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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1159.99218" ID-Pensoft-Pub="1313-2970-1159-87" ID-Pensoft-UUID="ECCECB868071523DB7628E304850BD62" ID-ZooBank="CD97DD120549412F976508D8DE21605B" ModsDocID="1313-2970-1159-87" checkinTime="1682458941538" checkinUser="pensoft" docAuthor="Okada, Ryohei, Jaitrong, Weeyawat &amp; Wewalka, Guenther" docDate="2023" docId="FAAD1C24D633582CAA7DE3F55B9F8B09" docLanguage="en" docName="ZooKeys 1159: 87-119" docOrigin="ZooKeys 1159" docPubDate="2023-04-25" docSource="http://dx.doi.org/10.3897/zookeys.1159.99218" docTitle="Microdytes shepardi Wewalka 1997" docType="treatment" docVersion="1" id="ECCECB868071523DB7628E304850BD62" lastPageNumber="87" masterDocId="ECCECB868071523DB7628E304850BD62" masterDocTitle="A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae)" masterLastPageNumber="119" masterPageNumber="87" pageNumber="87" updateTime="1682458941538" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Okada, Ryohei</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8488-0660</mods:nameIdentifier>
<mods:affiliation>Thailand Natural History Museum, National Science Museum, Technopolis, Pathum Thani, Thailand &amp; Coleopterological Society of Japan, National Museum of Nature and Science, Tsukuba, Japan</mods:affiliation>
<mods:nameIdentifier type="email">wasserinsekt@kub.biglobe.ne.jp</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jaitrong, Weeyawat</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1362-0754</mods:nameIdentifier>
<mods:affiliation>Thailand Natural History Museum, National Science Museum, Technopolis, Pathum Thani, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wewalka, Guenther</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2994-8001</mods:nameIdentifier>
<mods:affiliation>Starkfriedgasse 16 / 1 / 3, A - 1190 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1159</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>119</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1159.99218</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1159.99218</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1159-87</mods:identifier>
<mods:identifier type="ZooBank">CD97DD120549412F976508D8DE21605B</mods:identifier>
<mods:identifier type="Pensoft-UUID">ECCECB868071523DB7628E304850BD62</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:FAAD1C24D633582CAA7DE3F55B9F8B09" httpUri="http://treatment.plazi.org/id/FAAD1C24D633582CAA7DE3F55B9F8B09" lastPageNumber="87" pageId="0" pageNumber="87">
<subSubSection pageId="0" pageNumber="87" type="nomenclature">
<paragraph pageId="0" pageNumber="87">
<taxonomicName ID-CoL="42WF5" LSID="FAAD1C24-D633-582C-AA7D-E3F55B9F8B09" authority="Wewalka, 1997" authorityName="Wewalka" authorityYear="1997" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes shepardi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="shepardi">Microdytes shepardi Wewalka, 1997</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="description">
<paragraph pageId="0" pageNumber="87">
<figureCitation captionStart="Figures 1837" captionStartId="F7" captionText="Figures 18 - 37. Habitus of 18 Microdytes akitai 19 M. balkei 20 M. dimorphus 21 M. feryi (paratype, from Myanmar) 22 M. franzi 23 M. gabrielae 24 M. heineri 25 M. maculatus 26 M. mariannae 27 M. menopausis 28 M. paoloi 29 M. pasiricus 30 M. pederzanii 31 M. rocchii 32 M. schoedli 33 M. schoenmanni (from Myanmar) 34 M. schwendingeri 35 M. shepardi 36 M. shunichii 37 M. zetteli. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures18-37" httpUri="https://binary.pensoft.net/fig/842227" pageId="0" pageNumber="87">Fig. 35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="reference_group">
<paragraph pageId="0" pageNumber="87">
<taxonomicName authorityName="Wewalka" authorityYear="1997" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes shepardi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="shepardi">Microdytes shepardi</taxonomicName>
Wewalka, 1997: 37;
<bibRefCitation author="Wewalka, G" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="87" pagination="21 - 39" refId="B24" refString="Wewalka, G, 2011. New species, new records and taxonomic updates on Microdytes Balfour-Browne, 1946. Koleopterologische Rundschau 81: 21 - 39" title="New species, new records and taxonomic updates on Microdytes Balfour-Browne, 1946." volume="81" year="2011">Wewalka 2011</bibRefCitation>
: 33;
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Etudes Entomologiques Motschulsky" pageId="0" pageNumber="87" publicationUrl="http://www.norrent.se" refId="B13" refString="Nilsson, AN, Hajek, J, 2023. A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1.I.2023. http://www.norrent.se" title="A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2023." url="http://www.norrent.se" year="2023">
Nilsson and
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
2023
</bibRefCitation>
: 212.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="type locality">
<paragraph pageId="0" pageNumber="87">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="87">Thailand, Phetchabun Province, Nam Nao NP.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="materials_examined">
<paragraph pageId="0" pageNumber="87">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="87">
<materialsCitation collectingDate="2021-03-20" collectorName="R. Okada" country="Thailand" elevation="430" location="Ping Khong St." municipality="Chiang Dao District" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiang Mai Province">
<collectingCountry name="Thailand">Thailand</collectingCountry>
:
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
.
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Chiang Dao District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:FAAD1C24D633582CAA7DE3F55B9F8B09:0DC28DF6DBC2928B695578A6D8821FE0" country="Thailand" municipality="Chiang Dao District" name="Ping Khong St." stateProvince="Chiang Mai Province">Ping Khong St.</location>
226 (alt.
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.3" unit="m" value="430.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.3" unit="m" value="430.0">430 m</elevation>
</quantity>
),
<collectingDate value="2021-03-20">20.III.2021</collectingDate>
,
<collectorName>R. Okada</collectorName>
leg. (CRO)
</materialsCitation>
;
<materialsCitation collectingDate="2019-08-02" collectorName="R. Okada" country="Thailand" elevation="440" location="Muang Yai St." municipality="Wiang Kaen District" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Rai Province">
<collectingRegion country="Thailand" name="Chiang Rai">Chiang Rai Province</collectingRegion>
.
<specimenCount type="male">1♂</specimenCount>
,
<collectingMunicipality>Wiang Kaen District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:FAAD1C24D633582CAA7DE3F55B9F8B09:C271287ED07CCE8C8F19D88359C30032" country="Thailand" municipality="Wiang Kaen District" name="Muang Yai St." stateProvince="Chiang Rai Province">Muang Yai St.</location>
71 (alt.
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">440 m</elevation>
</quantity>
),
<collectingDate value="2019-08-02">2.VIII.2019</collectingDate>
,
<collectorName>R. Okada</collectorName>
leg. (CRO)
</materialsCitation>
;
<materialsCitation collectingDate="2020-06-20" collectorName="R. Okada" country="Thailand" elevation="440" location="Kaen Makrut St." municipality="Ban Rai District" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Uthai Thani Province">
<collectingRegion country="Thailand" name="Uthai Thani">Uthai Thani Province</collectingRegion>
.
<specimenCount type="male">5♂♂</specimenCount>
,
<specimenCount type="female">2♀♀</specimenCount>
,
<collectingMunicipality>Ban Rai District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:FAAD1C24D633582CAA7DE3F55B9F8B09:4A99D1207F72704B0A140FCB47E0174E" country="Thailand" municipality="Ban Rai District" name="Kaen Makrut St." stateProvince="Uthai Thani Province">Kaen Makrut St.</location>
128 (alt.
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">440 m</elevation>
</quantity>
),
<collectingDate value="2020-06-20">20.VI.2020</collectingDate>
,
<collectorName>R. Okada</collectorName>
leg. (CGW, CRO, THNHM) (Fig.
<figureCitation captionStart="Figures 1837" captionStartId="F7" captionText="Figures 18 - 37. Habitus of 18 Microdytes akitai 19 M. balkei 20 M. dimorphus 21 M. feryi (paratype, from Myanmar) 22 M. franzi 23 M. gabrielae 24 M. heineri 25 M. maculatus 26 M. mariannae 27 M. menopausis 28 M. paoloi 29 M. pasiricus 30 M. pederzanii 31 M. rocchii 32 M. schoedli 33 M. schoenmanni (from Myanmar) 34 M. schwendingeri 35 M. shepardi 36 M. shunichii 37 M. zetteli. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures18-37" httpUri="https://binary.pensoft.net/fig/842227" pageId="0" pageNumber="87">35</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="87">
<materialsCitation collectingDate="2002-05-11" collectingDateMax="2002-05-12" collectingDateMin="2002-05-11" collectorName="V. Kuban" country="Laos" elevation="750" latitude="19.583334" location="Thong Khan" longLatPrecision="1271" longitude="101.96667" specimenCount="1" specimenCount-male="1" stateProvince="Luang Prabang Province">
<collectingCountry name="Laos">Laos</collectingCountry>
:
<specimenCount type="male">1♂</specimenCount>
,
<collectingRegion country="Laos" name="Louangphabang">Luang Prabang Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:FAAD1C24D633582CAA7DE3F55B9F8B09:408C8A0556443B2EC505133B743D48A4" country="Laos" latitude="19.583334" longLatPrecision="1271" longitude="101.96667" name="Thong Khan" stateProvince="Luang Prabang Province">Thong Khan</location>
,
<geoCoordinate degrees="19" direction="north" minutes="35" orientation="latitude" precision="925" value="19.583334">19°35'N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="58" orientation="longitude" precision="925" value="101.96667">101°58'E</geoCoordinate>
, alt. ca.
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.5" unit="m" value="750.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.5" unit="m" value="750.0">750 m</elevation>
</quantity>
,
<collectingDate value="2002-05-11" valueMax="2002-05-12" valueMin="2002-05-11">11.-12.V.2002</collectingDate>
,
<collectorName>
V.
<normalizedToken originalValue="Kubáň">Kuban</normalizedToken>
</collectorName>
leg. (NMB)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="distribution">
<paragraph pageId="0" pageNumber="87">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="87">Thailand: Chiang Mai, Chiang Rai (first record), Mae Hong Son, Phetchabun, Uthai Thani (first record), Chaiyaphum and Sakon Nakhon provinces; Laos: Luang Prabang Province; China: Yunnan Province.</paragraph>
</subSubSection>
</treatment>
</document>