239 lines
30 KiB
XML
239 lines
30 KiB
XML
<document id="D6547F4A7C2BED9811066583EF708352" ID-DOI="10.11646/zootaxa.5446.4.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="11102071" ID-ZooBank="F82D6551-38AC-4732-B602-0BBDE4EA4377" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714635682967" checkinUser="plazi" docAuthor="Nel, André" docDate="2024" docId="7F4A8799FFF1FF82FF44F9F5FED1FC55" docLanguage="en" docName="zootaxa.5446.4.11.pdf" docOrigin="Zootaxa 5446 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5446.4.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gusagrion coloratum Nel 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="591" masterDocId="8373FFE1FFF0FF81FFD3FFC7FF80FFB8" masterDocTitle="Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae)" masterLastPageNumber="594" masterPageNumber="588" pageNumber="589" updateTime="1714636556268" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="9EB225A4A72985A0A9B6AA0765727C0C" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="198B4B4448793E28CEAB6D3F6A84C200">
|
||
<mods:title id="4A7D44767BB31A4744EE56C34DEA17A1">Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="19884DF39AA72B861912BA3182A5BE54" type="personal">
|
||
<mods:role id="03CEEE6683B03F35E8FA755570D74A24">
|
||
<mods:roleTerm id="A340043F9FD1787E3CAAD2DC32E9D1B0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D8A5F128947185C1D4FF65D8BF0F3118">Nel, André</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="B47E6CB85FED922DFCA22EEBC4DF6CE5">text</mods:typeOfResource>
|
||
<mods:relatedItem id="74B768269B648EAF7498FFF5F61E2033" type="host">
|
||
<mods:titleInfo id="0E301523BF69EF958B3AFF7F88F447A2">
|
||
<mods:title id="C27F9214948A8694498A61558126A8C1">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="3386F6586BC6B309681BE601B5108C0B">
|
||
<mods:date id="3FAD7950566FBC98EA0A771392747DA7">2024</mods:date>
|
||
<mods:detail id="B2EA375B581BCE8CD1C842765E614F70" type="pubDate">
|
||
<mods:number id="81E2A7FA7884B4A0609C0B320AC4A510">2024-05-02</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="D5C730C0AE962C06B72C025BC360E927" type="volume">
|
||
<mods:number id="A8C660DCD8FD55DB7A7E5F1B0344A8BC">5446</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="1425494C5BF09A65562919025B04F88D" type="issue">
|
||
<mods:number id="BA6A160E871916B5D1B29C1D0F3EB5EC">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="E91C84DF99CB80A2F44F0A725E332B2E" unit="page">
|
||
<mods:start id="2709166016E347299F90A2EC8A82DDFF">588</mods:start>
|
||
<mods:end id="E517DEA31D1197A55E9831CDC0561406">594</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="F111AA8AA362DA7AA4F5032F897AE8A9">
|
||
<mods:url id="1AEDC776CD3F2C111D40AC3BCC36A4F3">http://dx.doi.org/10.11646/zootaxa.5446.4.11</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="36891B1F04465154B50424073FF668D2">journal article</mods:classification>
|
||
<mods:identifier id="44D061A929A9A8F688AE1156D9FB1B43" type="DOI">10.11646/zootaxa.5446.4.11</mods:identifier>
|
||
<mods:identifier id="18C85843FBFFA548AF75E6A0F3B10516" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="3E0852ECC1AB6C5E2C74621DD3687066" type="Zenodo-Dep">11102071</mods:identifier>
|
||
<mods:identifier id="3F7D3C22634D56B7F7814BC4CF8124EC" type="ZooBank">F82D6551-38AC-4732-B602-0BBDE4EA4377</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="7F4A8799FFF1FF82FF44F9F5FED1FC55" ID-DOI="http://doi.org/10.5281/zenodo.11102084" ID-Zenodo-Dep="11102084" LSID="urn:lsid:plazi:treatment:7F4A8799FFF1FF82FF44F9F5FED1FC55" httpUri="http://treatment.plazi.org/id/7F4A8799FFF1FF82FF44F9F5FED1FC55" lastPageId="3" lastPageNumber="591" pageId="1" pageNumber="589">
|
||
<subSubSection id="BFF96504FFF1FF80FF44F9F5FD80F9F4" box="[151,512,1585,1612]" pageId="1" pageNumber="589" type="nomenclature">
|
||
<paragraph id="F75C368FFFF1FF80FF44F9F5FD80F9F4" blockId="1.[151,979,1585,1685]" box="[151,512,1585,1612]" pageId="1" pageNumber="589">
|
||
<heading id="AC1481E3FFF1FF80FF44F9F5FD80F9F4" bold="true" box="[151,512,1585,1612]" fontSize="11" level="1" pageId="1" pageNumber="589" reason="1">
|
||
<emphasis id="C597EA9DFFF1FF80FF44F9F5FD80F9F4" bold="true" box="[151,512,1585,1612]" pageId="1" pageNumber="589">
|
||
<taxonomicName id="30E34D0CFFF1FF80FF44F9F5FE1BF9F4" authority="Nel, 2024" authorityName="Nel" authorityYear="2024" box="[151,411,1585,1612]" class="Insecta" family="Thaumatoneuridae" genus="Gusagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="coloratum" status="sp. nov.">
|
||
<emphasis id="C597EA9DFFF1FF80FF44F9F5FE1BF9F4" bold="true" box="[151,411,1585,1612]" italics="true" pageId="1" pageNumber="589">Gusagrion coloratum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DEA457E6FFF1FF80FE70F9F5FD80F9F4" box="[419,512,1586,1612]" pageId="1" pageNumber="589" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFF96504FFF1FF80FF44F991FC53F92D" pageId="1" pageNumber="589" type="description">
|
||
<paragraph id="F75C368FFFF1FF80FF44F991FE88F9C8" blockId="1.[151,979,1585,1685]" box="[151,264,1622,1648]" pageId="1" pageNumber="589">
|
||
(
|
||
<figureCitation id="6FD82A0AFFF1FF80FF4CF991FF7DF9C8" box="[159,253,1622,1648]" captionStart="FIGURE 1" captionStartId="2.[151,250,1310,1334]" captionTargetBox="[151,1435,182,1286]" captionTargetId="figure-15@2.[151,1435,181,1286]" captionTargetPageId="2" captionText="FIGURE 1. Gusagrion coloratum gen. et sp. nov., holotype IP.358925. Photographs under alcohol of wing. A, part; B, xounterpart. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.11102073" httpUri="https://zenodo.org/record/11102073/files/figure.png" pageId="1" pageNumber="589">Figure 1</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="F75C368FFFF1FF80FF44F9BDFC53F92D" blockId="1.[151,979,1585,1685]" box="[151,979,1658,1685]" pageId="1" pageNumber="589">
|
||
<uri id="83723A8DFFF1FF80FF44F9BDFC53F92D" box="[151,979,1658,1685]" pageId="1" pageNumber="589">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="83450C5AFFF1FF80FE79F9BDFC53F92D" box="[426,979,1658,1685]" pageId="1" pageNumber="589">7A54A54E-4503-4BD6-B206-1FDF3EA84AA0</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFF96504FFF1FF80FF44F906FDF0F8F0" pageId="1" pageNumber="589" type="materials_examined">
|
||
<paragraph id="F75C368FFFF1FF80FF44F906FDE9F8B8" blockId="1.[151,1436,1729,1936]" pageId="1" pageNumber="589">
|
||
<emphasis id="C597EA9DFFF1FF80FF44F906FE86F963" bold="true" box="[151,262,1729,1755]" pageId="1" pageNumber="589">Material.</emphasis>
|
||
<materialsCitation id="478B3CD2FFF1FF80FEC1F905FDE9F8B8" country="United States of America" location="Invertebrate Paleontology Collection" municipality="Yale Peabody Museum" pageId="1" pageNumber="589" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="2858882DFFF1FF80FEC1F905FEFAF964" box="[274,378,1730,1756]" pageId="1" pageNumber="589" type="holotype">Holotype</typeStatus>
|
||
IP.358925 (part and counterpart of a complete wing), stored at
|
||
<location id="F23C6054FFF1FF80FBA6F905FE8FF8B8" LSID="urn:lsid:plazi:treatment:7F4A8799FFF1FF82FF44F9F5FED1FC55:F23C6054FFF1FF80FBA6F905FE8FF8B8" country="United States of America" municipality="Yale Peabody Museum" name="Invertebrate Paleontology Collection" pageId="1" pageNumber="589">Invertebrate Paleontology Collection</location>
|
||
,
|
||
<collectingMunicipality id="1738ACF5FFF1FF80FECAF921FD9EF8B8" box="[281,542,1766,1792]" pageId="1" pageNumber="589">Yale Peabody Museum</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="8FF4761FFFF1FF80FDFAF921FDE5F8B8" box="[553,613,1766,1792]" name="United States of America" pageId="1" pageNumber="589">USA</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="F75C368FFFF1FF80FF14F8CEFDF0F8F0" blockId="1.[151,1436,1729,1936]" pageId="1" pageNumber="589">
|
||
<emphasis id="C597EA9DFFF1FF80FF14F8CEFE18F89C" bold="true" box="[199,408,1801,1828]" pageId="1" pageNumber="589">Age and outcrop.</emphasis>
|
||
Eocene (Ypresian), Green River Formation, Parachute Creek member, Gus’ pit, two miles west of Rio Blanco store,
|
||
<collectingRegion id="3527F86DFFF1FF80FE69F8E9FDA5F8F0" box="[442,549,1838,1864]" country="United States of America" name="Colorado" pageId="1" pageNumber="589">Colorado</collectingRegion>
|
||
,
|
||
<collectingCountry id="8FF4761FFFF1FF80FDE3F8E9FDECF8F0" box="[560,620,1838,1864]" name="United States of America" pageId="1" pageNumber="589">USA</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFF96504FFF1FF80FF14F895FCC9F8D4" box="[199,841,1874,1900]" pageId="1" pageNumber="589" type="etymology">
|
||
<paragraph id="F75C368FFFF1FF80FF14F895FCC9F8D4" blockId="1.[151,1436,1729,1936]" box="[199,841,1874,1900]" pageId="1" pageNumber="589">
|
||
<emphasis id="C597EA9DFFF1FF80FF14F895FECEF8D4" bold="true" box="[199,334,1874,1900]" pageId="1" pageNumber="589">Etymology.</emphasis>
|
||
Named after the colored distal third of wing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFF96504FFF1FF80FF14F8B2FBAFF828" box="[199,1071,1909,1936]" pageId="1" pageNumber="589" type="diagnosis">
|
||
<paragraph id="F75C368FFFF1FF80FF14F8B2FBAFF828" blockId="1.[151,1436,1729,1936]" box="[199,1071,1909,1936]" pageId="1" pageNumber="589">
|
||
<emphasis id="C597EA9DFFF1FF80FF14F8B2FEC2F837" bold="true" box="[199,322,1909,1935]" pageId="1" pageNumber="589">Diagnosis.</emphasis>
|
||
As for the genus. A dark brown area covering apical third of wing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="A39C6607FFF2FF83FF44FAD9FE5DFAE2" ID-DOI="http://doi.org/10.5281/zenodo.11102073" ID-Zenodo-Dep="11102073" httpUri="https://zenodo.org/record/11102073/files/figure.png" pageId="2" pageNumber="590" startId="2.[151,250,1310,1334]" targetBox="[151,1435,182,1286]" targetPageId="2" targetType="figure">
|
||
<paragraph id="F75C368FFFF2FF83FF44FAD9FE5DFAE2" blockId="2.[151,1437,1310,1371]" pageId="2" pageNumber="590">
|
||
<emphasis id="C597EA9DFFF2FF83FF44FAD9FE98FA8E" bold="true" box="[151,280,1310,1334]" pageId="2" pageNumber="590">FIGURE 1.</emphasis>
|
||
<taxonomicName id="30E34D0CFFF2FF83FEF1FAD8FD81FA8F" authority="Nel, 2024" authorityName="Nel" authorityYear="2024" box="[290,513,1311,1335]" class="Insecta" family="Thaumatoneuridae" genus="Gusagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="590" phylum="Arthropoda" rank="species" species="coloratum" status="gen. et sp. nov.">
|
||
<emphasis id="C597EA9DFFF2FF83FEF1FAD8FD81FA8F" box="[290,513,1311,1335]" italics="true" pageId="2" pageNumber="590">Gusagrion coloratum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C597EA9DFFF2FF83FDDFFAD8FD3EFA8F" bold="true" box="[524,702,1311,1335]" pageId="2" pageNumber="590">
|
||
<taxonomicNameLabel id="DEA457E6FFF2FF83FDDFFAD8FD39FA8F" box="[524,697,1311,1335]" pageId="2" pageNumber="590" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
holotype IP.358925. Photographs under alcohol of wing. A, part; B, xounterpart. Scale bars = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="BFF96504FFF2FF82FF14FA6AFBEBFE15" lastPageId="3" lastPageNumber="591" pageId="2" pageNumber="590" type="description">
|
||
<paragraph id="F75C368FFFF2FF82FF14FA6AFBEBFE15" blockId="2.[151,1437,1453,2019]" lastBlockId="3.[151,1437,151,1005]" lastPageId="3" lastPageNumber="591" pageId="2" pageNumber="590">
|
||
<emphasis id="C597EA9DFFF2FF83FF14FA6AFED9FA7F" bold="true" box="[199,345,1453,1479]" pageId="2" pageNumber="590">Description.</emphasis>
|
||
A complete wing, with a dark brown area covering apical third, wing
|
||
<quantity id="301B9B6AFFF2FF83FB8EFA6AFB40FA7F" box="[1117,1216,1453,1479]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="2" pageNumber="590" unit="mm" value="28.8">28.8 mm</quantity>
|
||
long,
|
||
<quantity id="301B9B6AFFF2FF83FAD7FA6AFAD9FA7F" box="[1284,1369,1453,1479]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="2" pageNumber="590" unit="mm" value="8.4">8.4 mm</quantity>
|
||
wide, broad and with numerous small cells; distance from base to arculus
|
||
<quantity id="301B9B6AFFF2FF83FC55FA16FC5DFA53" box="[902,989,1489,1515]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="590" unit="mm" value="3.6">3.6 mm</quantity>
|
||
, from arculus to nodus
|
||
<quantity id="301B9B6AFFF2FF83FB32FA16FAB9FA53" box="[1249,1337,1489,1515]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="590" unit="mm" value="3.2">3.2 mm</quantity>
|
||
; primary antenodal crossveins Ax1 and Ax2 well-defined; Ax1
|
||
<quantity id="301B9B6AFFF2FF83FD2BFA32FCCFF9B7" box="[760,847,1525,1551]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="590" unit="mm" value="2.2">2.2 mm</quantity>
|
||
from wing base, distance between Ax1 and Ax2
|
||
<quantity id="301B9B6AFFF2FF83FAAAFA32FF46F98B" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="590" unit="mm" value="1.4">1.4 mm</quantity>
|
||
; Ax2 just distad arculus; no secondary antenodal crossveins; ca. 34 postnodal crossveins, not well aligned with postsubnodal crossveins; pterostigma rather short but broad,
|
||
<quantity id="301B9B6AFFF2FF83FC5FF9FAFC64F9EF" box="[908,996,1597,1623]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="590" unit="mm" value="2.4">2.4 mm</quantity>
|
||
long and
|
||
<quantity id="301B9B6AFFF2FF83FB89F9FAFB33F9EF" box="[1114,1203,1597,1623]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="590" unit="mm" value="0.8">0.8 mm</quantity>
|
||
wide, covering four cells; distal edge of pterostigma more oblique than proximal one; pterostigmal brace absent; one row of crossveins between C and RA distal of pterostigma; posterior part of arculus (= basal discoidal crossvein) present, therefore discoidal cell basally closed; discoidal cell rectangular, elongate, narrow (
|
||
<quantity id="301B9B6AFFF2FF83FC04F96EFBAEF97B" box="[983,1070,1705,1731]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="590" unit="mm" value="1.6">1.6 mm</quantity>
|
||
long,
|
||
<quantity id="301B9B6AFFF2FF83FBA5F96EFB4CF97B" box="[1142,1228,1705,1731]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="590" unit="mm" value="0.4">0.4 mm</quantity>
|
||
wide), and free of crossveins; origins of RP and MA separated in arculus; bend of ScP at nodus very abrupt and Z-like; nodal vein and subnodus oblique; subnodus aligned with nodal crossvein; no crossveins in antesubnodal area; median space free of crossveins; submedian space free of crossveins except for CuP-crossing; subdiscoidal cells free; anal area narrow (
|
||
<quantity id="301B9B6AFFF2FF83FF4CF8FEFF77F8EB" box="[159,247,1849,1875]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="590" unit="mm" value="0.3">0.3 mm</quantity>
|
||
wide in middle), not wider than subdiscoidal cell, with one crossvein below subdiscoidal cell; cubito-anal area max. 2.0 mm wide in its broadest part, with rows of cells; CuA well-defined and curved; CuA not forked; area between MP and CuA progressively broadened; MP regularly curved, not distally zigzagged, and ending between nodus and pterostigma; basal half of postdiscoidal area between MA and MP narrow with only one row of cells, but greatly widened distally, and with two long secondary longitudinal veins; MA ending well basad pterostigma level; MA, RP3/4, and IR2 rather straight, but distally curved towards hind margin; no antefurcal crossveins between basal parts of RP and MA; area between RP3/4 and MA narrow with one row of cells between them in basal half, but area distally widened with two long secondary longitudinal veins; area between RP3/4 and IR2 progressively broadened, with two long secondary longitudinal veins; RP3/4 ending beneath pterostigma; base of RP3/4
|
||
<quantity id="301B9B6AFFF3FF82FA95FEC4FA1DFEA5" box="[1350,1437,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="591" unit="mm" value="1.8">1.8 mm</quantity>
|
||
basad subnodus; base of IR2 below nodus; RP1/2 straight at subnodus; base of RP2 very far from subnodus, at 13 cells distal from it; IR1 well-defined, long, and curved, originating four cells distad RP2; no lestine oblique vein ‘O’ between RP2 and IR2; RP2 and IR2 with only one row of cells between them until level of base of pterostigma; RP1 without distinct bend at pterostigmal brace; wings well petiolated, petiole
|
||
<quantity id="301B9B6AFFF3FF82FC06FE54FBABFE15" box="[981,1067,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="591" unit="mm" value="2.8">2.8 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFF96504FFF3FF82FF14FE70FED1FC55" pageId="3" pageNumber="591" type="discussion">
|
||
<paragraph id="F75C368FFFF3FF82FF14FE70FBF3FDD9" blockId="3.[151,1437,151,1005]" pageId="3" pageNumber="591">
|
||
<emphasis id="C597EA9DFFF3FF82FF14FE70FEB0FE69" bold="true" box="[199,304,439,465]" pageId="3" pageNumber="591">Remark.</emphasis>
|
||
This damselfly wing fits quite well with those of the two eodysagrionine genera and species
|
||
<taxonomicName id="30E34D0CFFF3FF82FF44FE1CFC7BFE4D" authority="Bechly, Garrouste, Aase, Karr, Grande & Nel, 2021" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[151,1019,475,501]" class="Insecta" family="Dysagrionidae" genus="Tynskysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="brookeae">
|
||
<emphasis id="C597EA9DFFF3FF82FF44FE1CFE28FE4D" box="[151,424,475,501]" italics="true" pageId="3" pageNumber="591">Tynskysagrion brookeae</emphasis>
|
||
<bibRefCitation id="93724B7EFFF3FF82FE63FE1CFC7BFE4D" author="Bechly, G. & Garrouste, R. & Aase, A. & Karr, J. A. & Grande, L. & Nel, A." box="[432,1019,475,501]" pageId="3" pageNumber="591" pagination="1373 - 1402" refId="ref3284" refString="Bechly, G., Garrouste, R., Aase, A., Karr, J. A., Grande, L. & Nel, A. (2021) The damselfly palaeofauna from the Eocene of Wyoming and Colorado, USA (Insecta, Odonata, Zygoptera). Papers in Palaeontology, 7 (3), 1373 - 1402. https: // doi. org / 10.1002 / spp 2.1346" type="journal article" year="2021">Bechly, Garrouste, Aase, Karr, Grande & Nel, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="30E34D0CFFF3FF82FBE7FE1CFD02FDA1" authority="Bechly, Garrouste, Aase, Karr, Grande & Nel, 2021" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" class="Insecta" family="Dysagrionidae" genus="Oreodysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="tenebris">
|
||
<emphasis id="C597EA9DFFF3FF82FBE7FE1CFABFFE4D" box="[1076,1343,475,501]" italics="true" pageId="3" pageNumber="591">Oreodysagrion tenebris</emphasis>
|
||
<bibRefCitation id="93724B7EFFF3FF82FA9BFE1CFD02FDA1" author="Bechly, G. & Garrouste, R. & Aase, A. & Karr, J. A. & Grande, L. & Nel, A." pageId="3" pageNumber="591" pagination="1373 - 1402" refId="ref3284" refString="Bechly, G., Garrouste, R., Aase, A., Karr, J. A., Grande, L. & Nel, A. (2021) The damselfly palaeofauna from the Eocene of Wyoming and Colorado, USA (Insecta, Odonata, Zygoptera). Papers in Palaeontology, 7 (3), 1373 - 1402. https: // doi. org / 10.1002 / spp 2.1346" type="journal article" year="2021">Bechly, Garrouste, Aase, Karr, Grande & Nel, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
because of the elongate and narrow discoidal cell, long petiole, nodus situated in a very basal position, distal third of wing very broad.
|
||
<taxonomicName id="30E34D0CFFF3FF82FCB5FDE4FC75FD85" authorityName="Rust, Petrulevicius & Nel" authorityYear="2008" box="[870,1013,547,573]" class="Insecta" family="Dysagrionidae" genus="Eodysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FCB5FDE4FC75FD85" box="[870,1013,547,573]" italics="true" pageId="3" pageNumber="591">Eodysagrion</emphasis>
|
||
</taxonomicName>
|
||
has a shorter discoidal cell, the nodus situated in a more distal position, and much less postnodal crossveins (
|
||
<bibRefCitation id="93724B7EFFF3FF82FC7CFD80FBE7FDD9" author="Rust, J. & Petrulevicius, J. & Nel, A." box="[943,1127,583,609]" pageId="3" pageNumber="591" pagination="709 - 713" refId="ref4386" refString="Rust, J., Petrulevicius, J. & Nel, A. (2008) The first damselflies from the Lowermost Eocene of Denmark, with a description of a new subfamily (Odonata, Zygoptera: Dysagrionidae). Palaeontology, 51 (3), 709 - 713. https: // doi. org / 10.1111 / j. 1475 - 4983.2008.00780. x" type="journal article" year="2008">
|
||
Rust
|
||
<emphasis id="C597EA9DFFF3FF82FC38FD80FBA3FDD9" box="[1003,1059,583,609]" italics="true" pageId="3" pageNumber="591">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F75C368FFFF3FF82FF14FDACFBD3FCE5" blockId="3.[151,1437,151,1005]" pageId="3" pageNumber="591">
|
||
The new fossil differs from
|
||
<taxonomicName id="30E34D0CFFF3FF82FE2BFDACFD19FD3D" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[504,665,619,645]" class="Insecta" family="Dysagrionidae" genus="Tynskysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FE2BFDACFD19FD3D" box="[504,665,619,645]" italics="true" pageId="3" pageNumber="591">Tynskysagrion</emphasis>
|
||
</taxonomicName>
|
||
in the much narrower anal area below the subdiscoidal cell, at most as broad as subdiscoidal cell, as in
|
||
<taxonomicName id="30E34D0CFFF3FF82FE27FD48FD1DFD11" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[500,669,655,681]" class="Insecta" family="Dysagrionidae" genus="Oreodysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FE27FD48FD1DFD11" box="[500,669,655,681]" italics="true" pageId="3" pageNumber="591">Oreodysagrion</emphasis>
|
||
</taxonomicName>
|
||
vs. as broad as subdiscoidal plus discoidal cells in
|
||
<taxonomicName id="30E34D0CFFF3FF82FB17FD48FAE5FD11" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[1220,1381,655,681]" class="Insecta" family="Dysagrionidae" genus="Tynskysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FB17FD48FAE5FD11" box="[1220,1381,655,681]" italics="true" pageId="3" pageNumber="591">Tynskysagrion</emphasis>
|
||
</taxonomicName>
|
||
. The new fossil also has only one crossvein below subdiscoidal cell, vs. two in
|
||
<taxonomicName id="30E34D0CFFF3FF82FC19FD74FBF3FD75" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[970,1139,691,717]" class="Insecta" family="Dysagrionidae" genus="Oreodysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FC19FD74FBF3FD75" box="[970,1139,691,717]" italics="true" pageId="3" pageNumber="591">Oreodysagrion</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="301B9B6AFFF3FF82FB7BFD74FB6EFD75" box="[1192,1262,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="3" pageNumber="591" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
|
||
<taxonomicName id="30E34D0CFFF3FF82FB27FD74FA15FD75" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[1268,1429,691,717]" class="Insecta" family="Dysagrionidae" genus="Tynskysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FB27FD74FA15FD75" box="[1268,1429,691,717]" italics="true" pageId="3" pageNumber="591">Tynskysagrion</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="30E34D0CFFF3FF82FF65FD10FEDFFD49" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[182,351,727,753]" class="Insecta" family="Dysagrionidae" genus="Oreodysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FF65FD10FEDFFD49" box="[182,351,727,753]" italics="true" pageId="3" pageNumber="591">Oreodysagrion</emphasis>
|
||
</taxonomicName>
|
||
, the nodus is situated in a very basal position, 14% of the distance between Ax2 and wing apex vs. 19% in the species of
|
||
<taxonomicName id="30E34D0CFFF3FF82FE62FD3CFDD2FCAD" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[433,594,763,789]" class="Insecta" family="Dysagrionidae" genus="Tynskysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FE62FD3CFDD2FCAD" box="[433,594,763,789]" italics="true" pageId="3" pageNumber="591">Tynskysagrion</emphasis>
|
||
</taxonomicName>
|
||
. In the new fossil, the nodus is situated at 10% of the distance between Ax2 and wing apex, which better fits with
|
||
<taxonomicName id="30E34D0CFFF3FF82FD9BFCD8FD71FC81" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[584,753,799,825]" class="Insecta" family="Dysagrionidae" genus="Oreodysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FD9BFCD8FD71FC81" box="[584,753,799,825]" italics="true" pageId="3" pageNumber="591">Oreodysagrion</emphasis>
|
||
</taxonomicName>
|
||
. But the wing of
|
||
<taxonomicName id="30E34D0CFFF3FF82FC6FFCD8FBE5FC81" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[956,1125,799,825]" class="Insecta" family="Dysagrionidae" genus="Oreodysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FC6FFCD8FBE5FC81" box="[956,1125,799,825]" italics="true" pageId="3" pageNumber="591">Oreodysagrion</emphasis>
|
||
</taxonomicName>
|
||
is more slender than in the new fossil and in
|
||
<taxonomicName id="30E34D0CFFF3FF82FE8EFC84FE7EFCE5" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[349,510,835,861]" class="Insecta" family="Dysagrionidae" genus="Tynskysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C597EA9DFFF3FF82FE8EFC84FE7EFCE5" box="[349,510,835,861]" italics="true" pageId="3" pageNumber="591">Tynskysagrion</emphasis>
|
||
</taxonomicName>
|
||
. Thus the new fossil does not fit in these two genera.
|
||
</paragraph>
|
||
<paragraph id="F75C368FFFF3FF82FF14FCA0FED1FC55" blockId="3.[151,1437,151,1005]" pageId="3" pageNumber="591">
|
||
Lastly, the color pattern of the new wing is strongly different from those of
|
||
<taxonomicName id="30E34D0CFFF3FF82FB8AFCA0FAE8FC39" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[1113,1384,871,897]" class="Insecta" family="Dysagrionidae" genus="Oreodysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="tenebris">
|
||
<emphasis id="C597EA9DFFF3FF82FB8AFCA0FAE8FC39" box="[1113,1384,871,897]" italics="true" pageId="3" pageNumber="591">Oreodysagrion tenebris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="30E34D0CFFF3FF82FF44FC4CFE26FC1D" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[151,422,907,933]" class="Insecta" family="Dysagrionidae" genus="Tynskysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="brookeae">
|
||
<emphasis id="C597EA9DFFF3FF82FF44FC4CFE26FC1D" box="[151,422,907,933]" italics="true" pageId="3" pageNumber="591">Tynskysagrion brookeae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="30E34D0CFFF3FF82FE60FC4CFD3BFC1D" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[435,699,907,933]" class="Insecta" family="Dysagrionidae" genus="Oreodysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="tenebris">
|
||
<emphasis id="C597EA9DFFF3FF82FE60FC4CFD3BFC1D" box="[435,699,907,933]" italics="true" pageId="3" pageNumber="591">Oreodysagrion tenebris</emphasis>
|
||
</taxonomicName>
|
||
has a dark brown band between the base of RP2 and the basal side of pterostigma.
|
||
<taxonomicName id="30E34D0CFFF3FF82FE99FC68FDDAFC71" authorityName="Bechly, Garrouste, Aase, Karr, Grande & Nel" authorityYear="2021" box="[330,602,943,969]" class="Insecta" family="Dysagrionidae" genus="Tynskysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="brookeae">
|
||
<emphasis id="C597EA9DFFF3FF82FE99FC68FDDAFC71" box="[330,602,943,969]" italics="true" pageId="3" pageNumber="591">Tynskysagrion brookeae</emphasis>
|
||
</taxonomicName>
|
||
has a completely darkened wings or with a narrow hyaline band in distal half of the wing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |