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<document id="A5E8AAC74011A004231A07D08D9D0A77" ID-CLB-Dataset="276296" ID-DOI="10.1093/zoolinnean/zlaa057" ID-GBIF-Dataset="c569b071-a44d-4681-8a63-3d54b9bcfac2" ID-ISSN="0024-4082" ID-Zenodo-Dep="5300077" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630095302060" checkinUser="felipe" docAuthor="Haase, Martin, Meng, Stefan &amp; Horsák, Michal" docDate="2021" docId="B35287F0FF91953FFCE3F88EFBC623CF" docLanguage="en" docName="zlaa057.pdf" docOrigin="Zoological Journal of the Linnean Society 191 (3)" docSource="https://academic.oup.com/zoolinnean/article/191/3/720/5861197" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pupilla loessica Lozek 1954" docType="treatment" docVersion="6" masterDocId="4F6BFF88FF81952CFFD8FF9BFFF32336" masterDocTitle="Tracking parallel adaptation of shell morphology through geological times in the land snail genus Pupilla (Gastropoda: Stylommatophora: Pupillidae)" masterLastPageNumber="720" masterPageNumber="720" pageNumber="720" updateTime="1699150466301" updateUser="ExternalLinkService">
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<mods:title id="30AD8128CE5CA45D46A113021B42B33C">Tracking parallel adaptation of shell morphology through geological times in the land snail genus Pupilla (Gastropoda: Stylommatophora: Pupillidae)</mods:title>
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<treatment id="B35287F0FF91953FFCE3F88EFBC623CF" ID-GBIF-Taxon="217229020" LSID="urn:lsid:plazi:treatment:B35287F0FF91953FFCE3F88EFBC623CF" httpUri="http://treatment.plazi.org/id/B35287F0FF91953FFCE3F88EFBC623CF" lastPageId="19" pageId="16">
<subSubSection id="73E1656DFF91953CFCE3F88EFB61241B" box="[827,1170,1813,1837]" pageId="16" type="nomenclature">
<paragraph id="3B4436E6FF91953CFCE3F88EFB61241B" blockId="16.[827,1170,1813,1837]" box="[827,1170,1813,1837]" pageId="16">
<heading id="600C818AFF91953CFCE3F88EFB61241B" box="[827,1170,1813,1837]" centered="true" fontSize="9" level="2" pageId="16" reason="2">
<taxonomicName id="FCFB4D65FF91953CFCE3F88EFB61241B" ID-CoL="7SWS4" authority="Lozek, 1954" authorityName="Lozek" authorityYear="1954" box="[827,1170,1813,1837]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="16" phylum="Mollusca" rank="species" species="loessica">
<emphasis id="098FEAF4FF91953CFCE3F88EFB61241B" box="[827,1170,1813,1837]" italics="true" pageId="16">
Pupilla loessica
<bibRefCitation id="5F6A4B17FF91953CFBDBF88EFB61241B" author="Lozek V" box="[1027,1170,1813,1837]" pageId="16" pagination="327 - 342" refId="ref18470" refString="Lozek V. 1954. Novi mekkysi Ceskoslovenskho pleistocenu Vertigo pseudosubstriata sp. n., Pupilla muscorum densegyrata ssp. n. a Pupilla loessica sp. n. (Neue Mollusken aus dem tschechoslowakischen Pleistocen). Anthropozoikum 3: 327 - 342." type="journal article" year="1954">Ložek, 1954</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="73E1656DFF91953FFCE3F8A5FBC623CF" lastPageId="19" pageId="16" type="materials_examined">
<paragraph id="3B4436E6FF91953FFCE3F8A5FECE265E" blockId="16.[827,1442,1854,1906]" lastBlockId="19.[145,761,197,1630]" lastPageId="19" pageId="16">
The type locality of
<taxonomicName id="FCFB4D65FF91953CFBF5F8A5FADE2462" authority=", Predmosti" authorityName="Predmosti" box="[1069,1325,1854,1876]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="16" phylum="Mollusca" rank="species" species="loessica">
<emphasis id="098FEAF4FF91953CFBF5F8A5FB542465" box="[1069,1191,1854,1875]" italics="true" pageId="16">P. loessica</emphasis>
, Předmostí
</taxonomicName>
at Přerov in
<collectingRegion id="F93FF804FF91953CFC87F8C7FC352444" box="[863,966,1884,1906]" country="Czech Republic" name="Jihomoravsky" pageId="16">Moravia</collectingRegion>
(
<collectingCountry id="43EC7676FF91953CFC02F8C7FB572444" box="[986,1188,1884,1906]" name="Czech Republic" pageId="16">Czech Republic</collectingCountry>
), is a world-famous archaeological site. In loess sediments, mined for brick production in the past, a Gravettian-temporal cultural layer was excavated (e.g.
<bibRefCitation id="5F6A4B17FF90953DFE68F8DDFD77246B" author="Zebera K &amp; Lozek V &amp; Kneblova V &amp; Fejfar O &amp; Mazalek M" box="[432,644,1862,1886]" pageId="17" pagination="291 - 362" refId="ref20101" refString="Zebera K, Lozek V, Kneblova V, Fejfar O, Mazalek M. 1955. Zprava o II. etape geologickeho vyzkumu kvarteru v Predmosti u Prerova na Morave. Anthropozoikum 4: 291 - 362." type="journal article" year="1955">
Žebera
<emphasis id="098FEAF4FF90953DFDDEF8D3FDCC246A" box="[518,575,1863,1885]" italics="true" pageId="17">et al.</emphasis>
, 1955
</bibRefCitation>
). Because the type material of
<taxonomicName id="FCFB4D65FF90953DFBFDF891FB6D2429" box="[1061,1182,1802,1823]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="loessica">
<emphasis id="098FEAF4FF90953DFBFDF891FB6D2429" box="[1061,1182,1802,1823]" italics="true" pageId="17">P. loessica</emphasis>
</taxonomicName>
has been destroyed (
<collectionCode id="5DEAAE23FF90953DFCE8F8B2FCB72408" box="[816,836,1833,1854]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="17" type="Museum">V</collectionCode>
. Ložek and curator E. Kadlecová, pers. comm.), a
<typeStatus id="E4408844FF90953DFCE7F8D3FC6F246B" box="[831,924,1864,1885]" pageId="17" type="neotype">neotype</typeStatus>
should be defined in order to anchor this essential Pleistocene indicator species and preclude taxonomic confusion of this morphologically and ecologically variable and (historically) widespread species. For differentiating features see the
<emphasis id="098FEAF4FF92953FFD4BFEBAFD0B2200" box="[659,760,289,310]" italics="true" pageId="19">Material</emphasis>
section above and our present comparisons. Today, the excavation site virtually does not exist anymore and the important geological profiles including Ložeks profile in the former western mine and Kneblovás profile in the former northern mine near HradiŠtko [according to
<bibRefCitation id="5F6A4B17FF92953FFE98FE43FDCF22D9" author="Zebera K &amp; Lozek V &amp; Kneblova V &amp; Fejfar O &amp; Mazalek M" box="[320,572,472,496]" pageId="19" pagination="291 - 362" refId="ref20101" refString="Zebera K, Lozek V, Kneblova V, Fejfar O, Mazalek M. 1955. Zprava o II. etape geologickeho vyzkumu kvarteru v Predmosti u Prerova na Morave. Anthropozoikum 4: 291 - 362." type="journal article" year="1955">
Žebera
<emphasis id="098FEAF4FF92953FFE79FE41FE2A22D8" box="[417,473,473,495]" italics="true" pageId="19">et al</emphasis>
. (1955)
</bibRefCitation>
] are no longer accessible. According to
<bibRefCitation id="5F6A4B17FF92953FFE72FE63FDB22138" author="Lozek V" box="[426,577,504,526]" pageId="19" pagination="327 - 342" refId="ref18470" refString="Lozek V. 1954. Novi mekkysi Ceskoslovenskho pleistocenu Vertigo pseudosubstriata sp. n., Pupilla muscorum densegyrata ssp. n. a Pupilla loessica sp. n. (Neue Mollusken aus dem tschechoslowakischen Pleistocen). Anthropozoikum 3: 327 - 342." type="journal article" year="1954">Ložek (1954)</bibRefCitation>
, the type series came directly from the HradiŠtko limestone hill from the younger loess (Würmian
<collectionCode id="5DEAAE23FF92953FFE34FDADFDF8217D" box="[492,523,566,587]" pageId="19">III</collectionCode>
or Weichselian
<collectionCode id="5DEAAE23FF92953FFD12FDADFD18217D" box="[714,747,566,587]" pageId="19">III</collectionCode>
). In the vicinity of the plateau, there are still loess outcrops, however, consisting only of an uppermost layer of loess, presumably from the high cold period of the
<collectionCode id="5DEAAE23FF92953FFF06FD2BFEE921F3" box="[222,282,688,709]" country="Russia" httpUri="http://grbio.org/cool/z8zf-y76s" name="Leningrad School of Mines" pageId="19">LGM</collectionCode>
of the Weichselian glacial. This layer is sterile, virtually without any mollusc shells (S. Meng, pers. obs. based on processing a 10 L sample taken during a visit on
<date id="4F451026FF92953FFEAAFC97FD9A2014" box="[370,617,780,802]" pageId="19" value="2019-09-13">13 September 2019</date>
). The layer where Ložek found the original material of
<taxonomicName id="FCFB4D65FF92953FFD5EFCB0FD0B2009" box="[646,760,810,832]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="19" phylum="Mollusca" rank="species" species="loessica">
<emphasis id="098FEAF4FF92953FFD5EFCB0FD0B2009" box="[646,760,810,832]" italics="true" pageId="19">P. loessica</emphasis>
</taxonomicName>
must have been beneath the examined uppermost layer of loess. Because HradiŠtko is an important cultural and natural monument, further excavations are prohibited. However, in Ložeks collection in the National Museum of
<collectingRegion id="F93FF804FF92953FFE51FC5FFE2D20EF" box="[393,478,964,985]" country="Czech Republic" name="Praha" pageId="19">Prague</collectingRegion>
, two series of
<taxonomicName id="FCFB4D65FF92953FFD5CFC5FFD0A20EF" box="[644,761,964,985]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="19" phylum="Mollusca" rank="species" species="loessica">
<emphasis id="098FEAF4FF92953FFD5CFC5FFD0A20EF" box="[644,761,964,985]" italics="true" pageId="19">P. loessica</emphasis>
</taxonomicName>
also collected from the profile Kneblová in 1955 are present. To the best of our knowledge, this is the only material from the type locality that has not been lost. It was taken from a depth of 9.511.0 m, thus it is much older than the lost type material and it probably originates from the Rissian or Saalian glacial complex (e.g.
<bibRefCitation id="5F6A4B17FF92953FFE40FB02FD722786" author="Zebera K &amp; Lozek V &amp; Kneblova V &amp; Fejfar O &amp; Mazalek M" box="[408,641,1177,1201]" pageId="19" pagination="291 - 362" refId="ref20101" refString="Zebera K, Lozek V, Kneblova V, Fejfar O, Mazalek M. 1955. Zprava o II. etape geologickeho vyzkumu kvarteru v Predmosti u Prerova na Morave. Anthropozoikum 4: 291 - 362." type="journal article" year="1955">
Žebera
<emphasis id="098FEAF4FF92953FFE2EFB00FDC62799" box="[502,565,1178,1200]" italics="true" pageId="19">et al.</emphasis>
, 1955
</bibRefCitation>
;
<bibRefCitation id="5F6A4B17FF92953FFD4BFB01FEE327F8" author="Svoboda J &amp; Lozek V &amp; Svobodova H &amp; Skrdla P" pageId="19" pagination="457 - 472" refId="ref19597" refString="Svoboda J, Lozek V, Svobodova H, Skrdla P. 1994. Predmosti after 110 years. Journal of Field Archaeology 21: 457 - 472." type="journal article" year="1994">
Svoboda
<emphasis id="098FEAF4FF92953FFF49FB21FF3927F8" box="[145,202,1209,1231]" italics="true" pageId="19">et al.</emphasis>
, 1994
</bibRefCitation>
). However, as the profile Kneblová is only a few hundred meters away from the original site and the material was more or less “approved” by the author, we selected the
<typeStatus id="E4408844FF92953FFE6DFA8EFDE0261C" box="[437,531,1301,1322]" pageId="19" type="neotype">neotype</typeStatus>
of
<taxonomicName id="FCFB4D65FF92953FFDE4FA8DFD44261C" box="[572,695,1301,1323]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="19" phylum="Mollusca" rank="species" species="loessica">
<emphasis id="098FEAF4FF92953FFDE4FA8DFD44261C" box="[572,695,1301,1323]" italics="true" pageId="19">P. loessica</emphasis>
</taxonomicName>
from this series of a total of four shells (
<figureCitation id="A3C02A63FF92953FFDEBFAAFFD77267F" box="[563,644,1332,1354]" captionStart="Figure 10" captionStartId="22.[164,242,1301,1323]" captionTargetBox="[163,1443,195,1261]" captionTargetId="figure-292@22.[163,1443,195,1261]" captionTargetPageId="22" captionText="Figure 10. Pupilla loessica. Series of four shells from type locality Předmostí at Přerov (Czech Republic, fossil from Saalian loess) in frontal and lateral view. A is the designated neotype. All deposited at the National Museum of Prague." figureDoi="http://doi.org/10.5281/zenodo.5496070" httpUri="https://zenodo.org/record/5496070/files/figure.png" pageId="19">Fig. 10</figureCitation>
) with the following data:
</paragraph>
<caption id="6F84666EFF90953DFF49F9FEFDB825E5" ID-DOI="http://doi.org/10.5281/zenodo.5300089" ID-Zenodo-Dep="5300089" httpUri="https://zenodo.org/record/5300089/files/figure.png" pageId="17" startId="17.[145,226,1637,1659]" targetBox="[145,1425,209,1597]" targetPageId="17">
<paragraph id="3B4436E6FF90953DFF49F9FEFDB825E5" blockId="17.[145,1426,1637,1747]" pageId="17">
<emphasis id="098FEAF4FF90953DFF49F9FEFF0D254C" bold="true" box="[145,254,1637,1659]" pageId="17">Figure 6.</emphasis>
Thin-plate splines illustrating (theoretical) transitions in shape, side view, exaggeration 5 times. A, fossil
<taxonomicName id="FCFB4D65FF90953DFF49F918FF0B25AE" box="[145,248,1666,1688]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="loessica">
<emphasis id="098FEAF4FF90953DFF49F918FF0B25AE" box="[145,248,1666,1688]" italics="true" pageId="17">P. loessica</emphasis>
</taxonomicName>
→ extant
<taxonomicName id="FCFB4D65FF90953DFEBBF918FE3825AE" box="[355,459,1666,1688]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="loessica">
<emphasis id="098FEAF4FF90953DFEBBF918FE3825AE" box="[355,459,1666,1688]" italics="true" pageId="17">P. loessica</emphasis>
</taxonomicName>
. B, fossil
<taxonomicName id="FCFB4D65FF90953DFDF4F918FD5025AE" baseAuthorityName="Clessin" baseAuthorityYear="1871" box="[556,675,1666,1688]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="pratensis">
<emphasis id="098FEAF4FF90953DFDF4F918FD5025AE" box="[556,675,1666,1688]" italics="true" pageId="17">P. pratensis</emphasis>
</taxonomicName>
(including
<emphasis id="098FEAF4FF90953DFCC0F918FC3F25AE" box="[792,972,1666,1688]" italics="true" pageId="17">P. m. densegyrata</emphasis>
) → extant
<taxonomicName id="FCFB4D65FF90953DFBE7F919FB4825AE" baseAuthorityName="Clessin" baseAuthorityYear="1871" box="[1087,1211,1666,1688]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="pratensis">P. pratensis</taxonomicName>
. C, fossil
<taxonomicName id="FCFB4D65FF90953DFAC2F918FA6225AE" baseAuthorityName="Clessin" baseAuthorityYear="1871" box="[1306,1425,1666,1688]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="pratensis">
<emphasis id="098FEAF4FF90953DFAC2F918FA6225AE" box="[1306,1425,1666,1688]" italics="true" pageId="17">P. pratensis</emphasis>
</taxonomicName>
(including
<emphasis id="098FEAF4FF90953DFED9F93BFE452580" box="[257,438,1696,1718]" italics="true" pageId="17">P. m. densegyrata</emphasis>
) → extant
<taxonomicName id="FCFB4D65FF90953DFDF4F93BFD642580" box="[556,663,1696,1718]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="alpicola">
<emphasis id="098FEAF4FF90953DFDF4F93BFD642580" box="[556,663,1696,1718]" italics="true" pageId="17">P. alpicola</emphasis>
</taxonomicName>
. D, extant
<taxonomicName id="FCFB4D65FF90953DFCD2F93BFC712580" baseAuthorityName="Clessin" baseAuthorityYear="1871" box="[778,898,1696,1718]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="pratensis">
<emphasis id="098FEAF4FF90953DFCD2F93BFC712580" box="[778,898,1696,1718]" italics="true" pageId="17">P. pratensis</emphasis>
</taxonomicName>
<taxonomicName id="FCFB4D65FF90953DFC7DF93BFBE32580" box="[933,1040,1696,1718]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="alpicola">
<emphasis id="098FEAF4FF90953DFC7DF93BFBE32580" box="[933,1040,1696,1718]" italics="true" pageId="17">P. alpicola</emphasis>
</taxonomicName>
. E, extant
<taxonomicName id="FCFB4D65FF90953DFB5BF93BFB182580" box="[1155,1259,1696,1718]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="loessica">
<emphasis id="098FEAF4FF90953DFB5BF93BFB182580" box="[1155,1259,1696,1718]" italics="true" pageId="17">P. loessica</emphasis>
</taxonomicName>
<taxonomicName id="FCFB4D65FF90953DFAD0F93BFA802580" box="[1288,1395,1696,1718]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="alpicola">
<emphasis id="098FEAF4FF90953DFAD0F93BFA802580" box="[1288,1395,1696,1718]" italics="true" pageId="17">P. alpicola</emphasis>
</taxonomicName>
. F, fossil
<taxonomicName id="FCFB4D65FF90953DFF15F925FEA025E5" box="[205,339,1726,1747]" class="Gastropoda" family="Pupillidae" genus="Pupuilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="muscorum">
<emphasis id="098FEAF4FF90953DFF15F925FEA025E5" box="[205,339,1726,1747]" italics="true" pageId="17">P. muscorum</emphasis>
</taxonomicName>
→ extant
<taxonomicName id="FCFB4D65FF90953DFE18F925FDB525E5" box="[448,582,1726,1747]" class="Gastropoda" family="Pupillidae" genus="Pupuilla" kingdom="Animalia" order="Stylommatophora" pageId="17" phylum="Mollusca" rank="species" species="muscorum">
<emphasis id="098FEAF4FF90953DFE18F925FDB525E5" box="[448,582,1726,1747]" italics="true" pageId="17">P. muscorum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="6F84666EFF93953EFF7BF949FA50246B" ID-DOI="http://doi.org/10.5281/zenodo.5496064" ID-Zenodo-Dep="5496064" httpUri="https://zenodo.org/record/5496064/files/figure.png" pageId="18" startId="18.[163,244,1746,1768]" targetBox="[322,1283,197,1705]" targetPageId="18">
<paragraph id="3B4436E6FF93953EFF7BF949FA50246B" blockId="18.[163,1444,1746,1885]" pageId="18">
<emphasis id="098FEAF4FF93953EFF7BF949FEE325D1" bold="true" box="[163,272,1746,1768]" pageId="18">Figure 7.</emphasis>
Shell microsculpture (SEM micrographs). AT,
<taxonomicName id="FCFB4D65FF93953EFCFBF948FC6225DE" box="[803,913,1746,1768]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="18" phylum="Mollusca" rank="species" species="loessica">
<emphasis id="098FEAF4FF93953EFCFBF948FC6225DE" box="[803,913,1746,1768]" italics="true" pageId="18">P. loessica</emphasis>
</taxonomicName>
. UZ,
<taxonomicName id="FCFB4D65FF93953EFC03F948FBBF25DE" box="[987,1100,1746,1768]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="18" phylum="Mollusca" rank="species" species="alpicola">
<emphasis id="098FEAF4FF93953EFC03F948FBBF25DE" box="[987,1100,1746,1768]" italics="true" pageId="18">P. alpicola</emphasis>
</taxonomicName>
. ad,
<taxonomicName id="FCFB4D65FF93953EFB49F948FAF225DE" box="[1169,1281,1746,1768]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="18" phylum="Mollusca" rank="species" species="alpicola">
<emphasis id="098FEAF4FF93953EFB49F948FAF225DE" box="[1169,1281,1746,1768]" italics="true" pageId="18">P. alpicola</emphasis>
</taxonomicName>
(morphogroup
<emphasis id="098FEAF4FF93953EFF7BF96BFEBD2433" box="[163,334,1775,1797]" italics="true" pageId="18">P.m. densegyrata</emphasis>
). eh,
<taxonomicName id="FCFB4D65FF93953EFE56F96BFE072433" box="[398,500,1775,1797]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="18" phylum="Mollusca" rank="species" species="alpicola">
<emphasis id="098FEAF4FF93953EFE56F96BFE072433" box="[398,500,1775,1797]" italics="true" pageId="18">P. alpicola</emphasis>
</taxonomicName>
(lowland populations “
<taxonomicName id="FCFB4D65FF93953EFD0EF96BFCB42433" baseAuthorityName="Clessin" baseAuthorityYear="1871" box="[726,839,1775,1797]" class="Gastropoda" family="Pupillidae" genus="Pupilla" kingdom="Animalia" order="Stylommatophora" pageId="18" phylum="Mollusca" rank="species" species="pratensis">
<emphasis id="098FEAF4FF93953EFD0EF96BFCB42433" box="[726,839,1775,1797]" italics="true" pageId="18">P.pratensis</emphasis>
</taxonomicName>
”). ij,
<taxonomicName id="FCFB4D65FF93953EFC5BF96BFBF02433" box="[899,1027,1776,1797]" class="Gastropoda" family="Pupillidae" genus="Pupuilla" kingdom="Animalia" order="Stylommatophora" pageId="18" phylum="Mollusca" rank="species" species="muscorum">
<emphasis id="098FEAF4FF93953EFC5BF96BFBF02433" box="[899,1027,1776,1797]" italics="true" pageId="18">P.muscorum</emphasis>
</taxonomicName>
. AB, M_4559, Altai, Saylyugem (Russia). CD, M_4575, Altai, Saylyugem (Russia). EF, H_MC409, Altai, Dzhazator (Russia). GH, M_3970, Khatgal, shore of Lake Khövsgöl Nuur (northern Mongolia). IJ, M_2523_2, Yelantsy near Lake Baikal (Russia). KL, M_2523_1, Yelantsy near Lake Baikal (Russia). MN, M_994, Karsdorf (Saxony Anhalt, Germany), fossil from Early Saalian. OP, M_459, Zeuchfeld (Saxony
</paragraph>
</caption>
<paragraph id="3B4436E6FF92953FFF71FAEAFE982690" blockId="19.[145,761,197,1630]" pageId="19">
Ložek collection, National Museum
<collectingRegion id="F93FF804FF92953FFDE7FAEAFD6226B0" box="[575,657,1393,1414]" country="Czech Republic" name="Praha" pageId="19">Prague</collectingRegion>
(without collection number).
</paragraph>
<paragraph id="3B4436E6FF92953FFF71FA34FD892534" blockId="19.[145,761,197,1630]" pageId="19">
<materialsCitation id="8B933CBBFF92953FFF71FA34FD852534" ID-GBIF-Occurrence="4435229301" country="Czech Republic" county="Kneblova" latitude="49.467224" location="Predmosti" longLatPrecision="1" longitude="17.437971" municipality="Prerov" pageId="19" specimenCount="1" stateProvince="Jihomoravsky">
New
<typeStatus id="E4408844FF92953FFF3EFA34FEEA26F2" box="[230,281,1455,1476]" pageId="19">type</typeStatus>
locality:
<location id="3E24603DFF92953FFE5EFA34FE0826F2" LSID="urn:lsid:plazi:treatment:B35287F0FF91953FFCE3F88EFBC623CF:3E24603DFF92953FFE5EFA34FE0826F2" box="[390,507,1454,1476]" country="Czech Republic" county="Kneblova" latitude="49.467224" longLatPrecision="1" longitude="17.437971" municipality="Prerov" name="Predmosti" pageId="19" stateProvince="Jihomoravsky">Předmostí</location>
at
<collectingMunicipality id="DB20AC9CFF92953FFDFCFA34FD8326F2" box="[548,624,1454,1476]" pageId="19">Přerov</collectingMunicipality>
in
<collectingRegion id="F93FF804FF92953FFD40FA35FD0A26F2" box="[664,761,1454,1476]" country="Czech Republic" name="Jihomoravsky" pageId="19">Moravia</collectingRegion>
(
<collectingCountry id="43EC7676FF92953FFF40FA56FEAF26D5" box="[152,348,1485,1507]" name="Czech Republic" pageId="19">Czech Republic</collectingCountry>
), profile
<collectingCounty id="D2254E6AFF92953FFE15FA56FDB326D5" box="[461,576,1485,1507]" pageId="19">Kneblová</collectingCounty>
(
<geoCoordinate id="5ECF5021FF92953FFD8AFA56FD0026D4" box="[594,755,1485,1506]" degrees="49" direction="north" minutes="28" orientation="latitude" pageId="19" precision="1" seconds="02.0" value="49.467224">49°2802.0N</geoCoordinate>
,
<geoCoordinate id="5ECF5021FF92953FFF49FA77FED12534" box="[145,290,1516,1538]" degrees="17" direction="east" minutes="26" orientation="longitude" pageId="19" precision="1" seconds="16.7" value="17.437971">17°2616.7E</geoCoordinate>
) in the former northern mine
</materialsCitation>
.
</paragraph>
<paragraph id="3B4436E6FF92953FFF71F991FEF42508" blockId="19.[145,761,197,1630]" pageId="19">
Age and geology: Saalian Complex, loess, depth
<quantity id="FC039B03FF92953FFF49F9B2FEF02508" box="[145,259,1577,1599]" metricMagnitude="0" metricUnit="m" metricValue="9.65" metricValueMax="9.8" metricValueMin="9.5" pageId="19" unit="m" value="9.65" valueMax="9.8" valueMin="9.5">9.59.8 m</quantity>
.
</paragraph>
<paragraph id="3B4436E6FF92953FFF71F9D3FDD4256B" blockId="19.[145,761,197,1630]" box="[169,551,1607,1630]" pageId="19">
Height,
<quantity id="FC039B03FF92953FFEDDF9D3FE99256B" box="[261,362,1608,1629]" metricMagnitude="-3" metricUnit="m" metricValue="2.91" pageId="19" unit="mm" value="2.91">2.91 mm</quantity>
; width
<quantity id="FC039B03FF92953FFE65F9DCFDD0256B" box="[445,547,1607,1629]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="19" unit="mm" value="1.63">1.63 mm</quantity>
.
</paragraph>
<paragraph id="3B4436E6FF92953FFC99FF5EFBC623CF" blockId="19.[809,1425,197,250]" pageId="19">
For further data including the accompanying fauna see
<bibRefCitation id="5F6A4B17FF92953FFC8CFF79FBC223CF" author="Zebera K &amp; Lozek V &amp; Kneblova V &amp; Fejfar O &amp; Mazalek M" box="[852,1073,226,250]" pageId="19" pagination="291 - 362" refId="ref20101" refString="Zebera K, Lozek V, Kneblova V, Fejfar O, Mazalek M. 1955. Zprava o II. etape geologickeho vyzkumu kvarteru v Predmosti u Prerova na Morave. Anthropozoikum 4: 291 - 362." type="journal article" year="1955">
Žebera
<emphasis id="098FEAF4FF92953FFC71FF7FFC2A23CF" box="[937,985,228,249]" italics="true" pageId="19">et al</emphasis>
. (1955)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>