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205 lines
24 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1159.90120" ID-Pensoft-Pub="1313-2970-1159-151" ID-Pensoft-UUID="EE0AEE0FF72A593BB9BCA14345E5298A" ID-ZooBank="DEC4657EFE2445DBB132E916EC4E3A10" ModsDocID="1313-2970-1159-151" checkinTime="1683131567836" checkinUser="pensoft" docAuthor="Wei, Mian, Wang, Shuqiao & Lin, Yucheng" docDate="2023" docId="A92F27A4AA615477BDD0879EAB711980" docLanguage="en" docName="ZooKeys 1159: 151-168" docOrigin="ZooKeys 1159" docPubDate="2023-05-02" docSource="http://dx.doi.org/10.3897/zookeys.1159.90120" docTitle="Luthela asuka Wei & Lin 2023, sp. nov." docType="treatment" docUuid="917ACCF0-7506-496C-8C29-0CF53D0C710D" docUuidSource="ZooBank" docVersion="1" id="EE0AEE0FF72A593BB9BCA14345E5298A" lastPageNumber="151" masterDocId="EE0AEE0FF72A593BB9BCA14345E5298A" masterDocTitle="Systematic notes on three new Luthela (Mesothelae, Heptathelidae) spiders from China, with their descriptions" masterLastPageNumber="168" masterPageNumber="151" pageNumber="151" updateTime="1683131567836" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematic notes on three new Luthela (Mesothelae, Heptathelidae) spiders from China, with their descriptions</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wei, Mian</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7348-8885</mods:nameIdentifier>
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<mods:affiliation>Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wang, Shuqiao</mods:namePart>
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<mods:affiliation>Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lin, Yucheng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5054-0633</mods:nameIdentifier>
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<mods:affiliation>Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
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<mods:nameIdentifier type="email">linyucheng@scu.edu.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-02</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1159</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>151</mods:start>
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<mods:end>168</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1159.90120</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1159.90120</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1159-151</mods:identifier>
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<mods:identifier type="ZooBank">DEC4657EFE2445DBB132E916EC4E3A10</mods:identifier>
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<mods:identifier type="Pensoft-UUID">EE0AEE0FF72A593BB9BCA14345E5298A</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:917ACCF0-7506-496C-8C29-0CF53D0C710D" httpUri="http://treatment.plazi.org/id/A92F27A4AA615477BDD0879EAB711980" lastPageNumber="151" pageId="0" pageNumber="151">
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<subSubSection pageId="0" pageNumber="151" type="nomenclature">
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<paragraph pageId="0" pageNumber="151">
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<taxonomicName LSID="https://zoobank.org/917ACCF0-7506-496C-8C29-0CF53D0C710D" authority="Wei & Lin" authorityName="Wei & Lin" authorityYear="2023" class="Arachnida" family="Heptathelidae" genus="Luthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Luthela asuka" order="Araneae" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="asuka" status="sp. nov.">Luthela asuka Wei & Lin</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="151">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="151" type="description">
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<paragraph pageId="0" pageNumber="151">
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. New species of Luthela A, B, G L. asuka sp. nov. from Longquanyi District, Chengdu C, D, H, I L. beijing sp. nov. from Zizhuyuan Park, Beijing E, F L. kagami sp. nov. from Guihua Township, Pengzhou City A, C, E male habitus, dorsal view B, D, F female habitus, dorsal view G female haibitus, ventral view H living female, dorsal view I burrow, vertical section, with red arrow pointing to the spider. Photographs by Chao Wu (H, I). Scale bars: 5.00 mm." figureDoi="10.3897/zookeys.1159.90120.figure3" httpUri="https://binary.pensoft.net/fig/844934" pageId="0" pageNumber="151">Figs 3A, B, G</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luthela asuka sp. nov. A male left palp bulb, prolateral view B male left palp bulb, ventral view C male left palp bulb, retrolateral view D left cymbium, ventral view E left palpal bulb, apical view F right palpal bulb, apical view G vulva, ventral view H vulva, dorsal view. Green arrows indicate small teeth on conductor. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1159.90120.figure4" httpUri="https://binary.pensoft.net/fig/844935" pageId="0" pageNumber="151">, 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="151" type="type material">
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<paragraph pageId="0" pageNumber="151">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="151">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="151">Holotype</emphasis>
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♂,
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<emphasis bold="true" pageId="0" pageNumber="151">China</emphasis>
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: Sichuan Province, Chengdu City, Longquanyi District, Longquan Mountain Forest Park, near Tiangong Temple,
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<geoCoordinate degrees="30.5305" direction="north" orientation="latitude" precision="5" value="30.5305">30.5305°N</geoCoordinate>
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,
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<geoCoordinate degrees="104.2709" direction="east" orientation="longitude" precision="5" value="104.2709">104.2709°E</geoCoordinate>
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, 636 m elev., 8.X.2019, M. Wei and Y. Shen leg.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="151">Paratypes</emphasis>
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1♀,
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<emphasis bold="true" pageId="0" pageNumber="151">China</emphasis>
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: Sichuan Province, Chengdu, Longquan District, Longquan Mountain Forest Park, near the expressway of Chengdu to Jianyang,
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<geoCoordinate degrees="30.5381" direction="north" orientation="latitude" precision="5" value="30.5381">30.5381°N</geoCoordinate>
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,
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<geoCoordinate degrees="104.3015" direction="east" orientation="longitude" precision="5" value="104.3015">104.3015°E</geoCoordinate>
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, 740 m elev., 16.X.2022, S. Wang leg.; 1♀,
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<emphasis bold="true" pageId="0" pageNumber="151">China</emphasis>
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: Sichuan Province, Chengdu, Longquan District, Longquan Mountain Forest Park, near the expressway of Chengdu to Jianyang,
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<geoCoordinate degrees="30.5381" direction="north" orientation="latitude" precision="5" value="30.5381">30.5381°N</geoCoordinate>
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,
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<geoCoordinate degrees="104.3015" direction="east" orientation="longitude" precision="5" value="104.3015">104.3015°E</geoCoordinate>
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, 740 m elev., 1.II.2023, S. Wang and M. Wei leg. Deposited in NHMSU.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="151" type="etymology">
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<paragraph pageId="0" pageNumber="151">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="151">
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The specific epithet is from "Asuka Langley Soryu", a fictional character wearing a red combat suit from the animation
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<normalizedToken originalValue="“Evangelion”">"Evangelion"</normalizedToken>
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(by the Japanese creator Hideaki Anno), refers to the body color; noun (name) in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="151" type="diagnosis">
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<paragraph pageId="0" pageNumber="151">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="151">
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Males can be distinguished from those of congeners, except
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<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. kagami" order="Araneae" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="kagami">
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<emphasis italics="true" pageId="0" pageNumber="151">L. kagami</emphasis>
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</taxonomicName>
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Wei & Lin, sp. nov., in lacking the BSC (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luthela asuka sp. nov. A male left palp bulb, prolateral view B male left palp bulb, ventral view C male left palp bulb, retrolateral view D left cymbium, ventral view E left palpal bulb, apical view F right palpal bulb, apical view G vulva, ventral view H vulva, dorsal view. Green arrows indicate small teeth on conductor. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1159.90120.figure4" httpUri="https://binary.pensoft.net/fig/844935" pageId="0" pageNumber="151">4A</figureCitation>
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), contrary to other species (cf.
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5091.1.5" author="Xu, X" journalOrPublisher="Zootaxa" pageId="0" pageNumber="151" pagination="131 - 154" refId="B22" refString="Xu, X, Yu, L, Liu, F, Li, D, 2022. Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae). Zootaxa 5091 (1): 131 - 154, DOI: https://doi.org/10.11646/zootaxa.5091.1.5" title="Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae)." url="https://doi.org/10.11646/zootaxa.5091.1.5" volume="5091" year="2022">Xu et al. 2022</bibRefCitation>
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: figs 3B, 5E, 6B, 7E, 10B, 12B, 14D), and in having the contrategulum bearning relatively dense, smaller serrated teeth (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luthela asuka sp. nov. A male left palp bulb, prolateral view B male left palp bulb, ventral view C male left palp bulb, retrolateral view D left cymbium, ventral view E left palpal bulb, apical view F right palpal bulb, apical view G vulva, ventral view H vulva, dorsal view. Green arrows indicate small teeth on conductor. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1159.90120.figure4" httpUri="https://binary.pensoft.net/fig/844935" pageId="0" pageNumber="151">4B, E</figureCitation>
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), rather than sparse and larger teeth in other species (cf.
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5091.1.5" author="Xu, X" journalOrPublisher="Zootaxa" pageId="0" pageNumber="151" pagination="131 - 154" refId="B22" refString="Xu, X, Yu, L, Liu, F, Li, D, 2022. Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae). Zootaxa 5091 (1): 131 - 154, DOI: https://doi.org/10.11646/zootaxa.5091.1.5" title="Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae)." url="https://doi.org/10.11646/zootaxa.5091.1.5" volume="5091" year="2022">Xu et al. 2022</bibRefCitation>
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: figs 3A, 5D, 6B, 7D, 10H, 12D, 14H). Males also differ from
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<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. kagami" order="Araneae" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="kagami">
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<emphasis italics="true" pageId="0" pageNumber="151">L. kagami</emphasis>
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</taxonomicName>
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sp. nov. in having two nearly invisible lateral teeth on the middle portion of the conductor and the longer TA (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luthela asuka sp. nov. A male left palp bulb, prolateral view B male left palp bulb, ventral view C male left palp bulb, retrolateral view D left cymbium, ventral view E left palpal bulb, apical view F right palpal bulb, apical view G vulva, ventral view H vulva, dorsal view. Green arrows indicate small teeth on conductor. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1159.90120.figure4" httpUri="https://binary.pensoft.net/fig/844935" pageId="0" pageNumber="151">4A-C, E, F</figureCitation>
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), rather than two relatively larger teeth and a shorter TA in the latter (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Luthela kagami sp. nov. A male left palp, prolateral view B male left palp, ventral view C male left palp, retrolateral view D right palpal bulb, ventral view E right palpal bulb, dorsal view F right palpal bulb, apical view G vulva, ventral view H vulva, dorsal view. Green arrows in B and F indicate small teeth on conductor. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1159.90120.figure7" httpUri="https://binary.pensoft.net/fig/844938" pageId="0" pageNumber="151">7B-D, F</figureCitation>
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). Females differ from those of congeners in having the paired receptacular clusters situated at the relatively short genital stalks and in the relatively smaller size (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luthela asuka sp. nov. A male left palp bulb, prolateral view B male left palp bulb, ventral view C male left palp bulb, retrolateral view D left cymbium, ventral view E left palpal bulb, apical view F right palpal bulb, apical view G vulva, ventral view H vulva, dorsal view. Green arrows indicate small teeth on conductor. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1159.90120.figure4" httpUri="https://binary.pensoft.net/fig/844935" pageId="0" pageNumber="151">4G, H</figureCitation>
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), rather than the long genital stalks and the larger size (cf.
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5091.1.5" author="Xu, X" journalOrPublisher="Zootaxa" pageId="0" pageNumber="151" pagination="131 - 154" refId="B22" refString="Xu, X, Yu, L, Liu, F, Li, D, 2022. Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae). Zootaxa 5091 (1): 131 - 154, DOI: https://doi.org/10.11646/zootaxa.5091.1.5" title="Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae)." url="https://doi.org/10.11646/zootaxa.5091.1.5" volume="5091" year="2022">Xu et al. 2022</bibRefCitation>
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: figs 4, 5H, I, 6H-M, 8, 9, 11, 13, 14H-M). Females differ from those of
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<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. kagami" order="Araneae" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="kagami">
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<emphasis italics="true" pageId="0" pageNumber="151">L. kagami</emphasis>
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</taxonomicName>
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sp. nov. in having the receptacular clusters relatively separated and the lateral pair larger than the middle pair (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luthela asuka sp. nov. A male left palp bulb, prolateral view B male left palp bulb, ventral view C male left palp bulb, retrolateral view D left cymbium, ventral view E left palpal bulb, apical view F right palpal bulb, apical view G vulva, ventral view H vulva, dorsal view. Green arrows indicate small teeth on conductor. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1159.90120.figure4" httpUri="https://binary.pensoft.net/fig/844935" pageId="0" pageNumber="151">4G, H</figureCitation>
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), rather than closer and nearly equal in size (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Luthela kagami sp. nov. A male left palp, prolateral view B male left palp, ventral view C male left palp, retrolateral view D right palpal bulb, ventral view E right palpal bulb, dorsal view F right palpal bulb, apical view G vulva, ventral view H vulva, dorsal view. Green arrows in B and F indicate small teeth on conductor. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1159.90120.figure7" httpUri="https://binary.pensoft.net/fig/844938" pageId="0" pageNumber="151">7G, H</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1159.90120.figure4" httpUri="https://binary.pensoft.net/fig/844935" pageId="0" pageNumber="151" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="151">
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<emphasis bold="true" pageId="0" pageNumber="151">Figure 4.</emphasis>
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<taxonomicName authorityName="Wei & Lin" authorityYear="2023" class="Arachnida" family="Heptathelidae" genus="Luthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Luthela asuka" order="Araneae" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="asuka">
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<emphasis italics="true" pageId="0" pageNumber="151">Luthela asuka</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="151">A</emphasis>
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male left palp bulb, prolateral view
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<emphasis bold="true" pageId="0" pageNumber="151">B</emphasis>
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male left palp bulb, ventral view
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<emphasis bold="true" pageId="0" pageNumber="151">C</emphasis>
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male left palp bulb, retrolateral view
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<emphasis bold="true" pageId="0" pageNumber="151">D</emphasis>
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left cymbium, ventral view
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<emphasis bold="true" pageId="0" pageNumber="151">E</emphasis>
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left palpal bulb, apical view
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<emphasis bold="true" pageId="0" pageNumber="151">F</emphasis>
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right palpal bulb, apical view
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<emphasis bold="true" pageId="0" pageNumber="151">G</emphasis>
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vulva, ventral view
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<emphasis bold="true" pageId="0" pageNumber="151">H</emphasis>
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vulva, dorsal view. Green arrows indicate small teeth on conductor. Scale bars: 0.50 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="151" type="description">
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<paragraph pageId="0" pageNumber="151">Description.</paragraph>
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<paragraph pageId="0" pageNumber="151">
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<emphasis bold="true" pageId="0" pageNumber="151">Male</emphasis>
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(holotype) (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. New species of Luthela A, B, G L. asuka sp. nov. from Longquanyi District, Chengdu C, D, H, I L. beijing sp. nov. from Zizhuyuan Park, Beijing E, F L. kagami sp. nov. from Guihua Township, Pengzhou City A, C, E male habitus, dorsal view B, D, F female habitus, dorsal view G female haibitus, ventral view H living female, dorsal view I burrow, vertical section, with red arrow pointing to the spider. Photographs by Chao Wu (H, I). Scale bars: 5.00 mm." figureDoi="10.3897/zookeys.1159.90120.figure3" httpUri="https://binary.pensoft.net/fig/844934" pageId="0" pageNumber="151">3A</figureCitation>
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). Carapace red; cervical and radial groove distinct. Cephalic region moderately raised. Chelicerae robust; fang furrow with 11 promarginal teeth of variable size. Sternum longer than wide. Abdomen pale yellow, with 5 large dorsal and 2 small posterior tergites, 4 tapering setae near posteromargin of 5 large tergites, and 2 on the rest. Seven spinnerets. Measurements: body 12.06 long. Carapace 5.59 long, 5.09 wide. Abdomen 5.92 long, 4.51 wide. Sternum 2.49 long, 1.91 wide. ALE> PLE> PME> AME. Leg I 18.00 (4.90 + 5.58 + 4.66 + 2.86), leg II 18.08 (4.54 + 5.42 + 5.04 + 3.08), leg III 19.31 (4.20 + 5.41 + 5.87 + 3.83), leg IV 26.76 (6.21 + 7.70 + 8.10 + 4.76).
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</paragraph>
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<paragraph pageId="0" pageNumber="151">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="151">Palp</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luthela asuka sp. nov. A male left palp bulb, prolateral view B male left palp bulb, ventral view C male left palp bulb, retrolateral view D left cymbium, ventral view E left palpal bulb, apical view F right palpal bulb, apical view G vulva, ventral view H vulva, dorsal view. Green arrows indicate small teeth on conductor. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1159.90120.figure4" httpUri="https://binary.pensoft.net/fig/844935" pageId="0" pageNumber="151">4A-F</figureCitation>
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): prolateral paracymbium pale, weakly sclerotized, with numerous setae and spines at distal and retrolateral surface. Contrategular margin denticulate, with large teeth on proximal part and smaller but denser teeth on distal part. Marginal apophysis of tegulum serrated, with tapering terminal apophysis of tegulum, margin of dorsal extension of terminal apophysis with teeth varied in size and distance. Conductor smooth, fused to embolic base, with large apical spine and 2 tiny lateral spines on middle portion. Embolus with translucent, flat opening and several ribbed ridges distally.
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</paragraph>
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<paragraph pageId="0" pageNumber="151">
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<emphasis bold="true" pageId="0" pageNumber="151">Female</emphasis>
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(one of paratypes) (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. New species of Luthela A, B, G L. asuka sp. nov. from Longquanyi District, Chengdu C, D, H, I L. beijing sp. nov. from Zizhuyuan Park, Beijing E, F L. kagami sp. nov. from Guihua Township, Pengzhou City A, C, E male habitus, dorsal view B, D, F female habitus, dorsal view G female haibitus, ventral view H living female, dorsal view I burrow, vertical section, with red arrow pointing to the spider. Photographs by Chao Wu (H, I). Scale bars: 5.00 mm." figureDoi="10.3897/zookeys.1159.90120.figure3" httpUri="https://binary.pensoft.net/fig/844934" pageId="0" pageNumber="151">3B, G</figureCitation>
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). Carapace red, with dark pattern; cervical and radial grooves distinct, with sparse spines. Cephalic region slightly elevated. Chelicerae more robust than male, fang furrow with 12 promarginal teeth of variable size, larger than male. Sternum longer than wide. Abdomen pale, with five large and five small tergites; chaetotaxy on tergites as in male. Seven spinnerets. Measurements: body 16.12 long. Carapace 7.02 long, 6.94 wide. Abdomen 8.93 long, 8.08 wide. Sternum 3.39 long, 1.86 wide. ALE> PLE> PME> AME. Leg I 14.84 (4.80 + 5.50 + 2.57 + 1.97), leg II 14.96 (4.59 + 4.66 + 3.32 + 2.39), leg III 14.70 (4.64 + 4.61 + 2.94 + 2.51), leg IV 22.24 (6.57 + 6.64 + 5.83 + 3.20).
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</paragraph>
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<paragraph pageId="0" pageNumber="151">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="151">Female genitalia</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luthela asuka sp. nov. A male left palp bulb, prolateral view B male left palp bulb, ventral view C male left palp bulb, retrolateral view D left cymbium, ventral view E left palpal bulb, apical view F right palpal bulb, apical view G vulva, ventral view H vulva, dorsal view. Green arrows indicate small teeth on conductor. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1159.90120.figure4" httpUri="https://binary.pensoft.net/fig/844935" pageId="0" pageNumber="151">4G, H</figureCitation>
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). Two pairs of receptacular clusters situated on short and thick stalks; lateral pair relatively larger than middle pair. Middle pair of receptacular clusters separated from each other, situated on anteromargin of bursa copulatrix; lateral receptacular clusters situated slightly dorsolaterally.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="151" type="distribution">
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<paragraph pageId="0" pageNumber="151">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="151">
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Known only from the type locality (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution records of 11 Luthela species. 1 = L. asuka sp. nov., 2 = L. badong, 3 = L. beijing sp. nov., 4 = L. dengfeng, 5 = L. handan, 6 = L. kagami sp. nov., 7 = L. luotianensis, 8 = L. schensiensis, 9 = L. taian, 10 = L. yiyuan, 11 = L. yuncheng." figureDoi="10.3897/zookeys.1159.90120.figure8" httpUri="https://binary.pensoft.net/fig/844939" pageId="0" pageNumber="151">8</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |