472 lines
66 KiB
XML
472 lines
66 KiB
XML
<document id="7B579EA985AD4C591748C28E03776901" ID-CLB-Dataset="55814" ID-DOI="10.1080/00222933.2013.791952" ID-GBIF-Dataset="857d6372-de50-463d-8471-717e72bdd3ba" ID-ISSN="1464-5262" ID-Zenodo-Dep="4608237" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615907007564" checkinUser="carolina" docAuthor="Schmidt, Joachim, Wrase, David W. & Sciaky, Riccardo" docDate="2013" docId="FA03878BD35BFF81FE74FBD5FBC03722" docLanguage="en" docName="JNATHIST.47.41-42.2671-2689.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 47 (41 - 42)" docSource="http://dx.doi.org/10.1080/00222933.2013.791952" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Eobroscus masumotoi Morita 1990" docType="treatment" docVersion="7" lastPageNumber="2688" masterDocId="063AFFF3D34AFF92FFDEFFF1FFE8331E" masterDocTitle="Description of Kashmirobroscus gen. nov. with two new species from the Northwest Himalaya, and remarks on the East Asian genus Eobroscus Kryzhanovskij, 1951 (Coleoptera: Carabidae: Broscini)" masterLastPageNumber="2689" masterPageNumber="2671" pageNumber="2686" updateTime="1698933453466" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2DE7C5C42713D85CAF4E501B16E9DD5B">Description of Kashmirobroscus gen. nov. with two new species from the Northwest Himalaya, and remarks on the East Asian genus Eobroscus Kryzhanovskij, 1951 (Coleoptera: Carabidae: Broscini)</mods:title>
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<mods:namePart id="6B16CF286E3CE9CEF5F149C921FC04FF">Schmidt, Joachim</mods:namePart>
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<mods:affiliation id="1145B1FE9E2501B8F249C3756D9CCDAC">Lindenstrasse 31, 18211 Admannshagen, Germany;</mods:affiliation>
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<mods:namePart id="5ED69A0CF388549A0DFC1C58F8E1CF74">Wrase, David W.</mods:namePart>
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<mods:affiliation id="D3E9793D77445FDC4C05967F00F0EA39">Dunckerstrasse 78, 10437 Berlin, Germany;</mods:affiliation>
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<mods:namePart id="AA13E4883B36C32980E5F73E19583A74">Sciaky, Riccardo</mods:namePart>
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<mods:affiliation id="E20A4EFA32F9DA7990834280325385B3">Via Fiamma 13, 20129 Milano, Italy</mods:affiliation>
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<mods:title id="AF8DAB009F484CEE649B5BE8DB3FCC7C">Journal of Natural History</mods:title>
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<mods:date id="FDCBCD075276B9805002512363BAD76C">2013</mods:date>
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<mods:title id="2314C82CC8B9E6609A219E770ED0DB7D">J. Nat. Hist.</mods:title>
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<mods:number id="C241560A816B1377AAA5059E9A97FB67">41 - 42</mods:number>
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<treatment id="FA03878BD35BFF81FE74FBD5FBC03722" ID-DOI="http://doi.org/10.5281/zenodo.6502457" ID-GBIF-Taxon="194854825" ID-Zenodo-Dep="6502457" LSID="urn:lsid:plazi:treatment:FA03878BD35BFF81FE74FBD5FBC03722" httpUri="http://treatment.plazi.org/id/FA03878BD35BFF81FE74FBD5FBC03722" lastPageId="19" lastPageNumber="2688" pageId="17" pageNumber="2686">
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<subSubSection id="3AB06516D35BFF83FE74FBD5FCDC3722" box="[426,820,1060,1085]" pageId="17" pageNumber="2686" type="nomenclature">
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<paragraph id="7215369DD35BFF83FE74FBD5FCDC3722" blockId="17.[426,820,1060,1085]" box="[426,820,1060,1085]" pageId="17" pageNumber="2686">
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<taxonomicName id="B5AA4D1ED35BFF83FE74FBD5FCDC3722" ID-CoL="6FJ27" authority="Morita, 1990" authorityName="Morita" authorityYear="1990" box="[426,820,1060,1085]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="2686" phylum="Arthropoda" rank="species" species="masumotoi">
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<emphasis id="40DEEA8FD35BFF83FE74FBD5FD713722" bold="true" box="[426,665,1060,1085]" italics="true" pageId="17" pageNumber="2686">Eobroscus masumotoi</emphasis>
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<bibRefCitation id="163B4B6CD35BFF83FD7FFBD5FCDC3722" author="Morita S" box="[673,820,1060,1084]" pageId="17" pageNumber="2686" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita, 1990</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="3AB06516D35BFF83FF51FB95FD103782" pageId="17" pageNumber="2686" type="materials_examined">
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<paragraph id="7215369DD35BFF83FF51FB95FD103782" blockId="17.[143,1103,1124,1180]" pageId="17" pageNumber="2686">
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<materialsCitation id="C2C23CC0D35BFF83FF51FB95FD1C3782" ID-GBIF-Occurrence="3757639322" collectingDate="1995-06-17" collectingDateMax="1995-06-26" collectingDateMin="1995-06-17" collectorName="Dalihod & P. Moravec" country="Taiwan" elevation="2400" location="Alishan" pageId="17" pageNumber="2686" specimenCount="2" specimenCount-female="2">
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<collectingCountry id="0ABD760DD35BFF83FF51FB95FEEB3762" box="[143,259,1124,1148]" name="Taiwan" pageId="17" pageNumber="2686">TAIWAN</collectingCountry>
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:
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<location id="77756046D35BFF83FED1FB95FE8F3762" LSID="urn:lsid:plazi:treatment:FA03878BD35BFF81FE74FBD5FBC03722:77756046D35BFF83FED1FB95FE8F3762" box="[271,359,1124,1148]" country="Taiwan" name="Alishan" pageId="17" pageNumber="2686">Alishan</location>
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[locus typicus],
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<elevation id="F987D1AED35BFF83FDC1FB95FD9D3762" box="[543,629,1124,1148]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="17" pageNumber="2686" unit="m" value="2400.0">2400 m</elevation>
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</quantity>
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,
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<date id="0614105DD35BFF83FD5FFB95FCA83762" box="[641,832,1124,1148]" pageId="17" pageNumber="2686" value="1995-06-17" valueMax="1995-06-26" valueMin="1995-06-17">
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<collectingDate id="1650E9B5D35BFF83FD5FFB95FCA83762" box="[641,832,1124,1148]" pageId="17" pageNumber="2686" value="1995-06-17" valueMax="1995-06-26" valueMin="1995-06-17">17–26 June 1995</collectingDate>
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</date>
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,
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<collectorName id="DF5F534BD35BFF83FC92FB95FC423762" box="[844,938,1124,1148]" pageId="17" pageNumber="2686">Dalihod</collectorName>
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leg. (
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<specimenCount id="64ACFD14D35BFF83FC35FB95FBA33762" box="[1003,1099,1124,1148]" pageId="17" pageNumber="2686" type="female">1 female</specimenCount>
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, cWR); same data but:
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<collectorName id="DF5F534BD35BFF83FE55FB74FDEF3782" box="[395,519,1157,1180]" pageId="17" pageNumber="2686">P. Moravec</collectorName>
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leg. (
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<specimenCount id="64ACFD14D35BFF83FD99FB75FD4D3782" box="[583,677,1156,1180]" pageId="17" pageNumber="2686" type="female">1 female</specimenCount>
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, cSCI)
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</materialsCitation>
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.
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||
</paragraph>
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</subSubSection>
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<subSubSection id="3AB06516D35BFF81FF51FB11FBC03722" lastPageId="19" lastPageNumber="2688" pageId="17" pageNumber="2686" type="discussion">
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<paragraph id="7215369DD35BFF83FF51FB11FE2137E6" blockId="17.[143,1104,1248,1535]" box="[143,457,1248,1272]" pageId="17" pageNumber="2686">
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<heading id="295D81F1D35BFF83FF51FB11FE2137E6" box="[143,457,1248,1272]" fontSize="10" level="3" pageId="17" pageNumber="2686" reason="5">
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||
<emphasis id="40DEEA8FD35BFF83FF51FB11FE2137E6" box="[143,457,1248,1272]" italics="true" pageId="17" pageNumber="2686">Remarks on new synonymies</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="7215369DD35BFF83FF51FAF9FEA636E1" blockId="17.[143,1104,1248,1535]" pageId="17" pageNumber="2686">
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The genus
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<taxonomicName id="B5AA4D1ED35BFF83FECBFAF9FD6A363E" authority="Kryzhanovskij, 1951" authorityName="Kryzhanovskij" authorityYear="1951" box="[277,642,1288,1312]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="2686" phylum="Arthropoda" rank="genus">
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<emphasis id="40DEEA8FD35BFF83FECBFAF9FE6D363E" box="[277,389,1288,1312]" italics="true" pageId="17" pageNumber="2686">Eobroscus</emphasis>
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Kryzhanovskij, 1951
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</taxonomicName>
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was described for
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<taxonomicName id="B5AA4D1ED35BFF83FCB2FAF8FBC6363E" authorityName="Roubal" authorityYear="1928" box="[876,1070,1288,1312]" class="Insecta" family="Carabidae" genus="Broscus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="2686" phylum="Arthropoda" rank="species" species="lutshniki">
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<emphasis id="40DEEA8FD35BFF83FCB2FAF8FBC6363E" box="[876,1070,1288,1312]" italics="true" pageId="17" pageNumber="2686">Broscus lutshniki</emphasis>
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</taxonomicName>
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, a species from the Siberian Far East and
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<collectingCountry id="0ABD760DD35BFF83FDBBFAD8FD42365E" box="[613,682,1321,1344]" name="Japan" pageId="17" pageNumber="2686">Japan</collectingCountry>
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. More than 30 years later a second species,
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<taxonomicName id="B5AA4D1ED35BFF83FF32FAB8FDDA367E" authority="Morvan, 1982" authorityName="Morvan" authorityYear="1982" box="[236,562,1352,1376]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="2686" phylum="Arthropoda" rank="species" species="bhutanensis">
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<emphasis id="40DEEA8FD35BFF83FF32FAB8FE6F367E" box="[236,391,1352,1376]" italics="true" pageId="17" pageNumber="2686">E bhutanensis</emphasis>
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<bibRefCitation id="163B4B6CD35BFF83FE4EFAB8FDDA367E" author="Morvan P" box="[400,562,1352,1376]" pageId="17" pageNumber="2686" pagination="77 - 88" refId="ref7585" refString="Morvan P. 1982. Quatre nouveaux Carabiques du Bhoutan (4 e note) (Coleoptera: Carabidae, Broscini, Pterostichini, Anchomenini). Entomol Basiliensia. 7: 77 - 88." type="journal article" year="1982">Morvan, 1982</bibRefCitation>
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</taxonomicName>
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was described from
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<collectingCountry id="0ABD760DD35BFF83FCFDFAB9FC93367E" box="[803,891,1352,1376]" name="Bhutan" pageId="17" pageNumber="2686">Bhutan</collectingCountry>
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. Still later,
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<bibRefCitation id="163B4B6CD35BFF83FC21FAB9FF3F369E" author="Morita S" pageId="17" pageNumber="2686" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita (1990)</bibRefCitation>
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, when describing a third species (
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<taxonomicName id="B5AA4D1ED35BFF83FD89FA98FD18369E" authorityName="Morita" authorityYear="1990" box="[599,752,1385,1408]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="2686" phylum="Arthropoda" rank="species" species="masumotoi">
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<emphasis id="40DEEA8FD35BFF83FD89FA98FD18369E" box="[599,752,1385,1408]" italics="true" pageId="17" pageNumber="2686">E. masumotoi</emphasis>
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</taxonomicName>
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from Taiwan) created the subgenus
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<taxonomicName id="B5AA4D1ED35BFF83FF04FA79FEBF36BE" authorityName="Morita" authorityYear="1990" box="[218,343,1416,1440]" class="Insecta" family="Carabidae" genus="Orobroscus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="2686" phylum="Arthropoda" rank="genus">
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<emphasis id="40DEEA8FD35BFF83FF04FA79FEBF36BE" box="[218,343,1416,1440]" italics="true" pageId="17" pageNumber="2686">Orobroscus</emphasis>
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</taxonomicName>
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for the two species
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<taxonomicName id="B5AA4D1ED35BFF83FD99FA78FD0336BE" authorityName="Morvan" authorityYear="1982" box="[583,747,1416,1440]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="2686" phylum="Arthropoda" rank="species" species="bhutanensis">
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<emphasis id="40DEEA8FD35BFF83FD99FA78FD0336BE" box="[583,747,1416,1440]" italics="true" pageId="17" pageNumber="2686">E. bhutanensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="B5AA4D1ED35BFF83FCF5FA78FC2F36BE" authorityName="Morita" authorityYear="1990" box="[811,967,1417,1440]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="2686" phylum="Arthropoda" rank="species" species="masumotoi">
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<emphasis id="40DEEA8FD35BFF83FCF5FA78FC2F36BE" box="[811,967,1417,1440]" italics="true" pageId="17" pageNumber="2686">E. masumotoi</emphasis>
|
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</taxonomicName>
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. Finally, in 1995 the same author described a third species of that subgenus,
|
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<taxonomicName id="B5AA4D1ED35BFF83FC89FA58FBA336A1" authority="Morita, 1995" authorityName="Morita" authorityYear="1995" box="[855,1099,1447,1472]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="2686" phylum="Arthropoda" rank="species" species="uenoi">
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<emphasis id="40DEEA8FD35BFF83FC89FA58FC5836DE" box="[855,944,1449,1472]" italics="true" pageId="17" pageNumber="2686">E. uenoi</emphasis>
|
||
<bibRefCitation id="163B4B6CD35BFF83FC68FA56FBA336A1" author="Morita S" box="[950,1099,1447,1471]" pageId="17" pageNumber="2686" pagination="7 - 11" refId="ref7551" refString="Morita S. 1995. A new species of the genus Eobroscus (Coleoptera, Carabidae) from Northern Vietnam. Bull Natl Sci Mus Ser A. 21: 7 - 11." type="journal article" year="1995">Morita, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="0ABD760DD35BFF83FF0CFA36FEDE36C1" box="[210,310,1479,1503]" name="Vietnam" pageId="17" pageNumber="2686">Vietnam</collectingCountry>
|
||
. The distinguishing characters of
|
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<taxonomicName id="B5AA4D1ED35BFF83FD15FA39FCA036FE" authorityName="Morita" authorityYear="1990" box="[715,840,1480,1504]" class="Insecta" family="Carabidae" genus="Orobroscus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="2686" phylum="Arthropoda" rank="genus">
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<emphasis id="40DEEA8FD35BFF83FD15FA39FCA036FE" box="[715,840,1480,1504]" italics="true" pageId="17" pageNumber="2686">Orobroscus</emphasis>
|
||
</taxonomicName>
|
||
mentioned by
|
||
<bibRefCitation id="163B4B6CD35BFF83FC21FA36FEE436E1" author="Morita S" pageId="17" pageNumber="2686" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita (1990: 159)</bibRefCitation>
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were:
|
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</paragraph>
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<paragraph id="7215369DD35BFF83FF64F9D6FEF53541" blockId="17.[179,1103,1575,1695]" pageId="17" pageNumber="2686">(i) the occurrence of two setae at each side of the last abdominal sternite of the male,</paragraph>
|
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<paragraph id="7215369DD35BFF83FF6DF996FE1E3581" blockId="17.[179,1103,1575,1695]" pageId="17" pageNumber="2686">(ii) the presence of adhesive hairs on the proximal two (instead of three) protarsomeres in the male, and</paragraph>
|
||
<paragraph id="7215369DD358FF80FF31FF56FCF533A1" blockId="18.[239,797,167,191]" box="[239,797,167,191]" pageId="18" pageNumber="2687">(iii) the lack of dorsal strigosity of the onychium.</paragraph>
|
||
<paragraph id="7215369DD358FF80FF20FF19FBDE31E3" blockId="18.[211,1172,231,1690]" pageId="18" pageNumber="2687">
|
||
We had the opportunity to examine additional
|
||
<taxonomicName id="B5AA4D1ED358FF80FCF8FF16FC7E33E1" authorityName="Kryzhanovskij" authorityYear="1951" box="[806,918,231,255]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FCF8FF16FC7E33E1" box="[806,918,231,255]" italics="true" pageId="18" pageNumber="2687">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
material covering the whole geographical distribution of that genus and all the known species. As a result we observed the great similarity of the three taxa attributed to
|
||
<taxonomicName id="B5AA4D1ED358FF80FC7CFED6FBF73221" authorityName="Morita" authorityYear="1990" box="[930,1055,295,319]" class="Insecta" family="Carabidae" genus="Orobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FC7CFED6FBF73221" box="[930,1055,295,319]" italics="true" pageId="18" pageNumber="2687">Orobroscus</emphasis>
|
||
</taxonomicName>
|
||
as well as the remarkable variability of the characters used for the separation of the latter taxon. For example, concerning the setosity of the last abdominal sternite we observed that in both the species,
|
||
<taxonomicName id="B5AA4D1ED358FF80FE6CFE79FDA33281" authorityName="Morita" authorityYear="1990" box="[434,587,392,415]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="40DEEA8FD358FF80FE6CFE79FDA33281" box="[434,587,392,415]" italics="true" pageId="18" pageNumber="2687">E. masumotoi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5AA4D1ED358FF80FD5BFE79FCED3281" authorityName="Roubal" authorityYear="1928" box="[645,773,391,415]" class="Insecta" family="Carabidae" genus="Broscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="lutshniki">
|
||
<emphasis id="40DEEA8FD358FF80FD5BFE79FCED3281" box="[645,773,391,415]" italics="true" pageId="18" pageNumber="2687">E. lutshniki</emphasis>
|
||
</taxonomicName>
|
||
, some male specimens had a single seta on one side but two on the opposite side. In most of the
|
||
<taxonomicName id="B5AA4D1ED358FF80FC79FE59FBA132A1" authorityName="Morvan" authorityYear="1982" box="[935,1097,423,447]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD358FF80FC79FE59FBA132A1" box="[935,1097,423,447]" italics="true" pageId="18" pageNumber="2687">E. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
males we found two pairs of setae, but in a specimen from
|
||
<collectingCountry id="0ABD760DD358FF80FCFFFE37FC6932C0" box="[801,897,454,478]" name="Vietnam" pageId="18" pageNumber="2687">Vietnam</collectingCountry>
|
||
we found only one pair. Also the
|
||
<typeStatus id="AD11883FD358FF80FEE9FE17FE7332E0" box="[311,411,486,510]" pageId="18" pageNumber="2687" type="holotype">holotype</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="B5AA4D1ED358FF80FDF5FE16FCC932E0" authority="Morita, 1995" authorityName="Morita" authorityYear="1995" box="[555,801,486,510]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="uenoi">
|
||
<emphasis id="40DEEA8FD358FF80FDF5FE16FD6D32E0" box="[555,645,487,510]" italics="true" pageId="18" pageNumber="2687">E. uenoi</emphasis>
|
||
<bibRefCitation id="163B4B6CD358FF80FD55FE17FCC932E0" author="Morita S" box="[651,801,486,510]" pageId="18" pageNumber="2687" pagination="7 - 11" refId="ref7551" refString="Morita S. 1995. A new species of the genus Eobroscus (Coleoptera, Carabidae) from Northern Vietnam. Bull Natl Sci Mus Ser A. 21: 7 - 11." type="journal article" year="1995">Morita, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
, bears a single pair of setae at the apical margin of the last abdominal sternite. Moreover, we found an
|
||
<taxonomicName id="B5AA4D1ED358FF80FC18FDF6FBAB3100" authorityName="Roubal" authorityYear="1928" box="[966,1091,518,542]" class="Insecta" family="Carabidae" genus="Broscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="lutshniki">
|
||
<emphasis id="40DEEA8FD358FF80FC18FDF6FBAB3100" box="[966,1091,518,542]" italics="true" pageId="18" pageNumber="2687">E. lutshniki</emphasis>
|
||
</taxonomicName>
|
||
female specimen from Sakhalin that bears three setae on the right and two on the left side. Based on these observations we conclude that within the genus
|
||
<taxonomicName id="B5AA4D1ED358FF80FC45FDB7FBE33140" authorityName="Kryzhanovskij" authorityYear="1951" box="[923,1035,582,606]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FC45FDB7FBE33140" box="[923,1035,582,606]" italics="true" pageId="18" pageNumber="2687">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
the number of setae developed at the apical margin of the last abdominal sternite is not a speciesspecific constant. For this reason, abdominal setation seems inappropriate to be used as a differential character to define natural lineages within
|
||
<taxonomicName id="B5AA4D1ED358FF80FCBAFD57FC3C31A0" authorityName="Kryzhanovskij" authorityYear="1951" box="[868,980,678,702]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FCBAFD57FC3C31A0" box="[868,980,678,702]" italics="true" pageId="18" pageNumber="2687">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
. This conclusion was already reached by
|
||
<bibRefCitation id="163B4B6CD358FF80FE02FD34FD4B31C3" author="Morita S" box="[476,675,709,733]" pageId="18" pageNumber="2687" pagination="7 - 11" refId="ref7551" refString="Morita S. 1995. A new species of the genus Eobroscus (Coleoptera, Carabidae) from Northern Vietnam. Bull Natl Sci Mus Ser A. 21: 7 - 11." type="journal article" year="1995">Morita (1995: 11)</bibRefCitation>
|
||
in the paper with the description of
|
||
<taxonomicName id="B5AA4D1ED358FF80FBE7FD36FE8231E3" authority="Morita, 1995" authorityName="Morita" authorityYear="1995" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="uenoi">
|
||
<emphasis id="40DEEA8FD358FF80FBE7FD36FB7C31C3" box="[1081,1172,710,734]" italics="true" pageId="18" pageNumber="2687">E. uenoi</emphasis>
|
||
<bibRefCitation id="163B4B6CD358FF80FF0DFD14FE8231E3" author="Morita S" box="[211,362,741,765]" pageId="18" pageNumber="2687" pagination="7 - 11" refId="ref7551" refString="Morita S. 1995. A new species of the genus Eobroscus (Coleoptera, Carabidae) from Northern Vietnam. Bull Natl Sci Mus Ser A. 21: 7 - 11." type="journal article" year="1995">Morita, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
; however, the author still maintained the validity of
|
||
<taxonomicName id="B5AA4D1ED358FF80FC6CFD17FBC731E0" authorityName="Morita" authorityYear="1990" box="[946,1071,742,766]" class="Insecta" family="Carabidae" genus="Orobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FC6CFD17FBC731E0" box="[946,1071,742,766]" italics="true" pageId="18" pageNumber="2687">Orobroscus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7215369DD358FF80FF20FCF4FE8F3782" blockId="18.[211,1172,231,1690]" pageId="18" pageNumber="2687">
|
||
Concerning protarsal adhesive vestiture of males we confirm the observation of
|
||
<bibRefCitation id="163B4B6CD358FF80FF0DFCD4FE983023" author="Morita S" box="[211,368,805,829]" pageId="18" pageNumber="2687" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita (1990)</bibRefCitation>
|
||
who found adhesive vestiture on the proximal three tarsomeres of
|
||
<taxonomicName id="B5AA4D1ED358FF80FB91FCD7FEFC3043" authorityName="Roubal" authorityYear="1928" class="Insecta" family="Carabidae" genus="Broscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="lutshniki">
|
||
<emphasis id="40DEEA8FD358FF80FB91FCD7FEFC3043" italics="true" pageId="18" pageNumber="2687">E. lutshniki</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="EA912A18D358FF80FEFBFCB4FE723043" box="[293,410,837,861]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="18" pageNumber="2687">Figure 4D</figureCitation>
|
||
) and on the proximal two tarsomeres of
|
||
<taxonomicName id="B5AA4D1ED358FF80FCB8FCB7FBEE3043" authorityName="Morvan" authorityYear="1982" box="[870,1030,837,861]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD358FF80FCB8FCB7FBEE3043" box="[870,1030,837,861]" italics="true" pageId="18" pageNumber="2687">E. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="EA912A18D358FF80FBC8FCB4FB623043" box="[1046,1162,837,861]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="18" pageNumber="2687">Figure 4G</figureCitation>
|
||
) and of
|
||
<taxonomicName id="B5AA4D1ED358FF80FEF8FC97FE283063" authorityName="Morita" authorityYear="1990" box="[294,448,870,893]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="40DEEA8FD358FF80FEF8FC97FE283063" box="[294,448,870,893]" italics="true" pageId="18" pageNumber="2687">E. masumotoi</emphasis>
|
||
</taxonomicName>
|
||
. However, as a result of a comprehensive phylogenetic analysis of the tribe
|
||
<taxonomicName id="B5AA4D1ED358FF80FE8BFC74FD663083" authority="Roig-Junent (2000)" authorityName="Roig-Junent" authorityYear="2000" box="[341,654,901,925]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="tribe" tribe="Broscini">
|
||
Broscini
|
||
<bibRefCitation id="163B4B6CD358FF80FE66FC74FD663083" author="Roig-Junent SA" box="[440,654,901,925]" pageId="18" pageNumber="2687" pagination="1 - 90" refId="ref7621" refString="Roig-Junent SA. 2000. The subtribes and genera of the tribe Broscini (Coleoptera: Carabidae): cladistic analysis, taxonomic treatment and biogeographical considerations. B Am Mus Nat Hist. 255: 1 - 90." type="journal article" year="2000">Roig-Juñent (2000)</bibRefCitation>
|
||
</taxonomicName>
|
||
showed that this character has several parallel gains and losses, and it is therefore of low phylogenetic value (zero in the cladistic analysis of
|
||
<bibRefCitation id="163B4B6CD358FF80FEFAFC34FE0330C3" author="Roig-Junent SA" box="[292,491,965,989]" pageId="18" pageNumber="2687" pagination="1 - 90" refId="ref7621" refString="Roig-Junent SA. 2000. The subtribes and genera of the tribe Broscini (Coleoptera: Carabidae): cladistic analysis, taxonomic treatment and biogeographical considerations. B Am Mus Nat Hist. 255: 1 - 90." type="journal article" year="2000">Roig-Juñent 2000</bibRefCitation>
|
||
). Species of the same genus and even of the same subgenus have or lack this structure. Those conditions
|
||
<bibRefCitation id="163B4B6CD358FF80FD12FC15FC4B30E2" author="Roig-Junent SA" box="[716,931,996,1020]" pageId="18" pageNumber="2687" pagination="1 - 90" refId="ref7621" refString="Roig-Junent SA. 2000. The subtribes and genera of the tribe Broscini (Coleoptera: Carabidae): cladistic analysis, taxonomic treatment and biogeographical considerations. B Am Mus Nat Hist. 255: 1 - 90." type="journal article" year="2000">Roig-Juñent (2000)</bibRefCitation>
|
||
found also regarding the mesotarsal adhesive vestiture. The different extent of adhesive vestiture on male mesotarsomeres in
|
||
<taxonomicName id="B5AA4D1ED358FF80FE6DFBD4FDCB3723" authorityName="Kryzhanovskij" authorityYear="1951" box="[435,547,1061,1085]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FE6DFBD4FDCB3723" box="[435,547,1061,1085]" italics="true" pageId="18" pageNumber="2687">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
species is shown in
|
||
<figureCitation id="EA912A18D358FF80FCC9FBD5FC633722" box="[791,907,1060,1084]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="18" pageNumber="2687">Figure 4E</figureCitation>
|
||
(
|
||
<taxonomicName id="B5AA4D1ED358FF80FC41FBD4FBC93723" authorityName="Roubal" authorityYear="1928" box="[927,1057,1061,1085]" class="Insecta" family="Carabidae" genus="Broscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="lutshniki">
|
||
<emphasis id="40DEEA8FD358FF80FC41FBD4FBC93723" box="[927,1057,1061,1085]" italics="true" pageId="18" pageNumber="2687">E. lutshniki</emphasis>
|
||
</taxonomicName>
|
||
, adhesive vestiture covers approx. 75% of the ventral surface of tarsomere 2) and
|
||
<figureCitation id="EA912A18D358FF80FBC2FBB5FB7B3742" box="[1052,1171,1092,1116]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="18" pageNumber="2687">Figure 4H</figureCitation>
|
||
(
|
||
<taxonomicName id="B5AA4D1ED358FF80FF02FB94FE693762" authorityName="Morvan" authorityYear="1982" box="[220,385,1124,1148]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD358FF80FF02FB94FE693762" box="[220,385,1124,1148]" italics="true" pageId="18" pageNumber="2687">E. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
, adhesive vestiture covers less than 50% of the ventral surface of tarsomere 2).
|
||
</paragraph>
|
||
<paragraph id="7215369DD358FF80FF20FB55FD0E3625" blockId="18.[211,1172,231,1690]" pageId="18" pageNumber="2687">
|
||
Concerning the surface morphology of the onychium we found traces of dorsal longitudinal strigosity also in
|
||
<taxonomicName id="B5AA4D1ED358FF80FDFFFB34FD2937C2" authorityName="Morvan" authorityYear="1982" box="[545,705,1220,1244]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD358FF80FDFFFB34FD2937C2" box="[545,705,1220,1244]" italics="true" pageId="18" pageNumber="2687">E. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="EA912A18D358FF80FD29FB35FCB737C2" box="[759,863,1220,1244]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="18" pageNumber="2687">Figure 4I</figureCitation>
|
||
) and in
|
||
<taxonomicName id="B5AA4D1ED358FF80FC62FB34FBBB37C2" authorityName="Morita" authorityYear="1990" box="[956,1107,1221,1244]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="40DEEA8FD358FF80FC62FB34FBBB37C2" box="[956,1107,1221,1244]" italics="true" pageId="18" pageNumber="2687">E. masumotoi</emphasis>
|
||
</taxonomicName>
|
||
. This is in contrast to the statement of
|
||
<bibRefCitation id="163B4B6CD358FF80FD9BFB15FCF137E2" author="Morita S" box="[581,793,1252,1276]" pageId="18" pageNumber="2687" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita (1990: 156)</bibRefCitation>
|
||
that members of
|
||
<taxonomicName id="B5AA4D1ED358FF80FC03FB15FBB237E2" authorityName="Morita" authorityYear="1990" box="[989,1114,1252,1276]" class="Insecta" family="Carabidae" genus="Orobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FC03FB15FBB237E2" box="[989,1114,1252,1276]" italics="true" pageId="18" pageNumber="2687">Orobroscus</emphasis>
|
||
</taxonomicName>
|
||
have a smooth onychium dorsally. It is true that strigosity of the onychium is much more markedly developed in
|
||
<taxonomicName id="B5AA4D1ED358FF80FE09FAD5FDBE3622" authorityName="Roubal" authorityYear="1928" box="[471,598,1316,1340]" class="Insecta" family="Carabidae" genus="Broscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="lutshniki">
|
||
<emphasis id="40DEEA8FD358FF80FE09FAD5FDBE3622" box="[471,598,1316,1340]" italics="true" pageId="18" pageNumber="2687">E. lutshniki</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="EA912A18D358FF80FDBBFAD2FD3E3625" box="[613,726,1315,1339]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="18" pageNumber="2687">Figure 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7215369DD358FF80FF20FAB5FCAB3685" blockId="18.[211,1172,231,1690]" pageId="18" pageNumber="2687">
|
||
In summary, we state that separation of a subgenus
|
||
<taxonomicName id="B5AA4D1ED358FF80FCB1FAB5FC043642" authorityName="Morita" authorityYear="1990" box="[879,1004,1348,1372]" class="Insecta" family="Carabidae" genus="Orobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FCB1FAB5FC043642" box="[879,1004,1348,1372]" italics="true" pageId="18" pageNumber="2687">Orobroscus</emphasis>
|
||
</taxonomicName>
|
||
as defined by
|
||
<bibRefCitation id="163B4B6CD358FF80FF0DFA92FE9A3665" author="Morita S" box="[211,370,1379,1403]" pageId="18" pageNumber="2687" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita (1990)</bibRefCitation>
|
||
is highly questionable. Therefore, we propose
|
||
<taxonomicName id="B5AA4D1ED358FF80FCA5FA92FB7B3665" authority="Morita, 1990" authorityName="Morita" authorityYear="1990" box="[891,1171,1379,1403]" class="Insecta" family="Carabidae" genus="Orobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FCA5FA92FC103665" box="[891,1016,1379,1403]" italics="true" pageId="18" pageNumber="2687">Orobroscus</emphasis>
|
||
<bibRefCitation id="163B4B6CD358FF80FC21FA92FB7B3665" author="Morita S" box="[1023,1171,1379,1403]" pageId="18" pageNumber="2687" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName id="B5AA4D1ED358FF80FE05FA72FCD73685" authority="Kryzhanovskij, 1951" authorityName="Kryzhanovskij" authorityYear="1951" box="[475,831,1411,1435]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FE05FA72FDA33685" box="[475,587,1411,1435]" italics="true" pageId="18" pageNumber="2687">Eobroscus</emphasis>
|
||
Kryzhanovskij, 1951
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7215369DD358FF81FF20FA52FE2B3281" blockId="18.[211,1172,231,1690]" lastBlockId="19.[143,1104,167,1084]" lastPageId="19" lastPageNumber="2688" pageId="18" pageNumber="2687">
|
||
This decision is supported by our observation that species diversity in the genus
|
||
<taxonomicName id="B5AA4D1ED358FF80FF0DFA32FEAB36C5" authorityName="Kryzhanovskij" authorityYear="1951" box="[211,323,1475,1499]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD358FF80FF0DFA32FEAB36C5" box="[211,323,1475,1499]" italics="true" pageId="18" pageNumber="2687">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
is lower than expected: We could compare several specimens from
|
||
<collectingCountry id="0ABD760DD358FF80FBEDFA33FB7B36C4" box="[1075,1171,1474,1498]" name="Vietnam" pageId="18" pageNumber="2687">Vietnam</collectingCountry>
|
||
(coming from the type locality of
|
||
<taxonomicName id="B5AA4D1ED358FF80FD97FA12FCDE36E4" authority="Morita 1995" authorityName="Morita" authorityYear="1995" box="[585,822,1506,1530]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="uenoi">
|
||
<emphasis id="40DEEA8FD358FF80FD97FA12FD4C36E4" box="[585,676,1507,1530]" italics="true" pageId="18" pageNumber="2687">E. uenoi</emphasis>
|
||
<bibRefCitation id="163B4B6CD358FF80FD74FA13FCDE36E4" author="Morita S" box="[682,822,1506,1530]" pageId="18" pageNumber="2687" pagination="7 - 11" refId="ref7551" refString="Morita S. 1995. A new species of the genus Eobroscus (Coleoptera, Carabidae) from Northern Vietnam. Bull Natl Sci Mus Ser A. 21: 7 - 11." type="journal article" year="1995">Morita 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
) with the
|
||
<typeStatus id="AD11883FD358FF80FC78FA13FBE236E4" box="[934,1034,1506,1530]" pageId="18" pageNumber="2687" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="B5AA4D1ED358FF80FBF3FA12FE2C3504" authority="Morvan, 1982" authorityName="Morvan" authorityYear="1982" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD358FF80FBF3FA12FEFD3504" italics="true" pageId="18" pageNumber="2687">E. bhutanensis</emphasis>
|
||
<bibRefCitation id="163B4B6CD358FF80FEC1F9F2FE2C3504" author="Morvan P" box="[287,452,1538,1562]" pageId="18" pageNumber="2687" pagination="77 - 88" refId="ref7585" refString="Morvan P. 1982. Quatre nouveaux Carabiques du Bhoutan (4 e note) (Coleoptera: Carabidae, Broscini, Pterostichini, Anchomenini). Entomol Basiliensia. 7: 77 - 88." type="journal article" year="1982">Morvan, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
, and with additional specimens from
|
||
<collectingCountry id="0ABD760DD358FF80FCA4F9F3FC2A3504" box="[890,962,1538,1562]" name="China" pageId="18" pageNumber="2687">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="0ABD760DD358FF80FC0EF9F2FBA93505" box="[976,1089,1539,1563]" name="Myanmar" pageId="18" pageNumber="2687">Myanmar</collectingCountry>
|
||
,
|
||
<collectingCountry id="0ABD760DD358FF80FB91F9F3FB7C3504" box="[1103,1172,1538,1562]" name="Nepal" pageId="18" pageNumber="2687">Nepal</collectingCountry>
|
||
and northeast
|
||
<collectingCountry id="0ABD760DD358FF80FEA2F9D3FE533524" box="[380,443,1570,1594]" name="India" pageId="18" pageNumber="2687">India</collectingCountry>
|
||
, and we could determine identity. Based on a careful examination of this material we can state that all differential characters of
|
||
<taxonomicName id="B5AA4D1ED358FF80FC64F9B2FBFC3544" authorityName="Morita" authorityYear="1995" box="[954,1044,1603,1626]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="uenoi">
|
||
<emphasis id="40DEEA8FD358FF80FC64F9B2FBFC3544" box="[954,1044,1603,1626]" italics="true" pageId="18" pageNumber="2687">E. uenoi</emphasis>
|
||
</taxonomicName>
|
||
mentioned by
|
||
<bibRefCitation id="163B4B6CD358FF80FF24F993FE743564" author="Morita S" box="[250,412,1634,1658]" pageId="18" pageNumber="2687" pagination="7 - 11" refId="ref7551" refString="Morita S. 1995. A new species of the genus Eobroscus (Coleoptera, Carabidae) from Northern Vietnam. Bull Natl Sci Mus Ser A. 21: 7 - 11." type="journal article" year="1995">Morita (1995)</bibRefCitation>
|
||
fall in the variability of the species
|
||
<taxonomicName id="B5AA4D1ED358FF80FC92F992FC193564" authorityName="Morvan" authorityYear="1982" box="[844,1009,1634,1658]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD358FF80FC92F992FC193564" box="[844,1009,1634,1658]" italics="true" pageId="18" pageNumber="2687">E. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
: (i) body size
|
||
<quantity id="B5529B78D358FF80FF0DF973FE993584" box="[211,369,1666,1690]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" metricValueMax="1.61" metricValueMin="1.5699999999999998" pageId="18" pageNumber="2687" unit="mm" value="15.9" valueMax="16.1" valueMin="15.7">15.7–16.1 mm</quantity>
|
||
(
|
||
<quantity id="B5529B78D358FF80FEA1F973FDF63584" box="[383,542,1666,1690]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" metricValueMax="1.63" metricValueMin="1.59" pageId="18" pageNumber="2687" unit="mm" value="16.1" valueMax="16.3" valueMin="15.9">15.9–16.3 mm</quantity>
|
||
in
|
||
<taxonomicName id="B5AA4D1ED358FF80FD9EF972FD733584" authorityName="Morita" authorityYear="1995" box="[576,667,1667,1690]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2687" phylum="Arthropoda" rank="species" species="uenoi">
|
||
<emphasis id="40DEEA8FD358FF80FD9EF972FD733584" box="[576,667,1667,1690]" italics="true" pageId="18" pageNumber="2687">E. uenoi</emphasis>
|
||
</taxonomicName>
|
||
); (ii) coloration of dorsal side of body brown to black with +
|
||
<emphasis id="40DEEA8FD359FF81FE95FF59FEBF33DE" bold="true" box="[331,343,168,192]" italics="true" pageId="19" pageNumber="2688">/</emphasis>
|
||
– prominent metallic lustre (see
|
||
<figureCitation id="EA912A18D359FF81FD0FFF56FCAD33A1" box="[721,837,167,191]" captionStart="Figure 2" captionStartId="7.[143,210,1380,1402]" captionTargetBox="[145,1104,171,1346]" captionTargetId="figure-5@7.[144,1104,168,1347]" captionTargetPageId="7" captionText="Figure 2. Kashmirobroscus and Eobroscus in dorsal view. (A) Kashmirobroscus mirabilis sp. nov., holotype; (B) Eobroscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal." figureDoi="http://doi.org/10.5281/zenodo.4608241" httpUri="https://zenodo.org/record/4608241/files/figure.png" pageId="19" pageNumber="2688">Figure 2B</figureCitation>
|
||
; body almost black in
|
||
<taxonomicName id="B5AA4D1ED359FF81FF51FF39FF0233C1" authorityName="Morita" authorityYear="1995" box="[143,234,200,223]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="uenoi">
|
||
<emphasis id="40DEEA8FD359FF81FF51FF39FF0233C1" box="[143,234,200,223]" italics="true" pageId="19" pageNumber="2688">E. uenoi</emphasis>
|
||
</taxonomicName>
|
||
with greenish lustre on elytra); (iii) aedeagus relatively robust (see
|
||
<figureCitation id="EA912A18D359FF81FC0FFF36FBC633C1" box="[977,1070,199,223]" captionStart="Figure 5" captionStartId="11.[143,210,1350,1372]" captionTargetBox="[144,1103,170,1317]" captionTargetId="figure-5@11.[144,1104,167,1317]" captionTargetPageId="11" captionText="Figure 5. Aedeagal median lobe in left lateral view (A, C, E, G; H shows a cut-out of G) and in dorsal view (B, D, F, I; B and I additionally with parameres), and sternum IX (J) of Kashmirobroscus and Eobroscus. (A, B) Eobroscus lutshniki (Roubal, 1928), specimen from Ussurisk, Russia; (C–F) Eobroscus bhutanensis Morvan, 1982 (C, D, specimen from Qingcheng Houshan, China; E, F, specimen from Mt Fansipan, Vietnam); (G–J), Kashmirobroscus ortrudae sp. nov., paratype, male. Abbreviations: lap, latero-apical projections of internal sac; pl, left paramere; pr, right paramere." figureDoi="http://doi.org/10.5281/zenodo.4608247" httpUri="https://zenodo.org/record/4608247/files/figure.png" pageId="19" pageNumber="2688">Figure 5</figureCitation>
|
||
C– F; identical in
|
||
<taxonomicName id="B5AA4D1ED359FF81FEF0FF19FE6F33E1" authorityName="Morita" authorityYear="1995" box="[302,391,232,255]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="uenoi">
|
||
<emphasis id="40DEEA8FD359FF81FEF0FF19FE6F33E1" box="[302,391,232,255]" italics="true" pageId="19" pageNumber="2688">E. uenoi</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="163B4B6CD359FF81FE65FF16FD8933E1" author="Morita S" box="[443,609,231,255]" pageId="19" pageNumber="2688" pagination="7 - 11" refId="ref7551" refString="Morita S. 1995. A new species of the genus Eobroscus (Coleoptera, Carabidae) from Northern Vietnam. Bull Natl Sci Mus Ser A. 21: 7 - 11." type="journal article" year="1995">Morita 1995: 9</bibRefCitation>
|
||
); (iv) lateral and dorsal wing-like expansions of aedeagal median lobe markedly produced (see
|
||
<figureCitation id="EA912A18D359FF81FD1DFEF6FCCB3201" box="[707,803,263,287]" captionStart="Figure 5" captionStartId="11.[143,210,1350,1372]" captionTargetBox="[144,1103,170,1317]" captionTargetId="figure-5@11.[144,1104,167,1317]" captionTargetPageId="11" captionText="Figure 5. Aedeagal median lobe in left lateral view (A, C, E, G; H shows a cut-out of G) and in dorsal view (B, D, F, I; B and I additionally with parameres), and sternum IX (J) of Kashmirobroscus and Eobroscus. (A, B) Eobroscus lutshniki (Roubal, 1928), specimen from Ussurisk, Russia; (C–F) Eobroscus bhutanensis Morvan, 1982 (C, D, specimen from Qingcheng Houshan, China; E, F, specimen from Mt Fansipan, Vietnam); (G–J), Kashmirobroscus ortrudae sp. nov., paratype, male. Abbreviations: lap, latero-apical projections of internal sac; pl, left paramere; pr, right paramere." figureDoi="http://doi.org/10.5281/zenodo.4608247" httpUri="https://zenodo.org/record/4608247/files/figure.png" pageId="19" pageNumber="2688">Figure 5</figureCitation>
|
||
C–F; identical in
|
||
<taxonomicName id="B5AA4D1ED359FF81FC32FEF9FBA03201" authorityName="Morita" authorityYear="1995" box="[1004,1096,264,287]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="uenoi">
|
||
<emphasis id="40DEEA8FD359FF81FC32FEF9FBA03201" box="[1004,1096,264,287]" italics="true" pageId="19" pageNumber="2688">E. uenoi</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="163B4B6CD359FF81FF66FED6FE8B3221" author="Morita S" box="[184,355,295,319]" pageId="19" pageNumber="2688" pagination="7 - 11" refId="ref7551" refString="Morita S. 1995. A new species of the genus Eobroscus (Coleoptera, Carabidae) from Northern Vietnam. Bull Natl Sci Mus Ser A. 21: 7 - 11." type="journal article" year="1995">Morita 1995: 9</bibRefCitation>
|
||
). Therefore, we propose
|
||
<taxonomicName id="B5AA4D1ED359FF81FDA7FED9FD3C3221" authorityName="Morita" authorityYear="1995" box="[633,724,296,319]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="uenoi">
|
||
<emphasis id="40DEEA8FD359FF81FDA7FED9FD3C3221" box="[633,724,296,319]" italics="true" pageId="19" pageNumber="2688">E. uenoi</emphasis>
|
||
</taxonomicName>
|
||
mentioned by
|
||
<bibRefCitation id="163B4B6CD359FF81FCA3FED6FBF43221" author="Morita S" box="[893,1052,295,319]" pageId="19" pageNumber="2688" pagination="7 - 11" refId="ref7551" refString="Morita S. 1995. A new species of the genus Eobroscus (Coleoptera, Carabidae) from Northern Vietnam. Bull Natl Sci Mus Ser A. 21: 7 - 11." type="journal article" year="1995">Morita (1995)</bibRefCitation>
|
||
as a junior synonym of
|
||
<taxonomicName id="B5AA4D1ED359FF81FEB6FEB9FD5B3241" authority="Morvan, 1982" authorityName="Morvan" authorityYear="1982" box="[360,691,327,351]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD359FF81FEB6FEB9FDE13241" box="[360,521,327,351]" italics="true" pageId="19" pageNumber="2688">E. bhutanensis</emphasis>
|
||
<bibRefCitation id="163B4B6CD359FF81FDCFFEB9FD5B3241" author="Morvan P" box="[529,691,327,351]" pageId="19" pageNumber="2688" pagination="77 - 88" refId="ref7585" refString="Morvan P. 1982. Quatre nouveaux Carabiques du Bhoutan (4 e note) (Coleoptera: Carabidae, Broscini, Pterostichini, Anchomenini). Entomol Basiliensia. 7: 77 - 88." type="journal article" year="1982">Morvan, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
, a macropterous species with a wide distribution along the Himalayan chain and in neighbouring mountainous areas of South
|
||
<collectingCountry id="0ABD760DD359FF81FF07FE76FEAD3281" box="[217,325,391,415]" name="China" pageId="19" pageNumber="2688">East Asia</collectingCountry>
|
||
(
|
||
<figureCitation id="EA912A18D359FF81FE8BFE76FE5B3280" box="[341,435,391,415]" captionStart="Figure 7" captionStartId="14.[211,278,1572,1594]" captionTargetBox="[215,1172,870,1537]" captionTargetId="figure-151@14.[212,1172,866,1540]" captionTargetPageId="14" captionText="Figure 7. Distribution of Kashmirobroscus ortrudae sp. nov. (white square), Kashmirobroscus mirabilis sp. nov. (white triangle) and Eobroscus bhutanensis Morvan, 1982 (white circles) in High Asia." figureDoi="http://doi.org/10.5281/zenodo.4608251" httpUri="https://zenodo.org/record/4608251/files/figure.png" pageId="19" pageNumber="2688">Figure 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7215369DD359FF81FF64FE56FEE132E0" blockId="19.[143,1104,167,1084]" pageId="19" pageNumber="2688">
|
||
Hence, the genus
|
||
<taxonomicName id="B5AA4D1ED359FF81FE53FE56FE1532A1" authorityName="Kryzhanovskij" authorityYear="1951" box="[397,509,423,447]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD359FF81FE53FE56FE1532A1" box="[397,509,423,447]" italics="true" pageId="19" pageNumber="2688">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
encompasses actually no more than three species. We believe that this is an additional argument against splitting the genus into two subgenera.
|
||
</paragraph>
|
||
<paragraph id="7215369DD359FF81FF64FDF6FBC03722" blockId="19.[143,1104,167,1084]" pageId="19" pageNumber="2688">
|
||
In addition, we have doubts about the status of
|
||
<taxonomicName id="B5AA4D1ED359FF81FD1AFDF6FC1A3100" authority="Morita, 1990" authorityName="Morita" authorityYear="1990" box="[708,1010,518,542]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="40DEEA8FD359FF81FD1AFDF6FCB23100" box="[708,858,519,542]" italics="true" pageId="19" pageNumber="2688">E. masumotoi</emphasis>
|
||
<bibRefCitation id="163B4B6CD359FF81FCBEFDF7FC1A3100" author="Morita S" box="[864,1010,518,542]" pageId="19" pageNumber="2688" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
as a distinct species. Based on the comparison of two female specimens from the type locality with the
|
||
<taxonomicName id="B5AA4D1ED359FF81FF2DFDB6FE7B3140" authorityName="Morvan" authorityYear="1982" box="[243,403,582,606]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD359FF81FF2DFDB6FE7B3140" box="[243,403,582,606]" italics="true" pageId="19" pageNumber="2688">E. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
material listed above we have to reject four of the six differential characters mentioned by
|
||
<bibRefCitation id="163B4B6CD359FF81FE0FFD97FD4C3160" author="Morita S" box="[465,676,614,638]" pageId="19" pageNumber="2688" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita (1990: 164)</bibRefCitation>
|
||
because they fall within the variability of the species
|
||
<taxonomicName id="B5AA4D1ED359FF81FEEEFD76FE393180" authorityName="Morvan" authorityYear="1982" box="[304,465,646,670]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD359FF81FEEEFD76FE393180" box="[304,465,646,670]" italics="true" pageId="19" pageNumber="2688">E. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
: (i) the colour of the body surface; (ii) the length of the temples in proportion to the length of the eyes; (iii) the shape of the pronotum; (iv) the shape of the elytra. We have to confirm the observation of
|
||
<bibRefCitation id="163B4B6CD359FF81FC9EFD34FC3631C3" author="Morita S" box="[832,990,709,733]" pageId="19" pageNumber="2688" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita (1990)</bibRefCitation>
|
||
that specimens of the Taiwanese population (
|
||
<taxonomicName id="B5AA4D1ED359FF81FDEEFD17FD2231E3" authorityName="Morita" authorityYear="1990" box="[560,714,742,765]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="40DEEA8FD359FF81FDEEFD17FD2231E3" box="[560,714,742,765]" italics="true" pageId="19" pageNumber="2688">E. masumotoi</emphasis>
|
||
</taxonomicName>
|
||
) are somewhat smaller (
|
||
<quantity id="B5529B78D359FF81FC3DFD14FBA331E3" box="[995,1099,741,765]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="19" pageNumber="2688" unit="mm" value="14.9">14.9 mm</quantity>
|
||
;
|
||
<bibRefCitation id="163B4B6CD359FF81FF51FCF4FEB43003" author="Morita S" box="[143,348,773,797]" pageId="19" pageNumber="2688" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita 1990: 160</bibRefCitation>
|
||
:
|
||
<quantity id="B5529B78D359FF81FEB7FCF4FDE33003" box="[361,523,773,797]" metricMagnitude="-2" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.52" metricValueMin="1.45" pageId="19" pageNumber="2688" unit="mm" value="14.85" valueMax="15.2" valueMin="14.5">14.5–15.2 mm</quantity>
|
||
) than specimens from the continent (
|
||
<taxonomicName id="B5AA4D1ED359FF81FC1EFCF7FF433023" authorityName="Morvan" authorityYear="1982" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD359FF81FC1EFCF7FF433023" italics="true" pageId="19" pageNumber="2688">E. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="B5529B78D359FF81FF63FCD4FE8B3023" box="[189,355,805,829]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" metricValueMax="1.61" metricValueMin="1.5699999999999998" pageId="19" pageNumber="2688" unit="mm" value="15.9" valueMax="16.1" valueMin="15.7">15.7–16.1 mm</quantity>
|
||
). By comparing our preparations of
|
||
<taxonomicName id="B5AA4D1ED359FF81FCCAFCD7FC5F3023" authorityName="Morvan" authorityYear="1982" box="[788,951,805,829]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD359FF81FCCAFCD7FC5F3023" box="[788,951,805,829]" italics="true" pageId="19" pageNumber="2688">E. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
aedeagi (e.g.
|
||
<figureCitation id="EA912A18D359FF81FF51FCB4FF063043" box="[143,238,837,861]" captionStart="Figure 5" captionStartId="11.[143,210,1350,1372]" captionTargetBox="[144,1103,170,1317]" captionTargetId="figure-5@11.[144,1104,167,1317]" captionTargetPageId="11" captionText="Figure 5. Aedeagal median lobe in left lateral view (A, C, E, G; H shows a cut-out of G) and in dorsal view (B, D, F, I; B and I additionally with parameres), and sternum IX (J) of Kashmirobroscus and Eobroscus. (A, B) Eobroscus lutshniki (Roubal, 1928), specimen from Ussurisk, Russia; (C–F) Eobroscus bhutanensis Morvan, 1982 (C, D, specimen from Qingcheng Houshan, China; E, F, specimen from Mt Fansipan, Vietnam); (G–J), Kashmirobroscus ortrudae sp. nov., paratype, male. Abbreviations: lap, latero-apical projections of internal sac; pl, left paramere; pr, right paramere." figureDoi="http://doi.org/10.5281/zenodo.4608247" httpUri="https://zenodo.org/record/4608247/files/figure.png" pageId="19" pageNumber="2688">Figure 5</figureCitation>
|
||
C–F) with the figuring of the
|
||
<taxonomicName id="B5AA4D1ED359FF81FDE4FCB7FD3A3043" authorityName="Morita" authorityYear="1990" box="[570,722,838,861]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="40DEEA8FD359FF81FDE4FCB7FD3A3043" box="[570,722,838,861]" italics="true" pageId="19" pageNumber="2688">E. masumotoi</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="AD11883FD359FF81FD04FCB4FCD63043" box="[730,830,837,861]" pageId="19" pageNumber="2688" type="holotype">holotype</typeStatus>
|
||
specimen (
|
||
<bibRefCitation id="163B4B6CD359FF81FC64FCB4FF503063" author="Morita S" pageId="19" pageNumber="2688" pagination="155 - 165" refId="ref7524" refString="Morita S. 1990. Broscine carabid beetles of the genus Eobroscus (Coleoptera, Carabidae). Elytra Tokyo. 18: 155 - 165." type="journal article" year="1990">Morita 1990: 163</bibRefCitation>
|
||
) we also confirm that the median lobe of the latter is slightly shorter. In addition, lateral wing-like expansion on the right seems lesser developed in the
|
||
<taxonomicName id="B5AA4D1ED359FF81FC6BFC77FBA73083" authorityName="Morita" authorityYear="1990" box="[949,1103,902,925]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="40DEEA8FD359FF81FC6BFC77FBA73083" box="[949,1103,902,925]" italics="true" pageId="19" pageNumber="2688">E. masumotoi</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="AD11883FD359FF81FF51FC54FF1B30A3" box="[143,243,933,957]" pageId="19" pageNumber="2688" type="holotype">holotype</typeStatus>
|
||
than in specimens of
|
||
<taxonomicName id="B5AA4D1ED359FF81FE2CFC57FD7B30A3" authorityName="Morvan" authorityYear="1982" box="[498,659,933,957]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD359FF81FE2CFC57FD7B30A3" box="[498,659,933,957]" italics="true" pageId="19" pageNumber="2688">E. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
. However, apart from the
|
||
<typeStatus id="AD11883FD359FF81FC1BFC54FBC130A3" box="[965,1065,933,957]" pageId="19" pageNumber="2688" type="holotype">holotype</typeStatus>
|
||
no additional male specimens of
|
||
<taxonomicName id="B5AA4D1ED359FF81FE37FC37FD6C30C3" authorityName="Morita" authorityYear="1990" box="[489,644,966,989]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="40DEEA8FD359FF81FE37FC37FD6C30C3" box="[489,644,966,989]" italics="true" pageId="19" pageNumber="2688">E. masumotoi</emphasis>
|
||
</taxonomicName>
|
||
have been described or recorded in the literature and so, the true variability of the external shape of the aedeagus remains unknown for us. For this reason the decision to confirm or to reject the species status of
|
||
<taxonomicName id="B5AA4D1ED359FF81FF73FBD4FEAD3722" authorityName="Morita" authorityYear="1990" box="[173,325,1061,1084]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2688" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="40DEEA8FD359FF81FF73FBD4FEAD3722" box="[173,325,1061,1084]" italics="true" pageId="19" pageNumber="2688">E. masumotoi</emphasis>
|
||
</taxonomicName>
|
||
has to wait until further material from
|
||
<collectingCountry id="0ABD760DD359FF81FCDEFBD5FCBB3722" box="[768,851,1060,1084]" name="Taiwan" pageId="19" pageNumber="2688">Taiwan</collectingCountry>
|
||
becomes available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |