghruicus (16, 18), Aleochara fugax (19–21), and Atheta meghruica (22–23): (median lobe of) aedeagus in lateral and in ventral view (7–8, 13–16); female accessory sclerites (9); spermatheca (10, 21); habitus (11, 19, 22); thorax in ventral view (12); male antennomeres I–III (17–18); posterior margin of female sternite VIII (20); forebody (23). Scale bars: 11, 19, 22: 1.0 mm; 12, 23: 0.5 mm; 7–10, 13–16, 20–21: 0.1 mm; 17–18: without scale." figureDoi="http://doi.org/10.5281/zenodo.5742742" httpUri="https://zenodo.org/record/5742742/files/figure.png" pageId="61" pageNumber="152">Fig. 10</figureCitation>
.
</paragraph>
<paragraph id="786EE70D726B7176FCE263C7FAB3FAD1" blockId="61.[816,1428,255,1303]" pageId="61" pageNumber="152">
<emphasis id="4AA53B1F726B7176FCE263C7FD1DFEDE" bold="true" box="[816,1032,255,280]" pageId="61" pageNumber="152">Comparative notes</emphasis>
: The new species belongs to the
<taxonomicName id="BFD19C8E726B7176FCE26218FAFEFEF1" authorityName="GRAVENHORST" authorityYear="1806" box="[816,1003,288,312]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="caesum">
<emphasis id="4AA53B1F726B7176FCE26218FAFEFEF1" box="[816,1003,288,312]" italics="true" pageId="61" pageNumber="152">Omalium caesum</emphasis>
</taxonomicName>
group as defined by
<bibRefCitation id="1C409AFC726B7176FB0A6218FC85FEFE" author="ZANETTI, A." box="[1240,1424,288,312]" pageId="61" pageNumber="152" refId="ref82899" refString="ZANETTI, A. 1987: Fauna d'Italia. Coleoptera Staphylinidae Omaliinae. - Calderini, Bologna: 472 pp." type="book" year="1987">ZANETTI (1987)</bibRefCitation>
: ventral process of the aedeagus proximally much narrower than the basal bulb, parameres distinctly dilated and of intricate structure distally, with very long apical setae, tentorial impressions line-shaped. According to
<bibRefCitation id="1C409AFC726B7176FCE262F8FAF0FE1E" author="ZANETTI, A." box="[816,997,448,472]" pageId="61" pageNumber="152" pagination="45 - 63" refId="ref82967" refString="ZANETTI, A. 2002: Studies on Omalium GRAVENHORST, 1802 from Turkey, Cyprus, and the Caucasus region, with notes on some European and Asian species (Coleoptera, Staphylinidae: Omaliinae). - Bollettino del Museo Civico di Storia Naturale di Verona. Botanica Zoologia 26: 45 - 63." type="journal article" year="2002">ZANETTI (2002)</bibRefCitation>
, the following representatives of the
<taxonomicName id="BFD19C8E726B7176FCE262D8FA89FE31" authorityName="GRAVENHORST" authorityYear="1806" box="[816,924,480,503]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="caesum">
<emphasis id="4AA53B1F726B7176FCE262D8FA89FE31" box="[816,924,480,503]" italics="true" pageId="61" pageNumber="152">O. caesum</emphasis>
</taxonomicName>
group are distributed in
<collectingCountry id="00C6A79D726B7176FB6462D8FC14FE3E" box="[1206,1281,480,504]" name="Turkey" pageId="61" pageNumber="152">Turkey</collectingCountry>
,
<collectingCountry id="00C6A79D726B7176FADC62D8FC4EFE3E" box="[1294,1371,480,504]" name="Cyprus" pageId="61" pageNumber="152">Cyprus</collectingCountry>
and the Caucasus region:
<taxonomicName id="BFD19C8E726B7176FBF86138FC9AFDDE" authority="GRAVENHORST, 1806" authorityName="GRAVENHORST" authorityYear="1806" box="[1066,1423,512,536]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="caesum">
<emphasis id="4AA53B1F726B7176FBF86138FD80FDD1" box="[1066,1173,512,535]" italics="true" pageId="61" pageNumber="152">O. caesum</emphasis>
GRAVENHORST, 1806
</taxonomicName>
,
<taxonomicName id="BFD19C8E726B7176FCE26118FD59FDFE" authority="KRAATZ, 1857" authorityName="KRAATZ" authorityYear="1857" box="[816,1100,544,568]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="littorale">
<emphasis id="4AA53B1F726B7176FCE26118FAB5FDFE" box="[816,928,544,568]" italics="true" pageId="61" pageNumber="152">O. littorale</emphasis>
KRAATZ, 1857
</taxonomicName>
,
<taxonomicName id="BFD19C8E726B7176FB866118FC9AFDFE" authority="ZANETTI, 2002" authorityName="ZANETTI" authorityYear="2002" box="[1108,1423,544,568]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="wunderlei">
<emphasis id="4AA53B1F726B7176FB866118FDCFFDFE" box="[1108,1242,544,568]" italics="true" pageId="61" pageNumber="152">O. wunderlei</emphasis>
<bibRefCitation id="1C409AFC726B7176FB366118FC9AFDFE" author="ZANETTI, A." box="[1252,1423,544,568]" pageId="61" pageNumber="152" pagination="45 - 63" refId="ref82967" refString="ZANETTI, A. 2002: Studies on Omalium GRAVENHORST, 1802 from Turkey, Cyprus, and the Caucasus region, with notes on some European and Asian species (Coleoptera, Staphylinidae: Omaliinae). - Bollettino del Museo Civico di Storia Naturale di Verona. Botanica Zoologia 26: 45 - 63." type="journal article" year="2002">ZANETTI, 2002</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="BFD19C8E726B7176FCB66178FDFAFD9E" authority="MULSANT &amp; REY, 1880" authorityName="MULSANT &amp; REY" authorityYear="1880" box="[868,1263,576,600]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="rugatum">
<emphasis id="4AA53B1F726B7176FCB66178FAC8FD91" box="[868,989,576,599]" italics="true" pageId="61" pageNumber="152">O. rugatum</emphasis>
MULSANT &amp; REY, 1880
</taxonomicName>
). They are all clearly distinguished from
<taxonomicName id="BFD19C8E726B7176FB956158FDA4FDBE" authorityName="Assing &amp; Schülke" authorityYear="2019" box="[1095,1201,608,632]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="kociani">
<emphasis id="4AA53B1F726B7176FB956158FDA4FDBE" box="[1095,1201,608,632]" italics="true" pageId="61" pageNumber="152">O. kociani</emphasis>
</taxonomicName>
by long elytra, rather uniform and dark colour, and by the shape of the median lobe of the aedeagus (ventral process not narrowed at the base).
<taxonomicName id="BFD19C8E726B7176FCA461F8FDF8FD1E" authority="SMETANA, 1867" authorityName="SMETANA" authorityYear="1867" box="[886,1261,704,728]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="turcicum">
<emphasis id="4AA53B1F726B7176FCA461F8FD2AFD11" box="[886,1087,704,728]" italics="true" pageId="61" pageNumber="152">Omalium turcicum</emphasis>
SMETANA, 1867
</taxonomicName>
is very similar in coloration and in general shape, but differs by wider elytra, smaller eyes, more rounded temples and confluent punctation of the head, and an impunctate clypeus. However, the only reliable distinguishing character is the shape of the aedeagus: in
<taxonomicName id="BFD19C8E726B7176FBEC6058FDAFFCB1" authorityName="SMETANA" authorityYear="1867" box="[1086,1210,864,887]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="turcicum">
<emphasis id="4AA53B1F726B7176FBEC6058FDAFFCB1" box="[1086,1210,864,887]" italics="true" pageId="61" pageNumber="152">O. turcicum</emphasis>
</taxonomicName>
, the ventral process is gradually narrowed from the base to the obtuse apex (
<bibRefCitation id="1C409AFC726B7176FCE46098FAC0FC7E" author="ZANETTI, A." box="[822,981,928,952]" pageId="61" pageNumber="152" pagination="45 - 63" refId="ref82967" refString="ZANETTI, A. 2002: Studies on Omalium GRAVENHORST, 1802 from Turkey, Cyprus, and the Caucasus region, with notes on some European and Asian species (Coleoptera, Staphylinidae: Omaliinae). - Bollettino del Museo Civico di Storia Naturale di Verona. Botanica Zoologia 26: 45 - 63." type="journal article" year="2002">ZANETTI 2002</bibRefCitation>
: figure 58), whereas in
<taxonomicName id="BFD19C8E726B7176FB216098FC48FC7E" authorityName="Assing &amp; Schülke" authorityYear="2019" box="[1267,1373,928,952]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="kociani">
<emphasis id="4AA53B1F726B7176FB216098FC48FC7E" box="[1267,1373,928,952]" italics="true" pageId="61" pageNumber="152">O. kociani</emphasis>
</taxonomicName>
the ventral process is narrow at the base and enlarged in the middle, and the apex is more acute (
<figureCitation id="E0EAFB88726B7176FB7160D8FDF4FC3E" box="[1187,1249,992,1016]" captionStart="Figs 7–23" captionStartId="57.[178,218,1988,2010]" captionTargetBox="[184,1416,256,1980]" captionTargetId="figure-27@57.[177,1417,254,1983]" captionTargetPageId="57" captionText="Figs 7–23:Omalium kociani (7–10), Proteinus baculatus (11–14), Bryaxis armeniacus (15, 17), B. meghruicus (16, 18), Aleochara fugax (19–21), and Atheta meghruica (22–23): (median lobe of) aedeagus in lateral and in ventral view (7–8, 13–16); female accessory sclerites (9); spermatheca (10, 21); habitus (11, 19, 22); thorax in ventral view (12); male antennomeres I–III (17–18); posterior margin of female sternite VIII (20); forebody (23). Scale bars: 11, 19, 22: 1.0 mm; 12, 23: 0.5 mm; 7–10, 13–16, 20–21: 0.1 mm; 17–18: without scale." figureDoi="http://doi.org/10.5281/zenodo.5742742" httpUri="https://zenodo.org/record/5742742/files/figure.png" pageId="61" pageNumber="152">Fig. 8</figureCitation>
). In populations of
<taxonomicName id="BFD19C8E726B7176FC9F6738FAADFBD1" authorityName="GRAVENHORST" authorityYear="1806" box="[845,952,1024,1047]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="caesum">
<emphasis id="4AA53B1F726B7176FC9F6738FAADFBD1" box="[845,952,1024,1047]" italics="true" pageId="61" pageNumber="152">O. caesum</emphasis>
</taxonomicName>
with short elytra from the Alps, the ventral process of the aedeagus has a broad base (
<bibRefCitation id="1C409AFC726B7176FB396718FC9FFBFE" author="ZANETTI, A." box="[1259,1418,1056,1080]" pageId="61" pageNumber="152" pagination="45 - 63" refId="ref82967" refString="ZANETTI, A. 2002: Studies on Omalium GRAVENHORST, 1802 from Turkey, Cyprus, and the Caucasus region, with notes on some European and Asian species (Coleoptera, Staphylinidae: Omaliinae). - Bollettino del Museo Civico di Storia Naturale di Verona. Botanica Zoologia 26: 45 - 63." type="journal article" year="2002">ZANETTI 2002</bibRefCitation>
: figure 55).
<taxonomicName id="BFD19C8E726B7176FC796778FC0BFB9E" authority="FAUVEL, 1900" authorityName="FAUVEL, Am" authorityYear="1900" box="[939,1310,1088,1112]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="cribriceps">
<emphasis id="4AA53B1F726B7176FC796778FD6AFB9E" box="[939,1151,1088,1112]" italics="true" pageId="61" pageNumber="152">Omalium cribriceps</emphasis>
FAUVEL, 1900
</taxonomicName>
, which is distributed in
<collectingCountry id="00C6A79D726B7176FC196758FAEFFBB1" box="[971,1018,1120,1143]" name="Iran" pageId="61" pageNumber="152">Iran</collectingCountry>
,
<collectingCountry id="00C6A79D726B7176FBD46758FD21FBB1" box="[1030,1076,1120,1143]" name="Iraq" pageId="61" pageNumber="152">Iraq</collectingCountry>
,
<collectingCountry id="00C6A79D726B7176FBED6758FD62FBBE" box="[1087,1143,1120,1144]" name="Syria" pageId="61" pageNumber="152">Syria</collectingCountry>
, and
<collectingCountry id="00C6A79D726B7176FB616758FDEBFBBE" box="[1203,1278,1120,1144]" name="Turkey" pageId="61" pageNumber="152">Turkey</collectingCountry>
(
<bibRefCitation id="1C409AFC726B7176FADE6764FAC3FB5E" author="SCHULKE, M. &amp; SMETANA, A." pageId="61" pageNumber="152" pagination="304 - 1134" refId="ref82321" refString="SCHULKE, M. &amp; SMETANA, A. 2015: Staphylinidae, pp. 304 - 1134. - In: LOBL, I. &amp; LOBL, D. (eds), Catalogue of Palaearctic Coleoptera. New updated Edition. Volume 2. Hydrophiloidea - Staphylinoidea. Revised and updated edition. - Brill, Leiden: xxvi + 1702 pp." type="book chapter" year="2015">SCHÜLKE &amp; SMETANA 2015</bibRefCitation>
), is very similar to
<taxonomicName id="BFD19C8E726B7176FB7D67B8FC0CFB5E" authorityName="Assing &amp; Schülke" authorityYear="2019" box="[1199,1305,1152,1176]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="152" phylum="Arthropoda" rank="species" species="kociani">
<emphasis id="4AA53B1F726B7176FB7D67B8FC0CFB5E" box="[1199,1305,1152,1176]" italics="true" pageId="61" pageNumber="152">O. kociani</emphasis>
</taxonomicName>
in coloration, habitus, and the punctation of the head, but has an aedeagus of completely different shape (
<bibRefCitation id="1C409AFC726B7176FB3A67F8FC85FB1E" author="ZANETTI, A." box="[1256,1424,1216,1240]" pageId="61" pageNumber="152" pagination="45 - 63" refId="ref82967" refString="ZANETTI, A. 2002: Studies on Omalium GRAVENHORST, 1802 from Turkey, Cyprus, and the Caucasus region, with notes on some European and Asian species (Coleoptera, Staphylinidae: Omaliinae). - Bollettino del Museo Civico di Storia Naturale di Verona. Botanica Zoologia 26: 45 - 63." type="journal article" year="2002">ZANETTI 2002</bibRefCitation>
: figure 48) with wide median lobe and simple, linear parameres.
</paragraph>
</subSubSection>
<subSubSection id="30CBB486726B7176FCE26607FC00FA7E" pageId="61" pageNumber="152" type="distribution">
<paragraph id="786EE70D726B7176FCE26607FC00FA7E" blockId="61.[816,1427,1343,1464]" pageId="61" pageNumber="152">
<emphasis id="4AA53B1F726B7176FCE26607FD80FA91" bold="true" box="[816,1173,1343,1368]" pageId="61" pageNumber="152">Distribution and natural history</emphasis>
: This species is known only from the
<typeStatus id="A76A59AF726B7176FC156658FAE0FABE" box="[967,1013,1376,1400]" pageId="61" pageNumber="152">type</typeStatus>
locality in
<collectingCountry id="00C6A79D726B7176FBB86658FDD9FABE" box="[1130,1228,1376,1400]" name="Armenia" pageId="61" pageNumber="152">Armenia</collectingCountry>
. The typical series was collected in an alpine meadow by sifting humus and rotting herbs near a small residual snow field.
</paragraph>
</subSubSection>
</treatment>
</document>
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